Skip to main content

Full text of "Bulletins of American paleontology"

See other formats


+ 


? ee a. 


D: . > Dtea S 


HARVARD UNIVERSITY. 


LIBRARY 


OF THE 


MUSEUM OF COMPARATIVE ZOOLOGY 


gl 


Stk 
ae 
Ons 


i 


rr 
ft ea 


ines 


eigen 
eee a 


se 


BUMEE TINS 
AMERICAN 


PALEONTOLOGY 


VOL. 9 


March, 1921 — June, 1922 


Harris Co. 
Cornell Univ., Ithaca, N. Y. 
LOSES oa 


CONTENTS OF VOL. IX 
Bull. No. 
37 —New Eocene Species from Alabama. 
Bye eten VAUl cliriela ys) ae ee even Pl’s 1-3 


38 —Recent Mollusca of the Gulf of Mexico 
and Pleistocene and Pliocene 
Species from the Gulf States. 
Part 2.—Scaphopoda, Gastropoda, 
Amphineura and Cephalopoda. 
By Ce iaiMaunyansneeee eee aaeies 
39 —The Miocene of Northern Costa Rica 
with Notes on its General Strati- 
graphic Relations. 
hy Js oe (OSS0ysljgadenssessuasansose Pl’s 4-35 


Page 


1-32 


33-173 


174-482 


13, 96) 


BULLETINS 
OF 


AMERICAN PALEONTOLOGY 


Vol. 9 


No. 37 


NEW EOCENE SPECIES FROM ALABAMA 


BY 


T. H. ALDRICH 


March 6, 1921 


Ne 


< 
* Harris Co. 
Coriell Univ., Ithaca, N. Y. 
LB SEN 


PREEACEH. 


The late Herbert H. Smith, curator of the Alabama State 
Museum of Natural History University, Alabama, made exten- 
sive collections in the Tertiary of Alabama, and at the time of 
his sudden and lamented death was engaged in classifying and 
arranging his material. The collections were made on joint ac- 
count for the Carnegie Museum of Pittsburg and the Alabama 
Museum of Natural History. The condition of the material 
made it necessary to go over all of it, and classify it for division 
as well as to describe what was considered new. The writer has 
also added a few new species from his collecting. It is regretted 
that Mr. Smith was not spared to complete his labors, because - 
his ability in specific determinations was almost unique. The 
types of the new species collected by him are all in the Univer- 
sity Museum. We are indebted te—Dr. W. H. Dall, Dr. Paul 
Bartsch, Prof. G. D. Harris and Dr. F. A. Smith for generous 
aid in both specific and generic determinations of doubtful forms. 


a ey 


leak 


NEw HocENE SPECIES FROM ALABAMA 
BY 


he ae eA RICE 


Mitramorpha turriculata, n. sp. lel iG Sebo 10 

Shell small, biconic, whorls eight, three embryonic and 
smooth, the fourth with strong longitudinal ribs, which continue 
over the other whorls, dying down some on the body whorl. 
Spiral strize numerous, and impressed, a strong double band bor- 
dering the suture, aperture narrow, smooth within. Length 
6 mm. 

Locality.—Choctaw Corner, Ala. Woods Bluff horizon. 

Type.—My cabinet. 

Remarks.—This little shell seems to be nearest to Witramor- 
pha, it has a glazed appearance ; there is some evidence of a 
slight notch on and above the periphery of the body whorl. 


Turris (Pleurotoma) nodoideus, n. sp. Pls te Riss ya 

Shell small, spire pointed, whorls nine ; first four embryon- 
ic, the fifth bearing longitudinal ribs in part, and revolving lines 
in part with the ribs also present but without the subsutural col- 
lar below. ‘The sixth, seventh and eighth whorls with the bor- 
dering part of the ninth carrying first, a subsutural collar which 
becomes doubled and thicker towards the body whorl and is 
marked with very numerous raised lines belonging to the fasciole; 
second, just below is a rather broad concave revolving space 
reaching to the peripheral parts of the whorls carrying very 
numerous close set semicircular anal markings, these continue 


6 BULLETIN 37 6 


over the body whorl in curved raised lines : third, the periphery 
sometimes carrying a couple of coarse lines followed below by 
from four to six raised lines, also raised longitudinal ribs from 
the periphery to suture. Body whorl with raised revolving lines 
extending to the end of the canal, carrying a few alternating 
ones belowthe periphery ; aperture smooth within ; canal slightly 
rostrate. 

Locality.—Cave Branch, 6 miles E. of Thomasville, Ala. 
Woods Bluff beds. 

Type.—Ala. Museum. University, Ala. 

Remarks.—This species seems to differ in some respects from 
any heretofore described, but it may be only a variety of some 
existing species like Zurris denticula Bast. et. al. The fact is 
thatthe Woods Bluff beds at Cave Branch contain so many numer- 
ous varieties that it is almost impossible to locate them property. 
The whole subject of species in this family needs careful study. 
The young are often obtuse, and differ in shape and ornamenta- 
tion from the adult so that they could be called new species, and 
pass muster while a series would connect them all together. Am 
not prepared to go into details but hope this subject will be 
looked into by some of our younger students. Some specimens 
may be hybrids, and are altogether perplexing. One ought to 
have authentic specimens of the described species to even begin 
the examination. All the species of this group from this locality 
seem somewhat stunted. 


Turris specus, n. sp., JEN Neb eR WAR se 

Shell small, whorls probably seven (the embryonic tip is de- 
fective) first three embryonic, then four spiral whorls ornamented 
with a strong subsutural collar, then a slightly concave revolving 
fasciolar space covering about one fourth of the whorl, followed 
by a coarse cancellated surface ; the ribs cut by about four im- 
pressed revolving lines making a series of nodes, body whorl with 
the same ornamentation above followed by coarse revolving lines 
growing finer to the base ; outer lip smooth inside, columella 
thickened, base somewhat rostrate. 

Locality,—Cave Branch, Ala. Woods Bluff beds. 


7 ALDRICH, KOCENE MOLLUSCA 7 


Type.—Ala. Mus. of Nat. History, University, Ala. 
Microdriilia rostratuia Csy. 

This species described by Col. Casey in 1903 bears a certain 
resemblance to Pleuvotoma insignifica Heilpr. (Fusus nanas Lea) 
and has been referred to this species, but specimens before me 
show differences as follows: Col. Casey says the ‘‘body whorls 
are about four in number.’’ My specimens have five, also in the 
embryo there is sometimes but one costulate whorl, while 
Pleuro. insignifica Heilpr. has but three embryonic whorls, all 
smooth, and no costulate ones. The two species seem to be 
quite distinct. The Texas form mentioned by Prof. Harris is 
probably the same as Heilprin’s species. 


SPECIES TO BE ADDED TO THE Woops BLUFF FAUNA— 
FROM CAVE BRANCH, ALA. 


Exilia pergracilis Con. 
Cylichna deKayt Lea. 
Cancellarta alveata Con. 

ae costata Lea. 
Pasithea guttula Lea (H. H. Smith). 
Cancellaria panones Harris. 
Ancillaria subglobosa Con. 
Levifusus sutert Aldr. 
Caricella dolita Con. 
Drillia lonsdalit Vea. 
Barnea alatoidea Aldr. 
Diplodonta nana Vea. 
Egerella subtrigonia Lea. 
Tellina leana Dall. 


Turris bimoniatus, n. sp. Pliitea), Hig: 

Shell small, whorls probably ten, spire on the type partially 
missing. Another specimen shows four embryonic smooth 
whorls. The four spiral whorls and part of the fifth ornamented 
with curved sinuosities covering the main part of the whorls, a 


8 BULLETIN 37 8 


strong collar below the sutures, the balance showing a double 
collar with a concave space between ; the whole of the surface 
with numerous spiral raised lines. Aperture about a third of 
the length of shell, lines on the base have a tendency to alter- 
nate. Length 14 mm. ! 

Locality.—Cave Branch, Ala. Woods Bluff beds. 

Type.—Ala. Museum, University, Ala. 

Remarks.—'This seems to combine the characteristics of two 
or more species, and if there were only one specimen, could be 
considered a variety. It seems to be related to 7. monzliata 
Heilpr. 


Pleurotoma (Peratotoma) Gardneri, n. sp. TEN sey) Teds ich 


Shell medium, substance rather thin, broadly conic ; whorls 
nine to ten, embryonic ones four in number, the spire very small, 
but the whorls rapidly expanding, the permanent whorls profuse- 
ly ornamented with both longitudinal and spiral raised lines, the 
peripheral part and below carrying nodes while the spirals are 
alternate in size and thickness. Body whorl large, shouldered, 
profusely ornamented, the spirals nodular ; just below the suture 
there are a couple coarser than the others and also at base ; aper- 
ture half the length of the shell, smooth within, inner lip witha 
thin callus ; canal recurved and slightly spatulate. Length 
21 mm. 

Locality.—Pugh’s branch of Satilpa Creek, Ala., Gosport 
Sand. 

Type.—Ala. Museum of Nat. History, University, Ala. 

Remarks.—This species is on the borderland between /uszs, 
Strepsidura and Pleurotoma. It isvery close to /l. hetlprini, 
nobis from Jackson, Miss. The largest specimen is 25 mm. in 
length. The slit is not pronounced. Named in honor of Miss 
Julia A. Gardner of the National Museum. 


Odostomia (Evalea) Bartschi, n. sp. eal ats abe ey, 

Shell small, whorls six, spire almost totally immersed, shell 
rather rapidly expanding. Basal whorl large, suture deep, aper- 
ture oblong. One strong, straight fold on the columella placed 


9 ALDRICH, HOCENE MoLLUSCA 9 


centrally; umbilicus open, inner lip somewhat recurved at base. 
Surface not smooth, carrying microscopic incised spiral lines. 
Length 3 mm. 
Locality.—Found at both Gregg’s and Bell’s Landings. 
Type.—From Bell’s Ldg. In the Ala. Museum. 


Turbonilla (Ptycheulimella) clinensis, n. sp. LAL eg) Vetkerh 10), 

Shell small, narrow, whorls eight, two embryonic, the next 
three longitudinally striated, balance smooth. Spire blunt, 
suture distinct, aperture small. Outer lip smooth within, inner 
lip with a strong fold curving up into the aperture and bordering 
the base. Length 3+ mm. 

Locality.—Gregg’s Landing, Ala. 

Type.—Alabama Museum, University, Ala. 

Remarks.—Seems to be a distinct species. There are three 
other species of 7urbonzlla in the State Collection but too imper- 
fect to warrant description. 


Turbonilla (Cinguiina) tuscahomensis, n. sp. enh 3640), Teme ath 
Shell small, whorls nine, the first three embryonic ; apex 
small, the fourth and fifth whorls with raised spirals, balance 
carrying five strong spirals, and somewhat cancellated ; suture 
deeply impressed. The space above the first or lowest spiral 
slightly wider than between the other four. Aperture broadly 
ovate ; inner lip slightly reflected ; base slightly spatulate. 
Length 4mm. Figured specimen from Bell’s Ldg. 
Locality.—Both Gregg’s and Bell’s Ldg., Alabama. 
Type.—Alabama Museum of Nat. History. 
Remarks.—\Looks like a Lzttium. 


Turbonilla (Ptycheulimella) tardiusculus, n. sp. Pie Rigen 2. 


Shell with nine whorls, the first two embryonic, the balance 
smooth and of a dull brown color, substance of shell rather solid, 
suture lightly impressed, banded below. Aperture small, inner 
lip reflected at base, imperforate ; no fold showing on the col- 
umella. Two specimens found. Length4% mm. 

Locality.—Cave Branch, Ala. Woods bluff beds. 

Type.—My collection. © 


fe) BULLETIN 37 10 


Pyramidetta (Iphiana) anita Ald. 

The Nautilus, Vol. XXI, No. I, p. 9, Pl. 1, Fig. 12, May, 1907. 

This species was described as a 7urbonilla, but is now put 
into the Pyramidellide by Dr. Bartsch. 


Pyramideila (Syrnola) mitchelliana Aldr. 

Bull. of American Paleontology, No. 22, P. 8, Pl. 3, Fig. 3, IgII. 

Is also removed from 7urbonzlla. 

The Ala. Museum has a number of new forms of small spe- 
cies, collected by Mr. Herbert A. Smith which are described 
herein. They have been submitted to Dr. Bartsch who has kind- 
ly indicated their generic position. 


Pyramidella (Iphiana) obtusoides, n. sp. IP yes Webel Tey 

Shell small, with four whorls capped by two embryonic ones 
turned abruptly to the back side of shell and partially immersed. 
The specimens, three in number, have a dull surface, apparently 
smooth, but under a glass numerous spiral lines are seen. 
Suture not deeply impressed ; aperture oblong-ovate ; outer lip 
sharp, smooth within ; inner lip reflected, somewhat twisted, 
carrying a single smalltooth. Length 3% mm. 

Locality.—Found at both Gregg’s and Bell’s Landings, Ala. 
River, Ala. 

Type.—Ala. Museum of Natural History. 

Remarks.—A short, dull-looking species. 


Epitonium munistriatum, n. sp. Pion. pehigse tA a5 


Shell small, aciculate whorls, probably toor more. Embry- 
onic ones three in number, apex pointed ; the next two or three 
with longitudinals only, balance cancellated. Whorls rounded, 
suture deep. Spirals 5 to 7, rather coarse, ribs raised with con- 
cave spaces between, base with smaller alternate spirals between 
the primaries ; intersections pointed. ‘The base of this species 
(15) is in the Museum from Gregg’s Landing, Ala. The other 
specimen is from Woods Bluff, Ala. The aperture shows a 
a thickened inner lip where it joins the base, also a thin callus. 

Locality.—As above. 

Type—Ala. Museum of Nat. History. 


II ALDRICH, KOCENE MOLLUSCA Il 


Epitonium multiliniferum, n. sp. Veda GD ahkeg atop 

All the specimens are fragmentary, but well marked. Em- 
bryonic whorls missing ; suture distinct. Ribs on body whorl 
34 in number, extending over the base, much weaker there ; 
very numerous close set spiral lines between the ribs. | Aperture 
round, inner lip thickened, no umbilicus. 

Locality.—Cave Branch, Ala. Woods Bluff beds. 

Type.—My cabinet. 

Remarks.—This is a distinct species and it deserves differen- 
tiation from the other forms. ‘The figure given by Prof. Harris 
in Bull. Am. Paleontology, No. 11, Pl. 12, Fig. 8, is probably a 
young example of this species. The sides are more nearly par- 
allel than in most Apztonzums. 

Epitonium subacutum, n. sp. Pir Bigs s 175 0o: 

Shell thin, narrowly acute, whorls 12. The first five are em- 
bryonic and smooth ; the balance spirally striated ; cancellated. 
The vertical ribs on the body whorl about twenty-six in number, 
curving into the umbilicus. They are thin or acute ; the pri- 
mary spirals about ten in number with finer alternate lines be- 
tween ; inner lip with a slight callus. Length 10 mm. 

Locality.—Cave Branch, Ala. Woods Bluff beds. 

Type.—In Ala. Museum. One specimen shows the base and 
the other the embryonic shell. 


Teinostoma subangulata Mr. var. Smithii, n. var. VEAL id ty |i dietary, ako}, VIO 

Shell small, flattened above, rounded below ; umbilicus not 
large ; a callus on the base which enters the umbilicus and con- 
tinues within. Bordering the suture is an appressed area raised 
above the whorls ; surface above with numerous close set revolv- 
ing lines ; periphery smooth with a few revolving lines below. 
Aperture circular ; outer lip somewhat thickened ; lines of 
growth perceptible ; interior somewhat pearly. Diameter 8 mm. 

Localtty,—Bell’s Landing, Ala. 

Type.—Ala. Museum of Natural History. 

Remarks.—Very much larger than the type of 7. subangu- 
lata Mr. Looks very much like a land shell. It differs some- 
what from Meyer’s species in the shape of the aperture. 


12 BULLETIN 37 12 


Teinostoma reguiaris, n. sp. Bie igs. 2 1-2» 


Shell small but larger than the ordinary species of this 
genus ; whorls five, depressed above ; periphery rounded ; sur- 
face covered with fine lines which are crossed with lines of 
growth. Under a glass the intersections give the surface a 
beaded appearance ; a depressed space just below the suture. 
The base has coarser lines with a smooth space around the um- 
bilicus which is open ; aperture oval. Diameter 4 mim. 

Locality.—Pugh’s Branch of Satilpa Creek, Clark Co., Ala. 
Gosport Sand. 

Type.—Ala. Museum at University, Ala. 

Remarks.—This little shell may possibly belong to the sub- 
genus Omphalius of Chlorostoma but it seems closer to Zeznos- 
toma. 


Ancillopsis Tuomoyi, n. sp. Teds ttys | JE hes\, Ae oy, 


Shell solid, thick, oblong, smooth and shining ; whorls 5-6 ; 
spire rather pointed ; the first four whorls small, the others rap- 
idly increasing ; a swollen callus reaching above the body whorl 
and also covering the front of this whorl and lapping over the 
back of this whorl. A deep suture reaches the aperture where 
it is rather wide ; a few irregular lines of growth are visible ; 
aperture about three-fourths the length of the shell, oblong-ovate; 
columella smooth, concave. The basal groove shows strongly at 
base, but only showing a short distance up where it is covered by 
the enamel. Length 28 mm ; breadth 21 mm. 

Locality.—Bell’s Landing, lower bed, also one example from 
Gregg’s Landing, Ala. The lower bed at Bell’s Landing is the 
same bed as at Gregg’s Ldg. ‘Type from Bell’s Landing, lower 
bed. 

Type.—Ala. Museum of Nat. History. 

Remarks.—This species resembles //acron in some respects 
but is evidently an Azeczllopsis. 


Nassa pleona, n. sp. Plvmeebigsy 25.26: 


Shell small, nuclear whorls broken off. Four remaining 
whorls carry longitudinal ribs that are crossed by two spirals, 


13 ALDRICH, KOCENE MOLLUSCA 13 


nodular at intersections. Two close set spirals border the suture 
above and below same; suture channelled ; the last whorl car- 
ries about Io widely spaced spirals, raised and nodular with con- 
cave spaces between. Outer lip thickened with the spirals ex- 
tending over same, denticulated ; inner lip enameled with the 
spirals showing through the same. Canal very short ending in 
a semicircular opening. Length 12+ mm. ‘Two specimens ob- 
tained. 

Localtty—Lisbon, Ala. 

Type.—My cabinet. 

Glivella semilignitica, n. sp. Te ebhers: By, ates 

Shell small, whorls six, four smooth ; the fourth and subse- 
quent ones bounded by a raised band just below the suture ; apex 
blunt, body whorl showing impressed lines of growth ; aperture 
asin the genus. Length 5% mm. 

Locality.—Clark County, Ala. Woods Bluff horizon. 

Type. —Ala. Museum of Nat. History. 

Remarks.—The sutural callosity pronounced on the last 
whorl only. Differs from O. mediavia Harris in being smaller, 
more blunt, with fewer folds on the columella, and by a raised 
sutural band. 


Gilbertia esteliensis, n. sp. AL Ds | Jaber eas 
Shell globular, whorls three ; the first embryonic ; the sec- 
ond spirally striated but very narrow ; the third very globose 
and spirally striated ; outer lip thickened and broadly reflected ; 
a single tooth projecting into the aperture from it ; inner lip 
thickened with a broad spatulate tooth above, and two prominent 
teeth near the base ; the outer lip partially covering the second 
whorl. Under a glass the strize show that they are pitted as in 
Actgon. Height 2% mm. ; breadth 2 mm. 
Locality.—Sucarnoochee beds, 3 miles south of Estella, Ala. 
Ty pe.—Ala. State Museum, University, Ala. 


Canceilaria Gilberti, n. sp. Ble ie REO 20% 


Shell with seven whorls remaining ; the spire is damaged ; 
cancellated ; suture deeply impressed and constricted, especially 


14 BULLETIN 37 14 


on the lower whorls. Below the suture is a raised cord marked 
with a spiral line upon the lower whorls; above the suture is a 
wider space bounded by strong raised spirals with other raised 
spirals above. Body whorl expanded with a strong spiral at the 
periphery ; those above this are close together, those below al- 
ternated with a finer spiral; lines of growth show on body whorl 
reversing near the base ; aperture oblong-ovate, the spiral lines 
rounding into same ; columella with two folds ; shell has a small 
umbilicus. Labrum lirate within. Length 9 mm.; breadth 4mm. 

Locality.—Grege’s Ldg., Ala. River, Ala. 

Type.—Ala. Museum of Nat. History. 

Remarks.—Only one specimen found. This species has 
many of the characters of Admete. 


Qvula reguiaroidea, nu. sp. LEI Se |) Deabezy evo 


Shell small, elongate ; both anterior and posterior with fine 
revolving lines ; main body of shell smooth ; outer lip thickened 
and crenulated from beak to base ; inner lip also crenulated its 
whole length ; a heavy callus on the outer surface of the col- 
umella at the posterior end with the canal extending beyond the 
callus. On the smaller of the two specimens before me there are 
fine revolving lines half way down from the posterior ; aperture 
narrow, expanding back of the anterior canal ; ends somewhat 
flattened. Length of largest specimen 14 mm. 

Locality.—Bell’s Landing mari, Ala. River, Ala. 

Type.—Ala. Museum of Nat. History. 


Cyprea estellensis, n. sp. ISR) erica 


Shell small ; rotund ; smooth ; aperture narrow ; outer lip 
curving into the apex which is open. Lengthi2mm. Breadth 
ro mm. 

Locality.—Sucarnoochee beds, Pursley Creek, Wilcox Co., 
Ala. 

Type.—Ala. State Museum. 

Remarks.—This specimen has only preserved the shell in 
part but the interior is hard rock and gives the form quite accur- 
ately. The aperture is all filled with matrix, hiding the denti- 


15 ALDRICH, HKOCENE MOLLUSCA 15 


tion. Prof. Harris mentions finding one species at Fort Gaines 
on the Chattahoochee and another (which is probably the one 
subsequently described % mile north of Ripley, Miss.) They 
are figured and named as they seem to be rare and peculiar to 
this horizon. 
Cyprea sp.? Pia i Bigsuauae 

Shell small ; surface smooth ; basal callus extending par- 
tially over the sides ; outer lip expanded ; extremities somewhat 
pointed. One fragment shows the dentition, consisting of over 
20 small teeth on the inner lip. Length of the larger fragment 
15 mm. 

Locality.—Sucarnoocchee beds, one and three miles south of 
Estella, Ala. on’ Pursley Creek. 

Types.—Ala. Museum of Nat. History. 

Seems to be rather close to Cyprea Smutht nobis from 
Grege’s Ldg., Ala. 


Siphonalia quadrilineata, n. sp. Panos Ak ie 


Shell resembling the Ausus subscalarinus Heilpr. Whorls 
eight, two embryonic, the balance more or iess cancellated ; 
longitudinal costee somewhat rounded; spiral striee very numer- 
ous, four fine ones between a coarser line ; the volutions are ap- 
pressed at the suture ; the spiral striz cross over the costee and 
the coarse lines make nodes of intersections ; canal reflected and 
twisted : the outer lip on the type is broken away but appears to 
have been smooth internally ; inner lip somewhat thickened. 
Length 17mm. 

Locality.— Pugh’s Branch of Satilpa Creek, Clark Co., Ala. 
Gosport Sand. 

Type.—In Ala. Museum, University Ala. 

Remark.—One is rather rash to attempt a new form herewith 
but the surface ornamentation is peculiar and seems to be new. 


Cerithiopsis esteliensis, n. sp. VEIL LN Deikex, 1K)s 

Shell minute ; whorls eleven or more, the first two smooth, 
the next two with raised ribs ; the balance showing two strong, 
heavily beaded spirals on the peripheral part of each whorl with 


16 BULLETIN 37 16 


a third spiral much finer just below the suture ; one or more 
whorls are missing from the basal part. 
Length 3 mm. 


Locatity.—Sucarnoochee clays, near Estella, Ala. 
Type.—in Ala. Museum at University, Ala. 


Leevibuccinum (Euryochetus ?) harrisi, n. sp. Ply 2h) VEtes. 7-9: 

Shell small ; whorls eight, three smooth, the other five be- 
low {with close-set raised lines ; outer lip smooth, slightly in- 
curved ; inner lip with a callus, thicker and slightly raised on the 
canal, which is short, rather wide ; shell somewhat truncated an- 
teriorly ; suture distinct. Length of largest specimen 44% mm. 

Locality.—Cave Branch, Ala., about 6 miles east of Thomas- 
ville, Ala. Woods Bluff beds. 

Type.—Ala. Museum of Nat. Hist., University, Ala. 

Remarks.—This shell is quite fragile, the body whorl break- 
ing away from the spire. Prof. Harris figured the spire of this 
species under Aesopus erectus (Proc. Acad. Nat. Sciences, Phila., 
1896, p. 476, pl. 21, fig. 3), which it resembles, but later on 
states it isnotthesame. ‘The Ala. Museum has three specimens, 
and there are two broken ones in my cabinet. 

Type.—Ala. Museum of Nat. History, University, Ala. 

TENUIACTAGN, n. g. 

Shell resembling Actegon but much more slender ; whorls 
nine ; aperture small, spire turned to the left and partially im- 
mersed ; suture deep; columella with one fold ; surface with 
numerous incised revolving lines. 


Tenuiactzon pertenuis, n. sp. Ply 237) Eig: wo: 


Shell small, slender ; whorls 9 ; surface with numerous re- 
volving incised lines, stronger at base but almost disappearing at 
the shoulder of body whorl ; then just below the suture showing 
two or three stronger ones ; apex rounded and turned to the left, 
partially immersed ; outer lip somewhat incurved ; base round- 
ed ; columella with one strong fold ; the inner lip reflected over 


the body whorl ; suture deep, giving each whorl the appearance 
of being inserted into the next succeeding one. Length 13 mm. ; 


breadth of body whorl 4 mm. ; aperture 5 mm. 


17 ALDRICH, EKOCENE MoLLUSCA 17 


Locality.— Gregg’s Ldg. marl at Bell’s Ldg. and Gregg’s 
Ldg., Ala. River, Ala. 
Type.—Ala. State Museum at University, Ala. 


Teredo ringens, n. sp. JEM as) i saab ao 

Shell substance thin, closed at the larger end like Auphus ; 
body of shell with raised, rather acute rings parallel to each 
other, and virtually at right angles to the longer diameter. 
Other fragments are not so stongly marked. Length of type 
‘specimen 37 mm. Average breadth 5 mm. 

Locality.—Sucarnochee clay bed, 3 miles south of Estelle, 
Ala. 

Type.—Alabama Museum, University, Ala. 


Tellina estellensis, n. sp. PEI aon | Deb vedsyy ney Yl 


Shell small, exceedigly thin, exterior smooth, except fine 
growth lines ; these are stronger near the ventral margin ; in- 
equilateral ; right valve with small laterals, larger in the oppo- 
site valve ; pallial sinus large, rounded ; anterior somewhat 
pointed ; posterior rounded. Length 13-15 mm. ; height 9-11 mm. 

Localityx—Sucarnochee beds, near Estelle, Ala. 

Type.—Ala. Museum of Nat. History, University, Ala. 

Remarks.—This species is the same figured by Prof. Harris 
(Bulletins Am. Pal., Vol. 1, p. 182) asacast. It is rather com- 
mon but so very fragile it was almost impossible to procure a per- 
fect example. 


Tellina cynoglossa Dall. 


This is the common species in the Woods Bluff beds. It 
seems to be only a variety of 7ellina subtriangularis Aldr., which 
was described from half grown specimens, which is more regu- 
larly rounded posteriorly and also lacks the fold seen in older 
specimens. The full grown examples from Woods Bluff on the 
Tombigbee River measure as much as 25 mm. in length 


Tellina bellsiana, n. sp. Pay Rigs.) 15) LO: 


Shell large, flat, thin ; nearly equilateral ; surface faintly 
concentrically striate ; lines:growing coarser on the anterior end ; 


18 BULLETIN 37 18 


beaks very small ; the dorsal area strongly bent, pallial area gib- 
bous ; teeth normal. Height about 24 mm. 

Locality.—Both Gregg’s and Bell’s Landings, Ala. River, 
Ala. 

Type.—Ala. Museum of Nat. Hist. 

Remarks.—One example in the Carnegie Museum. The 
type is broken some, shell thin for its size. 


Tellina semirotunda, n. sp. peas Maleate 1077, 


Shell medium ; nearly equilateral ; anterior somewhat 
pointed ; surface smooth, except lines of growth irregularly 
spaced ; a slightly depressed area extending from beak to base 
behind the anterior fold. Cicatrices rather large, are well 
marked ; pallial area large ; cardinals bifid. Length 16 mm. ; 
height 9 mm. 

Locality.—Both Gregg’s and Bell’s Ldg., Ala. River, Ala. 

Type.—Ala. Museum of Natural History. 


Tellina semipapyria, n. sp. Vey ay) Dene, u's), 


Shell medium, resembling 7Tel/ina semtrotunda nobis ; sur- 
face smooth except growth lines ; pallial sinus large, partially 
confluent with the pallial line ; only left valve known. 

Locality.—Jackson’s Rockhouse branch. Woods Bluff beds. 
Ala. Length 23 mm. ; height 9mm. 

Type.—Ala. Museum of Nat. History. 

Remarks.—Bears a strong resemblance to the Claibornian 
species Tellina papyria Con. but is somewhat more rotund. 


Semele langdoniana, n. sp. Jes mle, 16), AO: 


Shell oblong, oval ; surface practically smooth, showing 
very fine lines that are coarser on the umbonal slope ; some spec- 
imens show also fine radial lines reaching to the ventral margin ; 
posterior shortest ; ventral margin entire ; pallial sinus profound. 
Shell looks like a 7el/ina externally. Length of largest spect- 
men is about 27 mm. 

Locality.—Bell’s Landing, Monroe Co., Ala. River, Ala. 

Type.—(Left valve) Ala. Museum, University Ala. 

Remarks.—There are several fragmentary specimens in the 


19 ALDRICH, KOCENE MOLLUSCA 19 


lot. ‘This species is named after the late Dr. W. Langdon, an 
Assistant State Geologist of Alabama and the discoverer of the 
Floridian Miocene. 


Semele monroensis, n. sp. 2a REN SIS 2 22. 


Shell small, rather flat ; one right valve only in the collec- 
tion ; umbonal slope strongly bent ; surface smooth above with 
several widely spaced lines towards the ventral margin ; hinge 
stout ; ligament long for the genus ; the umbonal slope marked 
interiorly by a few radial lines. Length 5 mm. 

Locality.—Bell’s Landing, Ala. River, Ala. 

Type.—Ala. Museum of Nat. History. 

Remarks.—The specimen described is quite young but dis- 
tinct. Looks like a Weretrix in shape and ornamentation. 

Lucina primoidea, n. sp. RIM 2aNOnigSN 22h 2A. 

Shell with many raised lines ; beaks recurved, higher than 
long. ‘This species is figured because it is from an horizon close 
to the Cretaceous and is doubtless an ancestral form of Luczna 
cornuta Conr. of the Claibornian. The interior is not accessible. 

Locality.—Black Bluff, Tombigbee River, Ala., about 20 feet 
above the base. 

Type.—My collection. 


Martesia recurva, n. sp. RANI eS 5.20. 


Shell small, short, rotund; with two accessory plates just 
forward of the umbones ; groove running nearly vertical from 
beaks to base and strongly marked ; concentric strize strongly 
marked on the posterior end, bending abruptly at the groove and 
thence running up behind the umbones and under the accessory 
plates ; the anterior end rounded and smooth ; cardinal margin 
bordered with a thickened and raised process. Length 8 mm. ; 
height 4% mm. 

Locality.—Fleming’s Mill, Ala.. on Pea River. Nanafalia 
beds. 

Type.—My collection. 

Remarks.—This species is more obtuse than the usual forms. 
Three examples found. ‘The interior is not accessible. 


20 BULLETIN 37 | 20 


Rochefortia minuta, n. sp. Pl 2tyibiigss 27.928 


Shell, small, ovate ; nearly equilateral ; periostracum still 
adherent, surface showing a few incremental lines ; beaks point- 
ed, with a resiliary pit underneath with short lamelle on each 
side, the posterior one the longest ; muscular scar slightly im- 
pressed ; the right valve has a space for the left lateral ; pallial 
line simple. Resembles somewhat Rochefortia Stimpsont Dall. 
The space in our specimen under the beak looks as though it had 
been excavated out of the dorsal side, but such is not really the 
case. Prof. W. H. Dall has examined this specimen, and veri- 
fied its generic position. Am greatly indebted to him in a num- 
ber of cases. Length 3 mm. ; height 2 mm. 

Locality.—Bell’s Ldg., Alabama River, Ala. 

Type.—Ala. Museum. 


Montacuta bicuspidata, n. sp. * Pl. 2, Figs. 29, 30. 


Shell minute, oblong-ovate ; surface with incremental lines ; 
beaks carrying the prodissoconch, nearly central ; viewing the 
shell from the outside and above, it shows two tooth-like projec- 
tions, one on each side of the beak, rising from the dorsal mar- 
gin and another shorter one which does not show like the two 
first mentioned ; ends rounded ; base arcuate ; hinge with a small 
subtrigonal tooth and also a space for a lateral in the other valve. 
Pallial line simple ; adductors distinct but not deep. Length 234 
mim ; height 2 mm. 

Locality.—Bell’s Ldg., Ala. 

Lype.—Ala. Mus. of Nat. History, University, Ala. 


Basterotia ? prima, un. sp. PLE aaatkigs amt anne 

Shell small ; valves rather thick ; subquadrangular ; beaks 
subterminal ; shell profoundly gaping at the anterior ventral 
part ; a depressed space running from umbo to ventral margin ; 
surface nearly smooth, showing growth lines which also show in 
the interior. Anterior adductor scar deep, with a raised 
border buttressed to hinge plate ; posterior scar slightly im- 
pressed, pear-shaped and both rather large for the size of shell ; 
one cardinal tooth inright valve, corresponding to a deep socket 


20 ALDRICH, FOCENE MOLLUSCA 43 


in the other ; a longer tooth below. Three single valves in the 
collection. Length 8mm. ; height about 7 mm. 
Locality.—Gregg’s and Bell’s Landings, Ala. River, Ala. 
Type.—In Ala. Museum. 
Remarks.—This species is doubtfully placed in Basterota ; 
the large gap suggests an attachment to some other form, but is 


not irregular. Prof. Dall has kindly examined it, and suggests 
that it might be commensal. 


Saxicavella alabamensis, n. sp. Rl ae igs a Aays. 


Shell inequivalve ; thin ; posterior much broader than the 
anterior ; outer surface showing irregular lines of growth, 
stronger over the umbonal slope ; an upright cardinal tooth 
directly under the umbo, with a short lateral on posterior side in 
the right valve ; anterior muscular impression distinct ; pallial 
line faint. Only one valve in the collection. Length 6 mm. ; 
height 4% mm. 

Locality.—Bell’s Landing, Monroe Co., Ala. 

Type.—In Ala. Museum. 


Montacuta Herberti, n. sp. Pee eS MOL 7. 


Shell small; inequilateral ; dorsal and ventral margins 
rounded ; each end rather blunt; the posterior somewhat 
angulated ; surface smooth with very fine concentric striations, 
and a few lines of growth near the ventral margin ; shell inflat- 
ed ; cardinals feeble, showing in left valve only ; beaks low, 
rather pointed, a small buttress under them. Length 9 mm ; 
height 6% mm. 

Locality—Cave Branch, Ala. 

Type.—Ala. Museum of Nat. History. 

Remarks.—One valve in my collection. Named in honor of 
the late Herbert H. Smith. 

Ervilia lignitica, n. sp. TMi gin! | TEAS toys 

Shell small, inflated ; nearly inequilateral ; rather solid ; 
the posterior slightly longer than the anterior ; both ends round- 
ed ; surface carrying a few irregularly spaced grooves of growth; 
interior polished, shining ; pallial sinus small, not reaching to 


22 BULLETIN 37 22 


center ; umbones small, dentition as in the genus with short and 
strong teeth. Length 6 mm. ; height 3% mm. 

Locality.—Both Gregg’s and Bell’s Landing beds, Ala. 
River, Ala. 

Type.—Ala. Museum of Nat. History, University, Ala. 

| Remarks.—This species has a strong resemblance to A7vvzlia 
polita Dall from the Floridian Pliocene. The young seem to be 
higher in proportion to their length than the full grown speci- 
mens. : 


Panopea bellsensis, n. sp. AL Ss Seite, GO) 18) 6 


Shell rather large, thin, with a brown epidermis ; lines of 
growth irregularly spaced ; shell compressed, nearly equilateral ; 
beaks small, appressed to and partly covered by the reflected dor- 
sal margin; one large cardinal tooth ; ligamental attachment 
short ; interior porcellaneous with growth lines showing through; 
pedal scars small. Length 30 mm. ; height about 13 mm. 

Locality.—Bell’s Ldg., Ala. River, Ala. 

Type.—Ala. Museum of Natural History. 

Remarks.—One valve and some fragments in the collection. 
This species has its epidermis preserved. ‘The shell is extremely 
thin and delicate. 


Donax acutangula, n. sp. NG gy 2 IWikex, 102), 


Shell nearly equilateral ; pointed at both ends, the umbonal 
slope bent abruptly to almost a right angle in old specimens ; 
surface nearly smooth ; fine growth lines showing ; ventral 
margin entire ; beaks small, pointed, smooth ; hinge well devel- 
oped ; laterals very long, striated on the inner edges. Length 
about 31 mm. ; height 15 mm. 

Locality.—Bell’s Landing, Ala. River, Ala. 

Type.—Ala. Museum of Nat. History, University Ala. 

Remarks.—One whole valve and several fragments of both 
valves received. 


Arca (Cucullaria) ozarkensis, un. sp. Bee enbigs nian eran 
Shell small, rounded at both ends, giving the shell a quad- 


23 ALDRICH, HOCENE MOLLUSCA 23 


rangular outline ; beaks small, compressed and pointed ; surface 
with rather close set radial lines on the posterior slope ; balance 
of the surface smooth ; teeth as in the subgenus, three or four 
anterior and two posterior ; muscular scars shallow but plainly 
marked. Length 5 mm. ; height 3 mm. 

Locality.—Woods Bluff beds near Ozark, Ala. 

Type.—My cabinet. 


Ostrea intermedoides, n. sp. Ne Debby ans sip 

Shell medium, valves rather thin ; surface in older speci- 
men with numerous raised concentric ribs ; the lower valve bent 
downwards near the beak ; the umbo shows radial strize ; mar- 
gins crenulated internally ; a fragment of the upper valve ap- 
pears to be flat. Length of lower valve 32mm. ; breadth about 
20 mm. 

Locality.— Bell’s Landing marl, Bell’s Landing, Ala. River, 
Ala, 

Type.—Ala. Museum of Nat. History. 

Remarks.—Have hesitated to add another oyster to our Eo- 
cene, but I have been unable to find a place for this species. 
Four lower valves and a fragment of the upper valve are in the 
collection. The ornamentation recalls O. falco Dall from the 
Jacksonian. 

Psammobia Smiihi, n. sp. Ply sah iesa. l7, plo: 

Shell large ; substance of shell thin ; oblong-ovate ; rather 
compressed ; surface shining, with fine concentric sculpture 
medially ; both ends coarser and elevated ; on the posterior 
slope the laminze are rough and Somewhat mammillated ; beaks 
very sinall ; muscular scars distinct but shallow. Length 60 
mm. ; height 32 mm. ; Breadth about 12 mm. 

Locality.—Gregg’s Ldg. marl at Bell’s Landing, Ala. Riv- 
er, Ala. 

Type.—Alabama Museum of Nat. History, University Ala. 

Remarks.—Both valves of this splendid species are in the 
collection. It bears a general resemblance to P. /i/osa Conrad, 
but the dimensions are very different. Collected by the late H. 
H. Smith and named in his honor. 


24 BULLETIN 37 24 


Psammobia harrisi, n. sp. Pl. 12.) Bigs..19,, 20! 


Shell nearly equilateral ; truncate behind, with fine concen- 
tric sculpture, coarser at each end ; ligamental area short ; cica- 
trices distinct ; lateral tooth long ; posterior one short. Length 
about 37 mm. 

Locality.—Gregg’s Landing, Ala. River, Ala. 

Remarks.—Vhis species resembles P. ozarkana Harris but is 
more obtuse. ‘The type is not perfect. 


€ 


DIVERSA 
(Echinocyamus ?) meyeri, n. sp. PIS a Mies Ha 1 228 


Test small, ovate ; flattened below ; deeply pitted with ovate 
scars above which become nearly square on the sides and below ; 
substance of the test of moderate thickness ; apicial system near- 
ly central ; ambulacral petals relatively short, straight, with 
numerous round pores. Peristome is medium circular, and 
slightly depressed ; periproct not small, closer to the peristome 
than the side. Length 5 mm. ; width 3% mm. ; height 3 mm. 

Locality.—Gosport Sand, Claiborne, Ala. 

Type.—My cabinet. 

Remarks.—This species is peculiar in the deep pits covering 
the surface. It is larger than &. huxleyanus Meyer and more 
obese though egg-shaped. Received from Dr. Otto Meyer in 
whose honor I have named it. 


NOTES 
BLACK BLUFF OR SUCARNOCHEE HORIZON. 


Prof. H. H. Smith and wife collected a number of species 
from these clays not far from Camden, Ala., in the vicinity of 
Estelle, Ala. The deposit consists of an aluminous clay, dark 
chocolate in color, and contains the greater part of the species 
mentioned by Prof. G. D. Harris from the bluff at Fort Gaines 
on the Chattahoochee River where they seem to be represented 
principally by casts. Those near Estelle are rather numerous, 
but very fragile. A few new species are described in this paper. 


25 ALDRICH, HOCENE MoLLUusCcA 25 


A surprising fact is that a minute Corbula, Corbula (Aloid:s) 
milium Dall, originally described from the Wilcoxian is extreme- 
ly common. ‘The supposed differences in the specific fauna is 
explained. The two localities are over 100 miles apart. The 
list of species from near Hstelle is as follows : 


Leda saffordana Harris. 

‘“ parva Rogers. 
guercollis Harris. 
mitlameensts ‘* 
Corbula (Aloidis) milium Dall. 
Glycymeris cf aviculoides Con. 
Pinna sp. ? 
Teredo ringens Aldr. 
Yoldia eborea Con. 
Venericardia wilcoxensis Dall. 
Tellina estellensts Aldr. 
Protocardia Harrist Dall. 
Pecten alabamensis Aldr. 
Ostrea ? young. 
Nucula ovula Lea. 


6c 


6 


Ce. 


medtavia Harris. 
Modtola saffordi Gabb. 
Meretrix ripleyana Gabb. 
Cucullaa saffordt Gabb. 

i macrodonta Witt. 
Crassatellites sepuicollis Warris. 

a gabbi Safford. 
Arca sp.? 
Corbuls subcompressa Gabb. 
Strepsidura fleilprint Aldr. 
Volutilithes rugatus Con. 

a limopsts Con. 
Turritella tennesseensts Gabb. 

i mortont Con. 
Aumerosa Con. 
Turritella alabamiensts Whitt. 


be 


BULLETIN 37 


Triton showaltevz Con. 
Solarium periscelidum Dall. 
Solariclla alabamensis Ald. 
Rissoina alabamensis Ald. 
Pseudoliva unicarinata Ald. 
ng scalina Heilpr. 
Pleurotoma quercollis Harris. 
i persa Whitt. 
meedtavia Harris. 
os adeona Whitt. 
longipersa Harris (var.) 
Calyptraphorus velatus, var. compressus Aldr. 
Olivella mediavia Harris. 
Natica saffordia? ** 
‘¢  reversa Whité. 
onusta a 
eminula Con. 
Limulany, Wes 
Amaura tombigbeensis Harris. 
Mesalia watsonensis Harris. 
‘(pumila Gabb (var. ) 
ae alabamiensis Whitt. 
Levifusus pagoda Heilpr. 
: dalei Harris. 
Fusus quercollis Farris. 
PN NOLLOMES EN Gite 
Coll coneeles NN ad he 
‘< —mohrt Aldr. 
Dentalium mediaviense Harris. 
Cadulus turgidus Mr. 
Atys robustordes Aldr. 
Tornatella quercollis Harris. 
Cylichna meyert Aldr. 
Gilbertia estellensis Aldr. 
Cypre@a 2 sp. 


a 


29 


27 


ALDRICH, HOCENE MOLLUSCA 


Exilia pergracilis Con. 
Nautilus sp. Fragments. 


Flabellum conoideum Vaughan. 
Balanophylla haleana M. Ed. & H. 
Stenocyathus n. sp. 


Foraminifera, 12 species. 


Figure 


W7h 
19, 


Zip 
23, 
25; 
27, 


~ 


Il. 


BULLETIN 37 


EXPLANATION OF PLATE 1 


28 


Page 
Mitramorpha turriculata, n. sp., Choctaw Cor, Length6mm... 5 
Turris MODOLACUSMMENSD AN Gave ranle bate iecen in eneenette ae gimm... 5 
TUIGUSESPECH STIS DM CAN CM OBTA TCH ewer sere eeaititatsasesee 8mm... 6 
Turris bimoniatus, n. sp., Cave Branch..........2.. sese..0 14mm... 7 
Pleurotoma ( Peratotoma) Gardneri, n. sp., Pugh’s Br. 21 mm... 8 
Odostomia (Evalea) Barischi, n. sp., Bell’s Ldg............ Bi youbodlane |) tS) 
Turbonilla (Ptycheulimella) clinensis, n. sp., Gregg’s 

A Bfelr aX 6 Vester aunt aA Hae aa aaa iat Apa mtn 3mm... 9 
Turbonilla (Cingulina) tuscahomensis, n. sp., Gregg’s 

I Brahals boa bea Waa Re aS MOUNTS Wn A aa eUO Ue Plame Pa HSIN a ar 4mm... 9 
Turbonilia ( Ptycheulimella) tardiusculus, nu. sp., Cave 

Ba ee a Shaan Wa LE alate aid sabia a Meet ona 4.5mm... 9 
Pyramidella ([phiana) obtusoides, n. sp., Gregg’s Ldg. 3.5mm... 10 
Epitontuim muntstriatum, n. sp., Jackson’s Rock House 

UB} of 010) a Maaienieai aM MU ant Ore RE cS a ERIE 4mm... 10 
Epitonium munistriatum, n. sp., Gregg’s Landing ..... 3mm... 10 
Epitonium multiliniferum, n, sp., Cave Branch......... iomm... II 
Epitonium subacutum, n. sp., Cave Branch...............- Io mm... II 
Teinostoma subangulata Mr., var. Smithi, n. var., 

Bell Sy arra dani patie auetun wp iauabamacuinsta at blu ca uionuiaualyan 8 mm... II 
Teinostoma regularis, n. sp., Pugh’s Branch..............+. 4mm... 12 
Anctllopsis Tuomeyi, n. sp., Bell’s Landing............... 28 mm... 12 
IN ASSO PICOM ENTS D Aly ISON Meni ceeee een ieee eae 12+ mm... 12 
Ohwvella semilienitica ims spy Clarkin Con ieee ison 5.5mm... 13 
Cancellaria Gilberti, n. sp., Gregg’s Landing...............9 mm... 13 
Ovula regularoides, n. sp., Bell’s Landing........... 2... 14mm... 14 


Sy; 


a 
a 
So 
> 


é @ % 
17 
Za 


Meets 
megs 


29 


ALDRICH, KOCENE MOLLUSCA 


PLATE 2 
(2) 


ey. 
T5, 


27; 
29, 


28. 
30. 


BULLETIN 37 


EXPLANATION OF PLATE 2 


30 


Page 
Gypr aa estellen sts. Ne Sp) \eursleya Chascsscceseecensetesres ce 12mm... 14 
@ypraaispybursley | Cree iece you eee enestececee ee sseeeeeee I5 mm... 15 
Siphonalia quadrilineata, nu. sp., Pugh’s Branch......... Gp scobadlinn 105) 
Gilbertia estellensis, n. sp., Estelle.................eeeeeeee 2.5x2 mm... 13 
Levibuccinum (Euryochetus ?) harrisi, n.sp., Cave 
Branch); largest specimens wscerse.c ses -ssasses io 4.5 mm... 16 
Tenutacteon pretenuis, n. gen. and sp., Gregg’s Ldg....13 mm... 16 
Gerithtopsts esicllensis. 1.) iSps. ee Stelle enn accneee seeseceeee 3mm... 15 
IRA AHO) (HISTALS oo VYSUAVS ooecaodoabbacnasooseoya, oaticased 37 mm... 17 
Tellina estellensis, nu. sp., Estelle......... largest sp....I5XII mm... 17 
Tellina bellsiana, n. sp., Bell’s Ldg.; height.............. 24mm... 17 
Tellina semirotunda, n. sp., Gregg’s Ldg. ; length......16 mm... 18 
Tellina semipapyria, n.sp., Jackson’s Rockhouse 
Branchy wen oth cove sssucesaveseecscer sone eeee ees 23 mm... 18 
Semele langdoniana, n. sp., Bell’s Ldg.. ..............060++ 27 mm... 18 
Semele monroensts, nu. sp., Bell’s Ldg...........ceeeeeeeeeee ee 5 mm... 19 
Lucina primoidea, n. sp., Black Bluff................006 9x9x6 mm... 19 
Martesia vecurva, n. sp., Fleming’s Mill, Nanafalia 
| OY e10 USAMA AL a eGR Sc acy 8x4.5 mm... 19 
RROCHE/OV UG IHINULAA NeSp (Bells ie daaauee ses ee eee ere eee 3mm... 20 
Montacuta bicuspidata, n. sp., Bell’s Ldg............... 2x234 mm... 20 


Vol. 9, Pl. 2 


Paleont. 


Amer. 


Bull. 


Pl. 2, No. 37 


7; 
MY 


Zi < 
Ky 
_ 


<7 
< 


7 Tis. : ) 
See OZ 
Coe 

7 


ui ae 
ria 


wy) 


cae 


wr 
ho Sy 
leat 


31 


ALDRICH, EOCENE MOLLUSCA 


PLATE 3 
(3) 


aI 


20. 


22. 


BULLETIN 37 32 
EXPLANATION OF PLATE 3 

Page 
Basterotia ? prima, n. sp., Gregg’s Ldg.................--- 8x7 mim... 20 
Sartcavella alabamensis, n. sp., Bell’s Ldg.......... 6x4.5 mim.... 21 
WIONLACULA ELCLOCH LIM SPs CAVE MD ae tae etre eens ee sass ee g mm... 21 
Eyvilraligniieca.) Te Spi Belly Siy i, digmees | ean eseicusin: senate 6mm... 21 
Fanopem bellsensts, 1. Spi Bells ide ys eee esa 30X13 mm... 22 
Donax acutangula, n. sp., Bells’? Ldg............ ee. 31XI5 mm... 22 
Arca (Cucullaria) ozarkeusis, nu. sp., near Ozark...... 5X3 mim... 22 

Ostrea intermedoides, n. sp., Bell’s Ldg. No. 15, 24x19 mm. 
No. 16, 32x22 mim... 23 
Psammobia Simitht, n: sp., Bells Wde. tes 60x32 mm... 23 
Psamunooia harrvist,)m.\sp., Gregers dere ce sale cesens 37 mm... 24 
(Echinocyamus ?) meyer, n. sp., Claiborne...... 5X342x3 mm... 24 


Vol. 9, PI. 3 


Paleont. 


Amer. 


Bull. 


Pl. 3, No. 37 


Ce 
<— 


Bo 


te 


a 


hoe ; 


ce 
nes 


BULLETINS 


OF 


AMERICAN PALEONTOLOGY 


soy TE 


Vol 9 


sk 


No 38 
RECENT MOLLUSCA OF THE GULF OF MEXICO AND 
PLEISTOCENE AND PLIOCENE SPECIES FROM 
THE GULF STATES 


Part 2 
Scaphopoda, Gastropoda, Amphineura, Cephalopoda 
BG: 


CARLOTTA JOAQUINA MAuRY 


January 31, 1922 
[Part I, PELECYPODA is Bulletin No. 34, 1920. ] 


Harris Co. 
Cornell University, Ithaca, N. Y. 
Wi SeAw 


i 
My) 


a 


A MOLLUSCA OF THE GULF OF MEXICO 24 


GWASS V/SGNAai@  @ibDA 
ORDER SOLENOCONCHA 
Genus DENTALIUM Linnzeus 


laqueatum Verrill, Trans, Conn., Acad, Arts Sci., vol. 6. pt. 2, 
Pas pl 44. fi 18) 18857 Walli Bull} Musi Comp. Zool: 
EMO D AZO DI ts Tresor) Bill ara WLS A Wine. 
GIS ypu TOM ply 27.) fn pl AG. li Tey) T8SOr.bilsahyiand 
Shanp, yinvyon s Man. Conch.,.17) ps ro, (pli 70 fa\ta2-) pluis) 
Peso?) tlenderson,, Bulli ci0.2U. Ss. Nat. Maas. p. 23; 
PAT te Oni'7,), 1O2O. 


Distribution—Hatteras to Barbados, Gulf of Mexico: West 
Florida and off the Mississippi delta, 60-68 fms. 


texasianum Philippi, Zeitschr. Malak, p. 144, 1848; Pilsbry & 
Sharp, Tryon’s Man. Conch., 17, p. 22, 1897; Henderson, 
cr OR SaNe VE 1.) 28 nile Enea 5 Oe, LO2O. 


sexangulare Hilgard, House of Rep. Ex. Doc. 1, pt. 2, pp. 
37,1889, pl. 3, f. 7, 1878. Not of Lamarck: 


gouldiz Singley, 4th Ann. Rept. Texas Surv., p. 331, 1892; 
Dall, List Cameron Sh. Not the true gould7? of Dall. 
Note.—This may be identical with Chenu’s 2. americanum 
1843. Closely related to gouldzz Dall, with which it has 
been confused. 


Distribution.— N.Carolina to Yucatan, 3-9 fms. Gulf coast: 
Point au Fer and Cameron, La.; Galveston, Corpus Christi 
and Pass Cabello, Tex. Pleistocene, Lake Borgne bor- 
ings, La. 

texasianum cestum Henderson, Bull, 111, U.S. N.M., p. 29, 
DIN Ark MAA. 8) (O20: 


Distribution.—Pass Cabello, Texas. Probably in very 
shallow water. 


rebeccaensis Henderson, Bull. r11, U. S. N. M. pp. 16, 31,pl. 3, 
LN TO2@) 


5 BULLETIN 38 35 


Distribution.—West Florida, 7-16 fms. and the Tortugas. 


(Anialis) antilarum d’Orbigny, In de la Sagra’s, Hist Pol. y 
Natyisla deliCubal 2iipszo2t plist Ota reroll site 
& Sharp, Diyouis) Man. Conch) 17). 57, ple) 14) f 22-25) 
seven Jealssavaleqsropal,| JEU Wessse VOL, Sy ONY VES! oy Au jolly. 96, t= 
4, 6-8, 1920. 
Note.—Confined to shallow water. Deep water citations are 
reterable to different species. 


Distribution.—Florida to Barbados. Gulf Coast: Charlotte 
Harbor and other stations off West Florida, 2-14 fms. 


(Antalis) pseudohexagonum Ihering. See Henderson Bull. 111, 
Ue SNe Mee py oy ple Gu tet- 2 192e) 
Distribution.—Florida to Brazil. Gulf coast: Marco, Tampa 
and Charlotte Harbor, West Florida, 2-5 fms. 


(Antalis) disparile d’ Orbigny, Hist. Pol. y Nat. Isla de Cuba, 
2. Puzo plicit lati lo4o: (ci Vianattay) Proc mncade 
Ne SiPhilaissi) pe7isi741 19033 taenderson, Bulle now WUEioe 
ING Meo ae pln On hac on i O20: 


Note.— Frequently confused with D. aztillarum by authors 
althougn really closer to dexasianum. 


Distribution.—Florida to Rio de Janeiro, Brazil. Gulf 
coast; Tampa, Fla. Littoral zone. 


(Antalis) ceratum Dall, Bull. Mus. Comp. Zool., 9, p. 38, 1881; 
TOGO US. 0, MAM OL AG). Wel I ONL, \ Ag) sts yi atshexoy Dowoudly a), U). 
SNE ME pe On pl 20 Ss TOSOn me enderson is allay mune 
Sa) NE) Meipper49-514) ploy. hf. 227, 1920: 

Distribution—Florida to Barbados. Gulf of Mexico: Off 
Mississippi delta, 321 fms.; Yucatan Banks, 84 fms. 

(Antalis) taphrium Dall, Bull. M. C. Z., 18, p. 422, 1889; U.S. 
N.M.,p. 76, 1889; Pilsbry & Sharp, Tryon’s Man. 17, p. 58, 
L8O7 lenderson Bulltaian Ue SaiNEwver Nps 260 pluie ies 
1920. 


36 MOLLUSCA OF THE GULF OF MEXICO 6 


Distribution—Hatteras to Cuba, 22-182 fms. Type dredged 
between Mississippi delta and Cedar Keys in 30 fms. 


(Antalis) bartietti Henderson, Bull. 111, U.S. N. M., p. 55, pl. 
CMH hy LOZO: 


rmonliagine Dall Bulle Me ICA eZ Ni'o.)paig7s) USSt + Bullaia 7. 
eS SINE MO Dp, 76) (pars), “1889.) Not) anilarum)) a Or- 
bigny, 1846. 


Note.—True antidllarum is semi-littoral while déarflett7 is in 
deeper, colder waters. 


Distribution.—Florida to Martinique. Gulf of Mexico: Be- 
tween Mississippi delta and Cedar Keys, 227 fms.; Yucatan 
Banks, 399 fms. 


(Heteroschisma) cailithrix Dall, Bull. M. C. Z., 18, p. 427, pl. 27, 
LOMO CO MP oUluia 7. Sa Na Me pio, plea 7Mit:) TOmisse: 
ilsbryce Suarp, Uryons Many Conchin7)) p62, ply 7, f.13; 
Noone Elenderson, Bully 111) Uo S4) Ne Mi p57, plewsa fs 3, 
dks On) MOO’ 


Distribution—North Carolina to off Rio de la Plata, Ar- 
gentina. Gulf of Mexico: Between Mississippi delta and 
Cedar Keys, 1181 fms.; Yucatan Strait, 640 fms. (type lo- 
cality). 


(Fissidentalium) meridionale Pilsbry and Sharp, Tryon’s Man. 
Conehy17) 99.73, ple15, £. 32-34) 1897, \(as variety of ca7z= 
didum): Henderson, Bull. 111, U. S. N. M., p. 61, pl. 9, f. 
2TO2O: 


Lona sep alle alle NE KOCH) Z-6,) 9: 37.) kool iLO Ps) 4:25.00) LOOOs 
Proc. U. S. N. M. 12, p. 294, 1889. Not ceras Watson. 


Distribution—Type dredged by the Albatross, 240 miles south- 
east of Rio de Janeiro, 671 fms., 37° temperature bottom. 
Also found Yucatan Banks, 1568 fms. and off Barbados. 
The largest Dentalium of the western Atlantic and charac- 
terized by its unique, ashen color. 


7 BULLETIN 38 37 


(Grapiacme) eboreum Conrad, Proc. Acad. N.S. Phila., 3, p._ 

27, 1846; Pilsbry. and Sharp, Tryon’s Man. Conch., 17, p. 
89) pl: 16,\f.)'47-405)) 55, 56) 1897,;Ekenderson))) Ball) irae 
NGM ph OO. pl MO NEM sH5 4) SiO mip lamin canon ZO 

leptin) Bush ihrans, Conny Acad aw ArtSsGandsS cla wisere nite 
CD NAT OmpIUA SMe Sh Tea iT SS Seii Daly tele aa an Ole 
NMP pO LS! 

matara Mal OB ME COZ 3h pa AZO TS SO. 1 Ellas 
WS ceN MOMS So) 


Distribution.— Hatteras to Barbados. Gulf coast: Sarasota 
Pass (type locality), Charlotte Harbor, Fla. and between 
Mississippi delta and Cedar Keys. Beach to 111 fms. 


Note.—Mr. Henderson unites matara and Jleptum with 
eborcum, noting that szafara is the perfectly smooth adult 
form of the species. 


(Grapiacme) semistriolatum Guilding, Trans. Linn. Soc. Lon- 
doniin7 pty ips s4vplagi tet os read Chemin Comehemldile 
De Ty pia i iain Set itendersons, Hull mre Opn Sener 
Me piOG yp lenin stank 54S MO n O2@: 


Distribution.— An abundant littoral species. Florida to 
Barbados. Gulf coast; West Florida in shallow water, 
7-12 fms. 


(Graptacme) sericatum Dall, Bull. M. C. Z., 9, p. 37, 1881; 18, 
DV AZZ DL ZO MTs son a dll ary MONS. ING IMs: i mii oruple 
2On eT SSO. VElendersony ME mile MNT Oe IN an Veep Deals 
PMI TA OM O2Or 
Distribution.—Type only found. Dredged by the Alake, 640 
fms., Yucatan Banks. 


(Laevidentalium) callipeplum Dall, Bull. M. C. Z., 18. p. p. 410, 
OLN AAS Saaceon = CMa ifereayanies) IBY EVOL) ak AUS AN SI INI dy bas Ob 6) Ol! 2347 
fyi 2'O) Lacon lrans: | \Viaoner wimSt Cia 2) \p. (442 1sQoR 
Pilsbry and Sharp, Tryon’s Man. Conch., 17, p. 100, pl. 19, 


38 MOLLUSCA OF THE GULF OF MEXICO 8 


MPOMnso7)) Henderson) Bullen) (Ws) S\N Mpa 4. pls) 12), 
“its (Gy) OPee 

Distribution.—Florida to Porto Rico. Gulf of Mexico, 

dredged to 169 fms. between Mississippi delta and Cedar 

Keys, (type locality). 


(Laevidentalium) perlongum Dall, Bull., M. C. Z., 5, p. 61, 1878, 
Women nUudure) Gy 9) 26.1883) 18). Anos pler27 fs, 6, 
MoCo Bulls7 US. Ney Mes ps6) pla 27.) On Taso blen= 
derson Bull ir1r US) N2 Meio 75.0 pk. (o,f. 1) 1920: 


Distribution.—Hatteras to Rio de la Plata, 11—1330 fms. 
Gulf of Mexico between Mississippi delta and Cedar Keys 
at 940 and 1181 fms. also elsewhere in the Gulf at 1330 
fms. 


(Episiphon) sowerbyi Guilding, Trans. Linn. Soc. London, 17 
pass fy 7a T8345) Pilsbry and) Sharp,)nyonrs, Man,) 
Counc ptr 7.) pl. 20, f..2om1so7> blenderson)) Bull. 111 
We SHON VE pps 77-80, Diya ity 2a LOwTO2@: 

Za Dally VB ww 37 Ue Si Ne ME pal 76 T88O.) | Pro: 
bably not f/um of Sowerby, 1860, which was from the east 
Atlantic. 


Distribution.—Hatteras to Barbados. Gulf of Mexico: Off 
Mobile, Ala., 27 fms.; between Mississippi delta and Cedar 
Keys, 60 fms. 

sowerbyi pelliceri Henderson, Bull. 111, U. S. N. M., p. 80, 
IE NEL 7S) Out O2 0: 
Distribution.— Florida to Cuba, Cape Catouche, Yucatan, 
25 fms. 


(Bathoxiphus) ensiculus Jeffreys, Ann. and Mag. Nat. Hist. ser. 
ERO MOU SAN LS 7h: Me hOC. LOOM SOC PE OOOW plan 4Qute0 4); 
1882. Watson, Challenger Rept., (Scaphopoda), p. 12, pl, 
Zeb 2 Sos. LOall: (Bully 370 Ul Sa Ne Me py 7On18 36. 
lenderson, Bullieriny Ws oS: Ney Vi pa Stanplanie iat 14.5, 
TOW LQ2O: 


9 BULLETIN 38 39 


Distribution.—East and West Atlantic. A deep.water 
species of wide range. Gulf of Mexico: Yucatan Strait, 
640 fms. 

(Bathoxiphus) didymum Watson, Jour. Linn. Soc. London, 14, p. 
517, 1879; Henderson, Bull) 11.0. S. NM. p. 83) spline 


10, MOKA, 


Distribution.--Culebra Isl. and Barbabos. Doubtful specimen, 
Yucatan Banks, 25 fms. 


(Compressidens) pressum Pilsbry and Sharp, Tryon’s Man. Conch. 
TiO w 2A Ole mut. MIMD 22a NGOs 2h ed SO). Wlendersone 
Bull nee NG Mi ons Zep lealaaien ss soo, sLO2O. 


compressum Watson, Jour. Linn. Soc. London, 14, p. 516, 1879; 
DallsyBullwMe C7 Zo passa rons Watson, Challe kept 
(ScaphQeip: 0, pla at] Guinges malls ull 27. US. Niner 
p. 76, 1889. Not compressum of d’Orbigny, 1850. 


Distribution.—Florida to the Antilles. Gulf of Mexico: 
Off Cedar Keys, 196 fms.; between Mississippi delta and 
Cedar Keys, Fla. 


(Compressidens) ophiodon Dall, Bull. M. C. Z., 9, p. 38, 1881; 18, 
D427. pl 26) OE S8o; MBLIllNe TORS. UN Melt oe 7G ple 
ZO tO 1889" Itenderson,, Bulle minim Us So. NY) Met) Saki: 
WA ATOZ @. 


Distribution.—Florida to Barbados. Gulf of Mexico: West 
Florida, 169 fms. 


quadrangulare Harris, Bull. Amer. Paleontology, vol. 1, p. 95, 
1895. Upper Miocene, Galveston well at 2465-2871 feet 
(Harris). 

tetragonum Sowerby, Thesaurus Conch., 3, p. 103, pl. 224, f. 
21, 22, 1860. Upper Miocene, Galveston well, at 2158- 
2920 feet (Harris). 

Remark.—A fragment of an undetermined, Pleistocene, Den- 
talium was obtained from Knapp’s No. 2 well, Terrebonne 
Parish, La. at 1190-1430 feet. 


A0 MoLLUSCA OF THE GULF OF MEXICO 10 


Genus CADULUS Philippi 


(Polyschides) quadridentatus Dall, Bull. M. C. Z. 9, p. 36, 1881; 
SOM AZo Plit2 7 i 5 rssory BULLEN a 7U) 1) SIN Mp.) 76; 
Plea Tat. Su 889) lenderson, Bull) iin Wh) SNe) MES pp 97- 
TOOM MNT 7a the 221) LOZ! 

toms bush. Vln: Conn. Acad.,\6) p. 471.) pl 45, f, 20) 18855 

Dalal sree Sy NW IME pkigGh plo An. st) 201 8 5o 
Distribution.—Hatteras to Rio de la Plata, 3-52, fms. Gulf 
of Mexico: West Florida, 10-25 fms.; Yucatan Banks 25 
fms. 


(Poiyschides) carolinensis Bush. In Verrill, Rept. Fish Com. (for 
LISA PSST LSo54 Li Conny Acad’) 6.0 p. 47) Pla 5) be 
Momissse Dall Bull: 27 Ua SuiNe Mp. (48 (pars). ply au, 
f. 19, 1889; Pilsbry and Sharp, Tryon’s Man. Conch., 17, p. 
m2 pl. 25, 4. 64) pli 30, f. 241-127), 1898; Henderson, Bull! 
ORO EL SMN EUV) Os 102) Ply mhz Mt eNO) LOZO} 
Distribution.—Hatteras to Key West, 3-87 fms. Gulf of 
Mexico, off Mobile, in 27 fms. 

(Platyschides) poculum Dall, Bull., M. C. Z,, 18, p. 429, 1889; 
Ble 7 Oo. Ne MES pp) 765 1889 5) Pilsbry and) sharp; 
direyonys Vian. Conch 17. p41 72, Dleisgeten 50, 57) L398) 5 
itenderson, Bulli, U.S: Ni Mey py 10s pli nz, 1.18, 1920. 
Distribution.— Type found off St. Vincent, W. I., 424 fms. 
Aso dredged, 640 fms., Yucatan Banks. 

(Platyschides) parvus Henderson, Bull. 111, U.S.N. M., p. 113, 
OTS hor 52.04) LO20. 

amtantus Dall, Bull. 37, U, S. N. M., p. 78, 1889. Net Dall’s 
typical amzantus. 
Distribution.— Off Barbados, too fms. (type); Yucatan 
Banks, 1002 fms. 

(Platyschides) elongatus Henderson, Bull. 111, U.S. N. M., p. 
LAB. ‘solo WO, sey sy sO VAOY 


II BULLETIN 38 4r 


Distribution.—Gulf of Mexico, off the mouth of the Missis- 
sippi River, 68 fms. 
(Platyschides) arctus Henderson, Bull. 111, U. S. N. M., p. 124, 
DILLON LO MmmOZOs 
Distribution.—Gulf of Mexico off Cape San Blas, Florida, 
60 fms. 
(Gadila) mayori Henderson, Bull. 111, U. S. N. M., p. 133, pl. 
LOW Lao LOZ) 
Distribution.—Florida, Gulf of Mexico, off Cape San Blas, 
Fla., 60 fms. 
(Cadulus) transitorius Henderson, Bull. m1, p. 143, pl. 19, f. 6, 
1920. 
Distribution.—Florida to Old Providence Isl.,north of Colon. 
Gulf of Mexico, off Cape San Blas, Fla., 196 and 169 fms. 


42 MOLLUSCA OF THE GULF OF MEXICO [2 


GOMASS GINS @ 7 @ aX 
ORDER PTEROPODA 


Genus CAVOLINA Abiidgaard 


inflexa Lesueur Nouv. Bull. Soc. Philom., 3, p. 285, pl. 5, f. 3, 
Iok(aseeyalca). Dalit Bulle We S Nati Mus.) (37, p. 82, 
LOCO ETOC. Ueno. Nate MUS i 137.\\0.) 1.7) LOIO: 
Distribution.—West Atlantic, Lat. 42°N.—42° S., Hast 
Pacific, Lat. 13° N.—42° S. Gulf of Mexico, West Florida. 
Pelagic. Pleistocene to Recent. 

uncinata Rang, in d’Orbigny’s, Voy. Ame’r Me'’r., p. 93, pl. 5, 
MEME TS 26.8 (as Eyalea) Dalley Balle 27 WU AS: Ne Mars, 
MSZ Mlle G6, hy) 1lO} 18803) brocwU) SN Mius: 375 pa t98, 
1910; Johnson, Occ. Papers, Boston Soc. Nat. Hist., 7, p. 
156, 1915. 

Distribution.—West Atlantic, Lat. 40°N.-40°S. East Pacific, 
Lower California to Ecuador and Galapagos Isls. Gulf of 
Mexico; West Florida. Pelagic. Pleistocene to Recent. 


(Diacria) trispinosa Lesueur, in Blainville; Dict. des Sciences Nat. 
227 82, 1821, (as Ayalea); (Gould) inv. Mass.,) p: 504, 
ms7ou(as Yzacra); Wall, Bull. 375% pi'82, pl. 66) f)115,, 1889, 
Sumner, Bull. Bur. Fish., 31, pt. 2, p. 703, 1913. Johnson, 
Oces Papalbosk. SOC47,) DP: 15 7, LORS: 


Distribution.—West Atlantic, Lat. 60°N.—4o° S. Gulf of 
Mexico, West Florida. Pelagic. Pleistocene to Recent. 


Genus CLIO Linnceus 


pyramidata Linnzeus, Systema Nature, 12th ed., p. 1094, 1767; 
Dalle Bulbe7. Wie SevNG MN peahsonmTssoy. (as Clcodora): 
Broce Se Ne Mea 7p. 198) TOLOwmjonnson: (Ocei rap: 
BOSH SOCK 7a Dulisou LOLS. 

Cleodora lanceolata Souleyet, Zool. Bonite, 2, p. 179, pl. 6, f. 
17-25, 1852. 
Distrib ution.—Atlan tic, Spitzbergen to Lat. 40° S. South- 


3 BULLETIN 38 43 


eastern Pacific, Lat. 27° S. Gulf of Mexico, West Florida 
and Texas. Pelagic. Pleistocene to Recent. 


recunva) Children,’ Jour//Roye\lust.;) 15). 220, pli, fa 107.) 1820) 
(as Balantium); Dall, Bull. 37, p. 82, 1889; Johnson, Occ- 
Papi BOsts! SOC 7 i ps lon LONss 


Distribution.—Lat. 40° N.—33° S. Gulf of Mexico, West 
Florida. Pleistocene to Recent. 


Genus CRESEIS (Rang) Sowerby 


subula Quoy and Gaimard, Ann. des Sci. Nat., ser. I, 10, p. 233, 
pls Dy hea ins274 \(asi@/eocoxa) Walla Balle 27. paoos 
1889; Dall and Simpson, Bull. U. S. Fish Com. for 1900, 1, 
Dp: 360, (plNS 7) fo LOOP DallivenocwWES. INEM 37s. 
LQLO) Johnson. Occ) Lap.) Bost Soc 7.) p- 158s LOLS) (as 
S7yleola) Dall Ball.) 12) Wey SeeNataiMiidsy.0p4) 5O.y1o2ie 


Distribution.—Atlantic, Lat. 41° N.—40° S. Type locality, 
Teneriffe. Pacific, Peruvian. Gulf of Mexico, West 
Florida and Texas. Pelagic. Pleistocene to Recent. 


Genus STYLIOLA (Lesueur) Gray 


virgula Rang, Ann. Sci. Nat., ser. 1, 13, p. 316, f. 2, 1828, (as 
Greseis): Verrill, Trans. (Conn: “Acad.,5, -p. 557, 1882, (as 
styizola) Dall) Bull iag Ue SssuNe oe MiuS.. )\ 1). 80.) 7 Sonntas 
Gresezs) (Proc. US. INE MES 27ers 18, LOLO)(as”.Sty/zola))e 
Johnson, Occ. Pap. Bost. Soc., 7, p. 157, 1915, (as Cvesezs). 


Distribution.— Atlantic, Lat. 41°N. - 35° S. Pacific, off Juan 
Fernandez Isl. Gulf of Mexico, West Florida, Pelagic. 
Pleistocene to Recent. 


Genus VAGINELLA Daudin 
chipolana Dall, Trans. Wagner Inst. Sci.,3, pt. 2, p. 431, pl. 23, f. 
4, 5, 1892. 


Lower Miocene, Chipola beds, Bailey’s Ferry, Calhoun Co., 
Florida. An ancient, Cretaceous to Recent, genus. 


44 MOLLUSCA OF THE GULF OF MEXICO 14 


Genus CUVIERINA Boas 


columnelia Rang, Ann. des Sci. Nat., ser. 1, v. 12, p. 323, 1827, 
Atlas 4°, pl. 45 B, f. 1-8. (as Cuvierta); Pelseneer, Challen- 
gen) Eteropoda, Zool.,.22 art.) 1, p. 67, Teese; Dall Buall.27, Ul: 
Soe Marsh) 182) i ply 66s) faint Son na Waser inst: 
Sew synpte 2, ps 4ar i T892s Peck. Proc) U's) Nii Mus.) 16; 
PAAOe Less ty ito. re9ge! Dall Proc.) War Su Na Mie 7) 1: 
KOS) TOLO; Jolson) Occ Pap. Bost..S0C!)) 7p. L595 LOLS: 


Triptera columnella Verrill, Trans. Conn. Acad., 5, p. 557, 
L382) 62 py 2iA, 1884: 


Distribution.—West Atlantic, Lat. 43° N.—4o° 5. East 
Pacific, Lat. 23° N.—42° S. Gulf of Mexico, West Florida 
and Texas, recent. Lower Miocene Chipola bed at Alum 
Bluff, Calhoun Co. Florida. 


/ 


Note.—The specific name is often, but erroneously, written 
columella. "The correct form is colummnella since Rang prints 
it thus twice in the original description. The generic name 
Cuvieria is preoccupied. 


Genus EMBOLUS Jeffreys 


inflatus d’Orbigny, Voy. Ame’r. Me’r., p. 174, pl. 12, f. 16-19, 
HSZ6) (asdvanta)- Wall, Bull) 27 UK SaiN) Musi p. 80, 
MSOOMETOC Wis. Ni MEWS: 27) pa lOom Oro; ounson,)|Oce: 
Rap BOSt. SOC.) 7.) p) 155, 1915. (as “2zveacia)). 


Spirialts rostralis Verrill, Trans, Conn. Acad., 6, p. 431, 1885. 


Distribution.—West Atlantic, Lat. 42° N.—40°S. East 
Pacific. Lat. 42° N.—40° S. Gulf of Mexico, West Florida. 
Pelagic, Pleistocene to Recent. 


Genus PERACLE Forbes 
reticulata d’Orbigny, Voy. Ame’r, Me’r., p. 178, pl. 12, f. 32- 


C0 Jie 


BSsO, oso. (asia Hana) Dally Bulle 27a UL Sa Ne Muss 
PMCs sO roe Weis.) Ne MUS! Nam Ds nO LOLO: 


15 BULLETIN 38 45 


Distribution.—West Atlantic, Lat. 37° N.—o° S. East 
Pacific, Lat. 20° S. Mediterranean. Gulf of Mexico, West 
Florida. Pelagic. Pleistocene to Recent. 


ORDER OPISTHOBRANCHIATA 
Genus ACTAESN Montfort 


punctostriatus C. B. Adams, Jour. Bost. Soc. Nat. Hist., 3, p. 
3235 pli 3; £70; 11840, \as) Lozvatela)\ i Dally uli) 274 WER. 
ING Mus: ((p. 7840) ipl. (41 is A ols 2a 22 Sori dlbiey NVialon er 
Inst. Sci., 3. p. 14, 90; Singley, 4th Ann Rept. Geol. Surv. 
Texas, p.)332, 92; Pilsbry,) ainyon’s) Man.) Conch! sem im 
TS) DN 157, Dl Tet OS, OR plano 22" 22 0 co2.\)Dalilvannad 
Simpson, Bull: U.S. Fish Come an). 262)! 1901. Johnsons 
OcePapers) Bost) Soc (Ne Lie omar) NOS: 


punctatus d’Orbigny, In La Sagra, Hist. Pol. y Nat. Isla de 
Cuba, 1, p. 230, Atlas, pl. 17, f. 10-12, 1842; (as Zornatella): 
Dall Proc. US) Ne Me Ole pa e247 r88iae Not 1a pacar 
Ferussac, 1823. Not of Lea, 1833. Not of Piette, 1855. 


cubensis Gabb, Tr. Amer. Phil, Soc., 15, p. 245, 1873. 


Note.—The recent Cuban Acteon punctatus d’Orbigny 
(preoccupied, renamed cubenszs by Gabb) is now considered 
identical with the northern punctostriatus. Act@on punc- 
tatus Wea) (Contr (Geoltio. jiammiaply 4st. (96) 1s), Claiborue 
Eocene, Ala.; and, fide Dall, Pliocene; Caloosahatchie beds, 
Fla. Not Recent. 


Distribution.—Cape Cod to the Antilles, 2-63 fms. Miocene 
(of Shiloh, New Jersey) to Recent. Gulf coast: Cedar Keys, 
Fla,, Corpus Christi Bay, Tex. Pliocene, Caloosahatchie 
beds, Fla. 

exilis Jeffreys, Ann. & Mag. Nat. Hist., 4th, ser., 6, p. 85, 1870; 
Watson, Chall. Rept. Gastr., p. 624, ’86; Dall, Bull. M. C. 
Lp wharve Coll i 8i ipa gon .SOsmotillen a7 Onna No Mie anon 


46 MOLLUSCA OF THE GULF OF MEXICO 16 


B3o Meilspigy, Mane Conch 1 Ser, 154 pe 156) ply to, 1! 4-6,’ 93. 


Punic a cescuipra | Verrmill:| Proc.) W..5) N, M5 32," p. 381, 
1880. 


wavdus Nernst hr. Connl Acad), 5, p..540) pl. 58) f..21, 1882. 


Distribution.— West Atlantic, Martha’s Vineyard, Mass. to 
Florida. Also Mediterranean and Bay of Biscay. 92-1465 
fms!) (Gulf of Mexico; -Campeche Bank, at ' 200° fms. 
Pliocene to Recent. 


pusillus Forbes, Rept. Brit. Assoc. Adv. Sci., p. 191, 1843, (as 
Tornatella); Jeffreys, Ann. & Mag. N.H.,4th ser., 6, p. 84, 
5th ser. 10, p, 34; Watson, Chall. Rept. Gastr. p. 627; Dall, 
Bulls a7. Ws S.No M., p. 84, 1889: Pilsbry, Man? Conch: ; 
TSC NSHP 5ONI OOS: 
An unfigured species. 
Distribution.— Mediterranean, Madeira; West Indies. 20- 
450 fms. Gulf of Mexico, off Sand Key, West Florida, at 
eT istaNS 

perforatus Dall, Bull. M. C. Z., 9, p. 96, 1881; 18, p. 42, pl. 18, 
Peso bull 27 Ua oo) NaN DN nS 4a ula TSM iit 35.089) 
Bilobiyw Man Conch. 1 Ser. 5) pel 5Om ple 204 ta 2On ESO" 
Distribution.—Gulf of Mexico, 805 fms. 


McIsSUS Dall NEC 7G .i Ds 05.) SSI kon puraze lei toute TO: 
POOP BUN a TA UR e SHON). Men sp. San Olen 7a fil TOs 218.0): 
il Sbinyag Mani Conehis 1 Sets) 15. pa Loluinog” 


Distribution.—Yucatan Strait, off Cape San Antonio, 640 
fms. Also off Fernandina, N. EF. Fla., 294 fms. 


delicatus Dall. Bull. M. C. Z., 18, p. 41, pl. 17, f. 5, 1889: Proc. 
We Se Ne METZ spy 296, /8o 7. Bullet anne SUNG Ma hoc 7Sae 
plate peins 62S.) balsbtaya Mans Conch aisers 1.154) p41 102.-pl: 
DO, ty Bin, OB 
Distribution.—Gulf of Mexico, West Florida, region. Also 
Barbabos 73-400 fms. 


17 BULLETIN 35 47 


galvestonensis Harris, Bull. Amer. Pal., vol. 1, p. 96, pl. 3, f. 1, 
1895. 
Distribution—Upper Miocene, Galveston well at 2552-287: 
feet. 
Genus RNIGICULA Deshayes 


RitidaVeralh, Am, our Seu auisen i) 5 pu Lows 730 br. Cons 
INCAd Ms DAS wlan Tuan tact Dalla olleeNten Cp Zi nS aioe 
AB WSO Bite aan Da Ott MDl Marte norns @ qi Ens otivgy (UMM taltay 
Conchiy ni sen sy is5..p- 290.) ple downs 36 1O3 +) OLSON. Oces 
Pay eBOSt SOC INIa UL a0) Tio) i sliy | OMe 


leplockevla Brugnone,) Misc! Male ipuiriiuipls)) Py it. 073) L87se 
Morelet wound de (Conch) \ppuimerm28 5. pla ssa tem7 aor 
Agassiz, Three Cruises of the Blake, 2, p. 70, f. 291, 1888. 


Distribution.—Massachussetts to Pernanibuco, Brazil. Also 
Pliocene of Italy (Brugnone) and recent in the Mediter- 
ranean. 19-1075 fms. Gulf of Mexico, Yucatan Strait, 
640 fms. 


Genus ACTEOCINA Gray 
bullata Kiener, Sp. et Icon. Cog. Viv., p. 5, pl. 1, f. 4, (as 
Toratela) Wall Bulli 7 Onis Ne WME SA o oor 
Palsbirya) Mans (Concha Ser nuis 4) pus 30) ple 50, bt.) 301 pluj22. 
fa i7- tol (Os) iblarnis Bull yAcne mals ol it; ps OOumOsE 
Dall sr Simpsoni Bully Sakisky (Com) 1. p. 362, plisyn 
LOM uOO! 
Bulla canaliculata d Orbigny, Hist. Pol. y. Nat. Isla de Cuba, 
HpsLsee pla) Diswei2ic24u Noto Say. 
Tornatina olivula A. Adams, Thes. Conch., 2, p. 569, pl. 121, 
an gat 
Distribution.—Florida Strait to Trinidad Island. Upper 
Miocene, Galveston well at 2410-2425 ft. (Harris). 


recta d’Orbigny, In Sagra’s Hist. Pol. y. Nat. Isla de Cuba, 1, 
DIU tpl 4) bis) t4 Mazo m(aswauiia) WlOall yy Ball a7) UNS. 


48 


MoLLUSCA OF THE GULF OF MEXICO 13 


NGNME p84) 18897 (Pilsbry, Man) Conchvtiser,15,/p. 184, 
pl. 22, f. 13-15, ’93, (as Zornatina); Maury, Bull. Amer. 
PANO 20) pital plalai ty 2hTol7: 


Distribution.— Florida Keys, Antilles and St. Helena. Gulf 
coast, Tampa. Also Lower Miocene Santo Domingo and 
Jamaica. A closely related, Older Pliocene, species is 
Acteocina wetherilli Lea, from Deal, N. J. and the Orbitolite 
bed, Tampa, Fla. 


canaliculata Say. Jour.. Acad. Nat. Sci. Phila., 5, p. 211, 1822, 


(as Volvaria); Holmes, Post. Pl. Foss. S. Car., p. 78, pl. 12, 
Mme iian Gouldw Binney) vedu nips 259 401115 kw S704 CaS 
Werculus) + Wally Proc: Us SiiiNaiMe) Os p. 324.) S35 Bulle 37, 
WM Sa Ni Ives woS4) pli SohteM 27 Sonia (asl onmarz7a)): 
Singley, 4th Ann. Rept. Geol. Tex., p. 332, 92; Pilsbry, 
Mane Conch iiisene 15) ipl o4epI M22 ib 2 2M) ple 5 Ousn2'5, 
Pow og.) Johnson, Occ.) Pap: Bost oo0ci) Ni En (70. 247, 
Honorary Vall) Am, PaliineNow 2oqn pila.) Plush.) 2), 
IQI7. 


obstricta Gould, Silliman’s Jour. Sci. 38, p. 196, 1840, (as 


Bulla wAdams.))\ Hes), 2.) pae15OOut 2h.) 200), (as 
Tornatina). 


Distribution.—Portland, Maine to Haiti, 0-63 fms. Lower 
Miocene (Santo Domingo) to Recent. Gulf coast.—Recent: 
Cameron and Point au Fer, La., (very abundant); Galves- 
ton and Corpus Christi, Tex. Pleistocene: North Creek, 
Manatee and Labelle, Fla.; Grand Chenier, La , Knapp’s 
wells, Terrebonne Parish, No. 2 at 1050-1790 ft., No. 3 at 
258-1525 ft. Pliocene; Caloosahatchie River, Fla. Miocene; 
Gilbert well No. 10, Bateson, Tex., 323 ft. 


candei d’Orbigny, Moll. Cuba, 1, p. 128, pl. 4, f. 1-4, (as Bulla); 


Daal a 700) SING Me pa Saem Olas fs (13.339), 
Vanatta, Pr. A. N.S. Phila., 55, p- 757, 1903. 


Distribution. Hatteras to Martinique, o-48 fms. Gulf 


1g BULLETIN 38 49 


coast, St. Joseph’s Bay and Crooked Isl., West Fla. 


Genus RETUSA Brown 


frielei Dall; Bull. M. C. Z., 9, p. tor, 188%, (as Uirzculus): 18, 
Diaz pl 175) te 4s (SOs MOM STA MOE SisilNis av leis Omastoleunlicge 
tia.) 89°) Pilsbry., Man. Conchi (a isera Wins saps 2uO nla inte 
8, 1893. 

Note.—I follow Cossmann in placing Ufricudus Brown, 
1827 (not of Schumacher, 1817) in the synonymy of 
Retusa. 


Distribution.—Yucatan Strait, 640 fms. 


Genus SCAPHANDER Montfort 


punctostriatus Mighels, Pr. Bost. Soc. N. H., 1, p. 49, 1841, (as 
Bulla. Wall, Ball 127.) Ula S Nee Mees Op ba 2 sete Aan Ors 
Pilsbry Man: Conch.) 1isereng ep 246 (aol hah ommOes 
Johnson, Occ! Pa. Bost: S: Nee aie pai4on Doms. 


librarius Love’n, Index Moll. Scand., p. 10, 1846. 


Distribution.—Iceland & Norway to Bay of Biscay. Maine 
to Barbados, 46-1467 fms. Gulf of Mexico, Lat. 28° N., 
Lon. 88° W., 533 fms. Inhabits shallow water in its north- 
ern habitats. 


watsoni Dall, Bull) MC. Zo); pgonimSSir aime p52 ,iyole elie 
for 78o.0 Bull 271) Wi. Su Nee Mes py SONI aie mons, 
Distribution.—Hatteras to Barbados, 54-324 fms. Gulf of 


Mexico, West Fla. region, dredged U.S. Fish Com., bot- 
tom temperature 46° F. 


nobilis Merrill, Tr. Conn.) Acad: /Seni6, sp izogy plazas: 
Dall Ball. Moi Z.. 718, p53) 1869s leilsbmy.wNtaney © onelny 
TS CIAM EllS NDA 24 OmOoe 
Distribution.—Martha’s Vineyard 906-1309 fms., Delaware 
Bay log1-1209 fms., Tobago 880 fms., Gulf of Mexico, U. 
S. Fish Com. Station 2127, at 1639 fms. 


50 MOLLUSCA OF THE GULF OF MEXICO 20 


Genus SABATIA Bellardi 


bathymophila Dall, Bull. M. C. Z., 9, p. 98, 1881, (as Azys?);18, 
PSs to NOD. Son Bulls) 27. Wel SiNoy Meni ps 36, pl. 
Mt Os OD. SOx 
Distribution.—Fernandina to Guadeloupe, 294-1568 fms. 
Gulf of Mexico, Yucatan Strait, 740 fms. 


Genus CYLICHNELLA Gabb 


bidentata d’Orbigny, Moll. Cuba 1. p. 125, pl. 4, f. 13-16, 1841, 
Gsve7da) Gabb.) (Pr. AYN Se Phila 240 ou 27h) pla iO i 
ae alle (Bulle 37. Ul Se Neri py. 86) plu anaite) wa 780); 
Wenattay Pr AU NS. Phila. 5 5.yp9075 75) L903: 


Bulla biplicata Lea. Pr. Bost, Soc. N. H., 1, p. 204, 1844. 


Distribution.—Hatteras to Barbados, 7-168 fms. Pliocene 
to Recent. Gulf coast, Recent: West Florida and Texas. 
Pleistocene, Louisiana, Knapp’s No. 2 "well, Terrebonne 
Parish, 1519-1542 ft., Crowley No. 4 well, Jennings, 1663- 
1670 feet. 

bidentata var. gaivestonensis Harris, Bull. Am. Pal., vol. 1, p. 
GoOmpla it: 2) 1895; 
Distribution.—Upper Miocene, Galveston well at 2600-2733 


HECE: 
Genus BULLARIA Rafinesque 


striata Bruguiere, Ency. Meth., 1, p. 572, (as Bulla); Dall, 
Bullen 37, Wl SNe Me) 1p. 88,) 1889; Pilsbry,)) Man. Conch.) 1; 
Sell SD See i Dle a7 uty 42-40... Oze 
Distribution.—Recent, Mediterranean ahd Atlantic coasts of 
Portugal and Morocco. Also living at Clearwater Harbor, 
West Florida; New Harbor Isls. and Chandeleurs, Louisi- 
ana; Texas. Pliocene, Caloosahatchie marls, Fla. 


occidentalis A. Adams, In Sowerby’s Thes., 2, p. 577, pl. 123, f. 
VERO S On (as bulla) Wall ei Ui os Ni MEMO np 1324) 
Se uRully a7 UES) Ne Mi ip.) 88.0780; )oimeley; 4th Amn: 


21 BULLETIN 38 st 


Rept (Geol Surv We pag320 02 Naiatca, wet weno 
Phila 5 5. pai s7 se koos 
Distribution.—Florida to St. Vincent, W. I. Gulf coast: 
Cedar Keys, St. Marks, Tampa, Crooked Isl (Calhoun Co.), 
Fla.; Corpus Christi and Matagorda Bays, Texas. 

solida Gmelin, Syst. Nat., 13, p. 3434, (as Bulla); Dall, Bull. 
Bh nO SNe MESS py) SSeS ocubilsbiey ny Miany Conch imisets 
U4: Oy NS Olly AUG Teh Ia, VECO). 
Distribution.—West Indies, Vera Cruz and Maracaibo. 
Cited from Texas region, Gulf of Mexico (Dall, ’8q). 


abyssicola Dall, Bull. M. C. Z., 9, p. 97, 1881, (as Bulla); 18, 
SO) pla i7e tai So; Bulle es 7 aaOr oN in DSS) sp ka aly 
Tl SOselisbry, Man. (Comchessiasemen 5.) 15 23o. plan sor 
21, 2S93" 

pinguiscula Jeffreys, Ann. & Mag. N.H., p. 318, 1880, (women 
nudum); Watson, Chall. Exp. Gastr., p. 638, 1886. 
Distribution.—Bay of Biscay, Azores, Antilles and Yucatan 
Strait, 450-640 fms. 


Genus HAMINGEA Turton 


elegans Gray. Ann. of Philos. new ser. 9, p. 408; 1825; Index 
estac.) ‘Suppl. Apliii2) wigienuas) Pella) Pilsbry, Vian 
Conchre ar iseraerss ps5 5 ale wate 37-30), Pl 4ONmer Sos 
1893; Dall and Simpson, Bull. U. S. Fish Com., 1, p. 364, 
IQOI. 
guilding?t Swainson, Malacol, pp. 251, 360, f. 46, 1840; Adams, 
hes!) P4580.) pl m2aaikas 7 Somalia ull 277, Unis siN eaves 
p. 88, 1889. 


Note.—/ide Pilsbry, the true elegans of Gray is this species 
and not a European shell. 


Distribution.—Antilles to Rio de Janeiro. Gulf coast, West 
Florida and Texas. 


52 MOoLLUSCA OF THE GULF OF MEXICO 22 


Succinea Conrad, Pr. A. N. S. Phila., 3, p. 26, pl. 1, f. 5, 1846, 
Geyaiaye Dally eri ioaM Ney MeO ps 6324,1 08837 Bull, 
PUR Oe Nig Me pa 88, 1803) Mitchellt Mist) Mex) Sh; 
ilcbiye Many | Conch, 1Ser) 15.) py 35 jy Dla 484i 119,393; 
Dall and Simpson, Bull: U. S. Fish Com., 1, p. 364, Igot. 
Distribution.—Indian River, E. Fla., to Texas. West 
Florida at Tampa (Conrad’s type locality), and Cedar Keys; 
Chandeleurs, La. 


aniillarum d’Orbigny, Moll. Cuba 1, p. 124, pl. 4, f. 9-12; 
Mosrehy Mal Bl. 22) p. 1755) Walle, Bulls (37. U2 5. No M.; 
MSs Tsso-) Pilsbry, Mant iConchy. 1Sen) 15. ps1358,) ple 41, 
f. 35, 36, 1893; Mitchell, List Texas Shells. 


cerina Menke, Zeitschr. fur Mal., p. 142, 1853. 


Distribution.—W. Florida to Porto Rico and St. Thomas. 
Gulf coast at Tampa, Fla., and Espiritu Santo Bay, Texas. 


antillarum var. guadalupensis Sowerby, Conch. Icon., f. 14, 1868; 
eilsbraya Mian (Conch, 1 (ser. 5.) pa a5on) Dis) 405/11 30-34,, 
1893. 

Distribution.—Guadeloupe, Cuba &c., Gulf coast at White 
Water Bay, West Florida. 

petiti d’Orbigny, Moll. Cuba 1, p. 130, pl. 4, bis. f. 13-16, (as 
Bava Dally Bulli i272 Ui. SusNe a Mops Soup TSso a bllsbiny, 
Mane Conch... 1) Ser, 15, p. 359, plied) 234) 245, 1893. 
Distribution.—St. Thomas, W. I. and Tampa, West Fla. 

virescens Sowerby, Genera, No. 39, f. 2; Dall, Tr. W.1.5., 3, 
Pas 1890, kilsbry, Man. Conch:))1 ser, 15" p: 360, pl. 4o, 
temo Pl 43, 1-119; 1893. 

Distribution.—Recent on the Pacific coast, California to 


Mexico. Pliocene, Caloosahatchie beds, Fla. Not in the 
recent Atlantic fauna. 


Genus CYLINDROBULLA Fischer 
beaui Fischer, Jour. de Conch., p. 275, pl. 8, f. 8, 9, 1856; Dall, 


23 BULLETIN 38  ' 53 


Bull. 37, U. S. N M., p. 88, 1889; Pilsbry, Man. Conch., 1 
SehS ps 280 pla A 2uit IO 2 OMS aOeE 


Distribution.—Guadeloupe (type locality), St. Thomas, W. 
I.; Gulf coast at Cedar Keys, 2-95 fms. ‘This species is the 
genotype. 


Genus HYDATINA Schumacher 


physis Linnzeus, Syst. Nat., ed. x., p. 727, 1758, (as Bulla); 
Dall Bulli ie7 00s 1S INS Ores ais oOr ealsbinyan Wate 
Conch, 'ser1015, 9. 387.) plague 8037) Dallvand 
Simpson) /Bull7U.S. Fish Comayarip. 364) noon. 
Distribution.—Atlantic, Pacific and Indian Oceans. Gulf 
coast: Sarasota, West. Fla. 


Genus PHILINE Ascanius 


flexuosa M. Sars, Nyt. Mag. f. naturvidens., 17, p. 181, pl. 11, f. 
23-26; Christianiafjordens Fauna, pp. 69, 70, pl. 11, f. 23- 
26-4 Wall Bull Me iC Zins Som sou reso bull 27. Un Senin 
Mi p88, 1889; Pilsbry,)ManynConch!) im ser.) 16) ;pyi214iple 
4, f, 86-89, 1896. 


Distribution.—Christiania, Norway; Yucatan Strait, 640 fms. 


Genus TETHYS Linnzeus 


protea Rang, Hist. Nat. des Aplysiens, p. 56, pl. 10, f. 1-3,1828, 
(as Aplysia): d’ Orbigny, Moll. Cuba, 1, p. 117; Beau, Cat. 
Coq. Guadeloupe, p. 20; Dall, Pr. U.S. N. M., p. 324, 
193; Pilsbry,. Man) iConcheiiraseruno.np. 78. pl. 3 7an2o-22. 
1895; Dall & Simpson, Bull. U. Shi Fish Com: 1) pies: 
TQOL. 


Distribution.—Florida to Carthagena, Columbia. Very 
abundant in the Antilles. Gulf of Mexico, West Florida 
(Dall). 


willcoxi Heilprin, Pr. A. N. S. Phila., p. 364, 1886, (as Aplysza) ; 
Da Bialls337, Wi SaNeeNieMpa GO SO) loll SbivaiMiane 


54 MOLLUSCA OF THE GULF OF MEXICO 24 


Conch., I ser. 16, p. 80, pl. 35, f. 30-32, 1895; Johnson, Occ. 
aoe BOSte SOC. ING Tine 5 An) ROWS. 


Distribution.—Little Gasparilla Bay and Marco West 
Florida. Also Massachussetts coast. 


ORDER PULMONATA 


Genus AURICULA Lamarck 


(Auriculastrum) pellucens, Menke, Syn. Meth. Moll., p. 131, 1830; 
Daler WSN): MESA pha Tsui WTS. fay Sos (ioctl le 
BOS) Ne) Menip! oo. plac sete SenlS so: 


Distribution.—Cedar Keys, Florida, to Demerara, British 
Guiana. 
Genus PEDIPES Scopoli 


mirabilis Muhlfeld, Mag. Ges. Nat. Freude, Berlin, 8, p. 8, pl. 
MPN nae ay TSS 4) Dall Bullies SN Given pa O2)\ipll 
47, f. 17, 1889: Dall & Simpson, Bull. U. S. Fish Com., I, 
DP BOOW ples 53,11.3;) LOOl. 
guadridens Pfeiffer, 1839: ovalis Adams, 1849, ¢ridens Pfeiffer, 
1854. 
naticoides Stearns, Pr. Bost. S. N. H. 13, p. 108, 1869; Dall, 
Pr W)o2 INS )MiA8) 5.270) Pi=Ee me a7 Lo O5) 
Distribution.—Tampa, Florida,to Guadeloupe Island. 
elongatus Dall, Pr. U. S. N. M.,8, p. 279, pl. 18, f. 4, 1885; Bull. 
BUL SOHING ME o2s) pl. 47, ita ATs so. 


Distribution.—Marco, Southwest Florida. 


Genus MELAMPUS Montfort 


coffeus Linnzeus,Syst. Nat. ed. x, p. 729, 1758,(as Bulla); Dall, 
mB alle s7a Wy. 5. UNG Me ps 92, 9pl4azaih.1/35 R889, Dall & 
Simpson, Bull. U. S Fish Com. 1, p. 368, pl. 53, f. 13, Ig0T. 


coniformis Bruguiere, Ency. Me’th. 1, p. 339, (as Bulimus). 


25 BULLETIN 38 55 


Auricula ovula d’Orbigny, Moll. Cuba 1, p. 187, Atlas, pl. 13, 
Lea TS 534 
Distribution.—Florida to French Guiana. Pleistocene to 
Recent. Gulf coast, Cedar Keys, Fla. and Texas. 


coffeus var. gundlachi Pfeiffer, Vanatta, Pr. A. N.S. Phila., 55, 
P- 757, 1903. 
Distribution.—St. Andrews Bay, Washington Co., Fla. 


floridanus Shuttleworth, MSS., Pfeiffer, Mon. Aur. Viv., p. 36, 
7O56: (Mall, Bulle s70Ue Sine Mies ozanmln anit 2) 153.0. 


Distribution.—Tampa to Florida Keys. 


flavus Gmelin, Syst. Nat., p. 3436, 1792, (as Voluta); Binney, 
Jour.Bost.Soc.) Nat) Hist eipen 60. 1d03--land Galkin 
Water Shi-pt..2. p. 12,°65; 2DallkperanWens, IN: Mis) 8) ip:zom. 
plMmsit. 2) 85 ullie 27 OM Same enn p 4 O2n mol. Ar aleeoTee 
°89; Dall & Simpson, Bull. U.S. Fish Com., 1, p. 368, pl. 
54, f. 9, 1901; Mitchell, List Texas Shells. 


Voluta flava Gmelin, Syst. Nat. p. 3436, 1792. 

Bulimus monile Bruguiere, Ency. Me’th., 1, p. 338, 1780. 

torosa Moerch, Cat. Yoldi, 38, 1852. 

coronatus Adams, 1849; coronulus Adams, 1854. 
Distribution.—Florida to Guadelonpe. Gulf coast: Tampa, 
Cedar Keys, Fla.; Point at Fer, Wa.; Port Lavaca and 
Espiritu Santo, Tex. 
Note.—According to Mr. Mitchell, this species lives near 
shore, hiding in the day time under drift wood and sea 
weed and goes in search of food nocturnally. 

lineatus Say. Jour. A. N.S. Phila., 1 ser. 2, p. 246, 1822; Dall, 

Bullve7, Uns. NR IMe poe tnp lag pet) (L240 SQ~.Simedleye 
4th Ann. Geol. Rept. Texas, p. 352, ’92: Johnson, Occ. Pa. 
BOSE SOC. ENG Hal 74 (pn aiSeomse 

Wiaeniatus Says yout.) A. INGi Sikandar Sele 2) up) 2415 O22 
Dekay. INV.) Moll pansye Dias ateO2.) mln Zar cau enwar 


56 MoLLUSCA OF THE GULF OF MEXICO 26 


Binney, Land & Fr. W. Sh., 2, p. 10, ’65; Not dzdentatus 
Montagu, 1803, which is European, but introduced on New 
England coast. 


biplicatus Pfeiffer, Mon. Auric. Viv., p. 21. 


Auricula cornea Weshayes, Ency. Me’th., 2, p. 90, 1830; 
jaumet Mittre’, 1841. 
Distribution.—Maine to Tortola Isl., Gulf coast: Tampa, 
Cedar Keys, St. Mark’s, Fla.; Point au Fer. Chaudeleurs, 
Belle Isle, La ; Galveston, Tex. 


Note.—Northern specimens very distinct; southern grade 
towards flavus with which they may hybridize. Inhabits 
salt marshes. 

(Detracia) bulloides Montagu, Testacea Brit., p. 339, pl. 30, f. 
WISes. (as. Volutia). Dall) Pra) Ue S. Na) ME) 8) p.)285, pl 
Ieee SSSe) Bull..37), UW. Sa Nee so g2, plia7, f7 7, 89. 


Distribution—Cedar Keys, Fla. and the Antilles. Type of 
Gray’s subgenus Detracia. 

(Sayella) hemphilli Dall, Pr. U.S. N. M., 6, p. 323, pl. 10, f. 6, 
1883, (as Leuconza); Pr. U. S. N. M., 8, p. 286, pl. 18, f. 
Tso Bulle WS.) ING Mei 7) paoou pls Az rey ith 8O); 
Singley, 4th Ann. Rept. Tex. Surv. p. 343, ’92. 


Distribution.—Cedar Keys, Florida, and Corpus Christi, 
Texas. ‘Type of section Sayella Dall which is intermediate 
between Blauneria and Detracia. 

(Sayella) crosseana Dall, Pr. U. S.N. M., 8, p.,286, pl. 18, f. 10, 
sey Bulllis7. U.S. Na M., p. 92) pliy47,) {2 10,789") oingley, 
AthieAnn Rept. Lex. Surv., p-) 343,92). 


Distribution.—West Florida at Egmont Key (Tampa Bay); 
Corpus Christi, Texas and the Bahamas. 


Genus PLANORB1IS Guettard 
Ophis Dall Proc. U.S: N. M:, 46, p. 236;;pls 21, f:/3,'\45 1914. 


27 BULLETIN 38 57 


Distribution.—Upper Miocene or Pliocene. Well near Al- 
exandria, La., at 49 feet. Brackish water formation. 


Note.—The living species of this fresh water genus are not 
included in this Synopsis. 


Genus TRALIA Gray 
pusilla Gmelin, Syst. Nat. p. 3436, 1792, (as Voluta); Dall, Pr. 
Wa Sn NGM Secon a7 6a Wins limite sain eis) ECL a7 Oise Ne 
NE o2), (pl Ag in. sii Son mall Sai Simpson allen mss 
Fish) Com.) ps) 260; plisaiite 13hinoom. 


triplicata Donovan, 1802; ovula Bruguiere, 1789; z¢tens 
Wamarck ns22% 


Distribution.—Cedar Keys to Guadeloupe. 


minuscula Dall, Bull. 37, U.S N. M., p. 92, 1889. 
Distribution.—Tampa to Bahamas. 


(Alexia) myosotis Draparnaud, Tabl., p. 53, ’o1, (as Auvzcula); 
Dall PrUS MN OMS i Dwi 7 Ame se motile 2 Zep Ss) N tem ules 
DHOZ Mpls 25 fo 8o! 

denticulata Montagu, Test. Brit., p. 234, pl. 20, f. 5, ’02, (as 
Voluta) 


Distribution.—Europe; Jamaica (introduced); West Florida, 
San Francisco (introduced). 


Genus BLAUNERIA Shuttleworth 


heteroclita Montagu, Test. Brit., Suppl., p. 469, 08, (as Voluta); 
Arango) Moll; Cubanayp icoma7s Wall Pro) OL) So uN vies 
Bp. 287) pl. 17. fh GN SeiBuliene Wes.) NGM), ip-kozmiple 
Az, t.)t4, 80, Dall &iSimpsoum bully) S.)Rish Con: in woe 
369, I9O01. 
pellucida Pfeiffer, ’40, cubenszs Pir. 


Distribution.—Tampa and Marco, West Florida. Also 
Antillean. 


58 MOoLLUSCA OF THE GULF OF MEXICO 28 


Genus SIPHONARIA Sowerby 


alternata Say, Jour. A. N, S. Phila., 5, p. 215, ’26, (as Patella); 
Bimmer Nand Ge Fol Sis) 25) p. 0535) to. 2540005; Dall Px. 
eon NAN Ss pn 28797 S57 Bull i270 WS. Menai pe 192) 
1889. 
Distribution.—The variety brunnea Hanley (Pr. Z. S., pp. 
21,151, 58)is found at Sarasota Bay. West Fla.,and Bermuda. 


lineolata d’Orbigny, Moll. Cuba, 1. pl. 17, f. 13-15; Dall, Bull. 
BAUS SUN. IVD Sp o2) 80: 
Distribution.—Fernandina, Fla., to Braziland Texas region 
of the Gulf of Mexico. 
Genns WILLIAMIA Monterosato 
krebsi Moerch, Dall, Bull. 37, U.S. N. M., p. 92. ’89. 
Distribution.—West Florida to Barbados. 


Fenus YVERONICELLA Biainville 


floridana Binney, Terr. Moll. U.S., 2, pl. 67, f. 17, °51; Dall, 
Bulla 7. U5) N. M.; p- 90,1889. 
Distribution.—Charlotte Harbor, West Florida to Florida 
Keys. 
Note.—Since Blainville erroneously attributed a shell to this 


slug-like genus, Ferussac’s later name Vagznula is often 
used; but Vevonicella has priority. 


ORDER CTENOBRANCHIATA 


Genus TEREBRA Bruguiere 


cinerea Born, Test. Mus. Vind., p. 267, pl. 10, f. 11, 12, 1780; 
avon Manki 7p. 21 pl. Oo) tv Ova Toss Dalle spill vie C: 
ZL e POs so. Sineley,, 4th) Annee Rept) Dex. \py 332) 
qo2; Dall) & Simpson, Bulls) Us Ss ikishy Com: 1 ip. 382) 
IQOT. 
salleana Deshayes, jamatcensis C. B. Adams. 


29 BULLETIN 38 ; 59 


Distribution.— West Indies to Colombia. Gulf coast: 
Chandeleurs, La., Corpus Christi, Tex. Pleistocene, New 
Orleans Pumping Station, No. 7. 


dislocata Say. Jour. A. N. S. Phila., 2, p. 236, 1825, (as 
Cerithtum); Hilgard, House of Rep., Ex. Doc. 1, pt. 2, p. 
S87 785) bryou,) Manwi7, joy TSy (plein Og tino sm steledleiye 
4th Amn. Rept), Mexia.) a2. OZ Vanattann ai ya Neno: 
Philay (550 P7575, 2903. 
rudis Gray, petttiz Kiener. Not petit? Maury, 1917. 


Distribution.— Maryland to West Indies and Venezuela. 
Miocene to Recent. Gulf coast Recent; Cedar Keys, Fla.; 
Chandeleurs, La.; Galveston & Corpus Christi, Texas. 
Pleistocene of New Orleans pumping station No. 7; New 
Orleans artesian well of 1856; Lake Borgne borings; Knapp’s 
wells, Terrebonne Parish, No. I at 1600-1700, 2000-2150 ft., 
No. 2 at 1150-1290, 1731-1739, No. 3 at 1150- 1525 feet. 
Also of North Creek, Manatee and Labelle, West Fla. Up- 
per Miocene, Galveston well, 2158-2920 ft. 


concava Say. Jour. A. N.S. Phila., 5, p. 207, 1827; Dall, Tr. W. 
Inst. Sci., 3, p, 24, 1890; Singley, 4th Ann. Rept. Tex.; p. 
2325.92 Haris, Bulk vA Mralee wv oleer. ps O71 Os. 


Distribution.—N. Carolina, Georgia and Texas, at Galves- 
ton . Pleistocene of North Creek, Fla. Pliocene, Caloosahat- 
chie, Fla. Upper Miocene, Galveston well at 2552-2920 ft. 
Note.—The variety vinosa Dall (Bull. M. C. Z., £8, p. 64, 
*89) is found on the Gulf coast at Sarasota Bay and Char- 
lotte Harbor, West Fla. 

protexta) Conrad, Pr, A.) Nis) Phila’, 2), ps) 26) )184ai (as 
Cerithium): Cryon) Mantz 25)pl. (6, 1.168, 11805 salle 
Ball37) UliS: NoMa Vpsio4aNe sox air. Wagner lmst.eiah ps 
25, 90; Dall & Simpson, Bull, Fish Com. I, p. 382, 1901; 
Vaughan, Publ. No. 133, Carn. Iust., p. 171, 1910; Maury, 
Boll Am) Pall 20) ps2 aplaquvt tole 


60 MOLLUSCA OF THE GULF OF MEXICO 30 


Distribution.—Hatteras to Texas 2-50 fms. Lower Miocene 
to Recent. Gulf coast: Cedar Keys & St. Marks, Fla.; 
Cameron, La.; Galveston, Tex. Pleistocene, North Creek, 
Manatee and Labelle, Fla.. The variety J/utescens E. A. 
Smith, ranges from N. Carolinato West Florida. It is 
more distinctly reticulated than the typical form. 


nassula Dall, Bull. M. C. Z., 18, p. 66, pl. 36, f. 8, 1889; Dall 
& Simpson, Bull. U. S. Fish Com., 1, p. 383, 1901. 


Distribution.— Martinique. Also Gulf of Mexico, Lat. 23° 
N., Lon. 88° W., at 95 fms.; Yucatan Strait, 640 fms. 
limatula Dall, Bnll. M. C. Z.,18, p. 66, 1889; Bull. 37, U.S. N. 
M., p. 94, ’89. 
Distribution.—N. Carolina to Barbados. Also Gulf of Mex- 
ico between Mississippi delta and Cedar Keys at 111 fms. 
texana Dall, Nautilus, 12, p. 45, Aug., 1898; Pr. U. S. N. M., 
PMID aSO2 ply 2On te (Oo) 1902). 
Distribution.—Matagorda Island, Texas. Collected by Hon. 
J. D. Mitchell. Very rare. 
galvestonensis Harris, Bull. Am. Pal., vol. 1, p. 98, pl. 3, f. 5, 
1895. 
Distribution.—Upper Miocene, Galveston well at 2387-2871 
IQIO. 


calhounensis Maury, Bull. Am. Pal., No. 21, p. 4, pl. 1, f. 3, 
19IO. 


Distribution.—Lower Miocene, Chipola marl, Bailey’s 
Ferry, Fla., and Bascom No. 2 well Mobile, Alabama, at a 
depth of 1241 feet, Chipola horizon. 


indenta Conrad, Cited by Mr. Aldrich from the Bascom No. 1 
well, Mobile, Ala. at 1500-1556 feet. 

langdoni Dall, Pr. U. S. N. M., 18, p. 39, 1895; Tr. W. Inst., 
3, pl. 59, f. 27, 1903. 


BULLETIN 38 — 61 


Distribution.—Lower Miocene, Chipola marl, Fla.; Upper 
Miocene Galveston well at 2158-2920 feet (Harris). 


Genus CONUS Linnzus 


proteus Hwass, Enc. Meth. vers, 1, pt. 2, p. 682, 1789; Reeve, 
Conch Iicontit 210/00 Dalla Bull 27a UE SweN Mes omane 
1889 Vaughan Publ: Now 122 Carn. Imstcips 7p volo: 
Many.) Bulli) Am Pal No Zov meat pl Gn hanno 


leoninus Hwass, 1791, spurius auctores, as of Gmelin. 


Distribution.—West coast Florida, 19 fms. 
zuela. Pliocene, Caloosahatchie River, 


Labelle, Fla. 


South to Vene- 
and Pleistocene, 


centurio Born, Mus. Vind., pl. 7, f. Io, 1780; Tryon, Man., 6, p. 
33, f. 9, f£. 68,°1884: Dall, Bull. M. C. Z., 18, p. 69, °89. 


Distribution.—Antilles. Also Gulf of Mexico between 
Mississippi delta and Cedar Keys at 25 fms. 


delessertianus Recluz, Mag. de Zool., pl. 72, 1843; Tryon, Man., 


6))p!133.)) pl. Oo) fy 677) Sam Dall (asi delessextze i iaalleviere 
We Sh NEEM Di OAe SOR 


Distribution.—Hatteras to West Florida, 22-63 fms. Doubt- 
ful specimen, Porto Rico. 


flavescens Gray, Sowerby, Conch. IIl., f. 68; Tryon, Man., 6, p. 
86, pliimo) he 84,,rssacDallmB allies NOLS. Nien aa ae 
1889. 


Distribution.— Hatteras to Barbados, and West Florida, 15- 
170 fms. Pliocene to Recent. 

floridanus Gabb, Am. Jour. Conch., 4, p. 195, pl. 15, f. 4, 1868; 
Aryon,)Man:.6,) p)i28splimmeutaAnigh arsed Dalle vp imilia zai 
94, 1889; Ir. W. Inst. Sci., 3, p, 27, 1890. 


Distribution.—Hatteras to Florida Keys and West Fla. 
Pliocene of the Caloosahatchie beds. 


64 MOLLUSCA OF THE GULF OF MEXICO 32 


peali Green, Tr. Albany Inst.,1, p. 123,pl. 3, f. 3, 1830; Tryon, 
Mean 6. ps) Z0.)plai 10. )f2/89-92))) 1884; (Dall) Bulls MM: C. Z., 
18, p. 68,789. 
seannseconntad) yAm.4) foun Coneh. is) pi toa pl.) TO} /f.01, 
1869. 


Distribution.—Hatteras to Isthmus of Panama. Also Cedar 


Keys, Florida, o-5 fms. Pliocene, Caloosahatchie beds. 


pygmaeus Reeve, Pr. Z. S., p. 179, 1843; Kiener, Coq. Viv. 
Conus, p. 174,pl. 111, f. 1 a-b,1848; Dall and Simpson, Bull. 
WSS) Bish Com.) 1, p. 383. 190K: 


Distribution.—West Florida to TobagoIsl. Pliocene, Ca- 
loosahatchie beds. Pleistocene, North Creek, Fla. 
amphiurgus Dall, Bull. M, C. Z., 18, p. 70, 1889. 
Distribution.—Coast of Yucatan, 27 fms. 
chipolanus Dall, Pr. U. S. N.M., 18, p. 42, 1895. 
Distribution.—Lower Miocene, Chipola marls, Florida and 
of the Bascom No. 2 well, Mobile, Alabama, at 1241 feet. 
Genus TURRIS Bolten 


albida Perry, Conch. Expl., pl. 32, f. 4, 1811, (as Pleurotoma); 
Wall Bull27) We Ss.) IN.) ME p96) 7780; Wall Simpson, 
Bulle Wes. Hish! Com). 1) pi 284. 190r- | Maury. (Bulli Am, 
Bale INO: )29, (pp. 50; 1917: 
Distribution.—Recent, Cedar Keys to Barbados, 26-100 fms. 
Oligocene of Yicksburg, Miss., and of Tampa, Fla.; Lower 
Miocene, Western Florida and Santo Domingo; Upper Mio- 
cene, Galveston well at 2158-2950 feet. 


Note.—The variety ¢el/ea Dall (Bull. M. C. Z., 18, p. 72, 
1889) was dredged in the Gulf of Mexico between Missis- 
sippi delta and Cedar Keys. 
Genus LEUCOSYRINX Dall 
verrilli Dall, Bull, M. C. Z., 9, p. 57, 1881, (as Pleurotoma); 18, 


33 BULLETIN 38 63 


PA Siiple LO, 55. 18 So me ulle Brailes pORMensm OO! 
Distribution.—N. Carolina to Guadeloupe. Also Gulf of 
Mexico, Lat. 25° N., Lon. 84° W., 888 fms., and between 
Mississippi delta and Cedar Keys. 

sigsbeei Dall, Bull. M. C. Z., 9, p. 57, 1881, (as Pleurotoma) ; 
18.270, \pl. li tO. Sos Bulla TOE A SMU Nu NIE Map AOON 
Pla Ths OMe So: 
Distribution.—Yucatan Strait, 640 fms. Off Bequia, 1591 
fms. 

tenoceras Dall, Bull. M.C. Z:, 18; p: 76; pli 36, £. 5; 1889; 
Bullita7) OS. Nee pi oo" plaizoamanssnioo: 


Distribution.—N. Carolina to Guadeloupe. Gulf of Mexico 
between Mississippi delta and Cedar Keys at 724 fms. 


subgrundifera Dall, In Agassiz’s Three Cruises of the Blake, 2, 
p: 66, f: 282, 1388, (as Plezrotoma) Bulle Me Cy Zan sa: 
77) Oly Z8e 142) 789. Bills, 27.(U Se eNEy Mes DOO, please 
I Oy 
Distribution.—Cape Fear, N. C., to St. Kitt’s, W. I. Gulf 
of Mexico, Yucatan Strait, 640 fms., and between Missis- 
sippi delta and Cedar Keys, 940 fms., U. S. Fish Com. St. 
2384. 

Genus ANCISTROSYRINX Dall 

radiata Dall, Bull. M. C. Z., 18, p. 78, pl. 12, f. 12, 1889; Bull. 
87h Uns. Ne Me. p06) plana tegen So. all) Se. Simpsons 
Bull U.S. Kish Com, (1, pigsanimoor: 
Distribution. —Gulf of Mexico, Yucatan Strait,64o fms. and 
between Mississippi delta and Cedar Keys. Also Antillean. 

Genus GENOTA Adams 

mitrella Dall, Bull. M.C. Z., 9, p. 56, Aug., 1881, (as Pleuro- 
LOD) N18 op., 794 Plinnletaisy mySormbtt len 277. ns) SsuN aver 
TO) CO) ONY 107) 505 Be 1 Mesoy. 

didyma Watson, 1881. 


Distribution. —Yucatan Strait, 640 fms. Also off Sombrero. 
Weal. 


64 MoLLUSCA OF THE GULF OF MEXICO 34 


Genus DRILLIA Gray 


ostrearum Stearns, Pr. Bost. Soc. N. H., 15, p. 22, 1872; Tryon, 
Man Pap mor pla te 7 O a oar Oallem Bullen ise N 
M., p. 96, ’89; Dall & Simpson, Bull. U. S. Fish Com., 1, 
Pp. 385, 1901. 
Distribution.—Hatteras to Cape Catouche, Yucatan and An- 
tilles. 15-170 fms. West Florida: Recent, Cedar Keys; 
Upper Oligocene, Tampa; Lower Miocene, Chipola beds; 
Pliocene, Caloosahatchie beds. 


albicoma Dall, Bull. M. C. Z. 18, p. 83, pl. 10, f. 8, ’89; Bull. 
We Ss Ne Me; 37,0: 96; pl: 10,/f 8); 1889: 


Distribution.—Gulf of Mexico, Lat. 23° N., Lon. 89° W., 
84 fms. Also off Barbados, and St. Thomas. 


detecta Dall, Bull. M. C. Z., 9, p. 65, Aug., 1881: 18, p. 84, pl. 
Wei Toso.) bull. 37." ps 96) plaio han Sor 
circumvoluta Watson, Jour. Linn. Soc., 15, p. 465, Nov. 1881; 
ReptyChall, Gastr. p. 357, plai2ik, f.10. 188.5. 


Distribution.—Gulf of Mexico, 339 fms. Also Culebra 
Sin We 


alesideta var. macilenta Dall, Bull. M. C. Z., 18, p. 85, pl. 36, f. 
TSS On bulla a7 NU.) S.No VM pooh ply 26nihon 78o) 
Distribution.—Cape Fear, N. C. to Barbados. Gulf of 
Mexico between Mississippi delta and Cedar Keys, 111 fms. 
Typical form of species, off Hatteras. 


eucosmia var. canna Dall, Bull, M. C. Z., 18, p, 86, 1889; Bull. 
BROS ON Mie. 1: 96; 780. 
Distribution.—Cape Lookout, N. C., to Grenada, W. I. 
Also Gulf of Mexico, west of Florida, 50 fms. 


leucomya Dall, Proc, U.S. N. M., 6, p. 328, pl. 10, f. 8, 1883; 
Bel, Vy, Fs COR jello sig sy Sustskeye “Aes WGN DBalsie 7 hy yb eer 
1890. 


35 BULLETIN 38 65 


Distribution.—Shores of Gulf of Mexico from Sarasota, 
West Fla., to Yucatan, 3-5 fms. Pliocene, Caloosahatchie 
beds, Fla. 

albinodata Reeve, Zool. Proc., 6, 1846; Tryon, Man., 6, p. 197, 
plyags £5, 1884) DalleBull 27, We SiN MES: oO So: 
Distribution.—Charlotte Harbor, West Fla., to Santo 
Domingo. 


haliostrephis Dall, Bull. M. C. Z., 18, p. 86, pl. 13, f. 3, 1880; 
Baller UES. WING MLL pi Oo hp lem TigwEnoNs GO! 


Distribution.—Gulf of Mexico, West of Florida, 84 fms. 
acestra Dall) 'Bull’'37/°U) S: NVM poor ple ro, tf. 7) So. 


Distribution.— West Florida to Grenada 161-400 fms. 


pharcida Dall, Bull. M. C. Z., 18, p, 88, pl. 12, f. 2, 1889; Bull. 

B27 Mp UOOMple U2 teN2 Os 
exasperata Dall. Not of Reeve. 

Distribution.—Gulf of Mexico, west of Florida, 229 fms. 
Also East Fla. to Barbados 150-1002 fms. 

tristicha Dall, Bull. M. C. Z.. 18, p. 88, 1889; Bull. 37, p. 98, 
1889. 
Distribution.—Three stations Gulf of Mexico between Mis- 
sissippi delta and Cedar Keys, 111-210 fms. ‘Temperature 
522 - Ore 

ebur Reeves, Pr. Zool. Soc., p. 116, 1845, (as Pleurotoma); 
Conch!Tcon!) pli 20M ams as dinyony, Man.) 6) pumnser 
plas) £56, 84: Dall Bulle Ue Sa uNe EME .\p.O8aisox 
Distribution.—Hatteras to Sombrero Isl. Also Gulf of 
Mexico, West of Florida, Blake Station 10, at 34 fms. 


pagodula Dall, Bull. M. C. Z., 18, p. 90, pl. 13, f. 6, 1889; Bull. 
By pO8y ple 138 £6) Soe 
Distribution.— Gulf of Mexico, West of Florida, 50fms. 
Also Barbados. 


66 MOLLUSCA OF THE GULF OF MEXICO 36 


ineayDall Pr. U.S. NM.) 6, p. 328) pl) 10) f..5, 2883; Bull, M. 
CALS) Oo 7897) Bullvia7. Us Say Nav Muli. O8) pl. i48; 
eT SO): 

Distribution.— Hatteras to Florida Keys, and West Florida 
at Cedar Keys, 3-15 fms. 
Note.—The variety carminura Dall (Bull. M. C. Z., 18, p. 
gi, 1889) was dredged in Gulf of Mexico, U.S. Fish Com. 
Station 2402 at 111 fms. 

simpsoni Dall, Pr. Davenport Acad. Sci., 5, p. 54, 1887, (as 
ieuvovoma): Bull. M:C.Z., 68) p) or; 1880. 

Distribution.— Hatteras to Tampa Bay, West Florida, 15- 
18 fms. 

lissotropis Dall, Bull. M.C. Z., 9, p. 58, Ang. 1881; 18, p. 91, 
VieELINake4 TS SOs Bull a7 MOs,\ Plauen mien sunain SOs 

hypsela Watson, Jour. Linn, Soc., 15, p. 433, Oct. 1881; Chall. 
Caste s4r ply 21.0. 41885: 

Distribution.—Gulf of Mexico, West of Florida at 220 fms. 
Also Antillean. Variety ferpolita Dall (Tr. W. Inst., 3, p. 
36, ’90), Caloosahatchie Pliocene. 

dalli var. cestrota Dall, Bull. M. C. Z., 18, p. 92, 1889; Bull. 
375 B- 98, °39. 

Distribution.—Between the Mississippi delta and Cedar 
Keys, at 196 fms. Typical dallz Verrill, ranges from 
Martha’s Vineyard to Delaware Bay. 

nucleata Dall, Bull, M.C. Z., 9, p. 62) ’81; 18, p. 92, pl. rr, f. 
1 SH ausxexO)p 
Distribution.—Gulf of Mexico, 229-339 fms. Also Antillean. 
Pleurotoma amblia Watson may be a synonym. 

Ven Dalle Balle Me VC.) 59) sp. 68.) 188i TS.) pn Osi ple mony ik. 
ZR SO Ulla 2 7Aa Ow OS plese bie tei 2 sO. 


Distribution.—Gulf of Mexico, West of Florida, 220-310 fms. 


2 
~ 


BULLETIN 38 ; 67 


havanensis Dall, Bull. M. C, Z., 9, p. 67, 1881; 18, p. 93, pl. 11, 
PASSO! 
Distribution.—Yucatan Strait, 640 fms. Also off Havana. 


lithocolleta Watson, Jour. Linn. Soc., 15, p. 441, 1881; Chall. 
Rept. Gastr., p. 320, pl. 24, f. 6, 1885 (as Pleurotoma); 
Dalia whl MC Z ss Vie Os WO. 180: souls yee 
OVS Holl aiE aby Kn MAVO\s 


a 
i) 
— 
. 
4 
tI 

. 
Lanie) 


Distribution.—Hatteras to Guadeloupe,W. I. Also Gulf of 
Mexico between Mississippi delta and Cedar Keys, at 940 
fms. 


Genus CYMATOSYRINX Dall 


centimata Dall, Bull. M. C. Z., 18, p. 95, pl. 36, f. 9, 1889; Bull. 
27 AO Se NL... QS: spl Zowieor oO! 


Distribution.—Hatteras and Gulf of Mexico, Lat. 24° N., 
Lon. 84° W. at 1920 fms., and between Mississippi and 
Cedar Keys at 1181 fms. 

moseni) Wall, Bull. Mi Co Zur 8 hoof plagon 3 TSsor eb ulie 
BNW Se IN MES OS) pla gon tna SO! 
Distribution.--Cape Hatteras and Sarasota Bay, West Fla. 


Genus PLEURQTOMELLA Verrill 


packardi Verrill, Am. Jour. Sci., (3) 5, p. 15, 1873; Tr. Conn., 
ENCAG OS D145 25 /PlANAe tite oeiOlemorpmON Shane 2 


Distribution.—Gulf of Maineto Rhode Island. ‘The variety 
bencdicin Vii &S,,) (Dall Buller Se INGE paozaple 
14, f. 4, pl. 60, f. 70 a, ’89) ranges from Gulf of Maine to 
Beqttia, 1290-1507 fms.Gulf of Mexico, West Florida region. 


leucomata Dall, Bull. M. C Z., 9, p. 63, 1881, (as Drillia); 18, 
PMLZoepVictt, £) ne wsoy Bally ia7. ap LO2 ence eNtolnieliaun a. 
1889. 


68 MOLLUSCA OF THE GULF OF MEXICO 38 


Distribution.—Gulf of Mexico between Mississippi delta 
and Cedar Keys. Also Florida Strait. 533-940 fms. 


agassizi Verrill and Smith, Am. Jour. Sci., 22, p. p. 394, 1880; 
pittew Coma cade 54) pl 4 Say pl. 5y70 £308 ay oe 


Distribution.—Rhode Island to Cape Fear, N. C. The 
Matiety wcxicana Wall (Bull! 37) U) S. NoM., p: 104) pl. 11, 
f. 14, 1889) ranges from Western Florida and Yucatan re- 
gions of the Gulf to Martinique. 502-640 fms. 


filifera Dall, Bull. M. C. Z., 9, p. 56, 1881, (as Bela); 18, p. 123, 
MLO Ou Sonn Bill: 27.) pitOsNe Dyn 2 1 tanO toOs 


Distribution.—Gulf of Mexico, West of Florida, 331 fms. 


HaugaDale Bull) MC. Z.) 18, \p. T25) 18893) Bully) 37,\ pu 1084) 
1889. 


Distribution.—Cape Fear, N. C., 407 ims. Also Gulf of 
Mexico between Mississippi delta and Cedar Keys, at 1181 
fms. 


extensa Dall Bull: M.C.Z. 9, p. 55, Aug, 1881, (as var. of 
icurotoma blakeana), 18, p. 126, pl. 10) t.)2, 789; Bull. 37, 
DAMLOA ple TOs) f: 2° So! 


FPleurotoma streptophora Watson, Jour. Linn. Soc., 15, p. 464, 
Nov. 1881; Chall. Gastr., p. 366, pl. 10, £. 8 a-b) 1885. 


Distribution.—Gulf of Mexico, Lat. 23° N., Lon. 88° W., 
804 fms.; Yucatan Strait 640 fms. Also North Atlantic. 


blakeana Dall, Bull, M. C. Z., 9, p. 54, 1881; 18, p. 126, pl. 10, 
pio Oc ull 27. my TOA... Ply LOm th eaNoGs 


breois Neral, br. Conn. Acad!,6, pij4ar7s plyiaa tf: 82785. 


Distribution.—Gulf of Maine to Santa Cruz, W. I., 100-1608 
fms. Also Yucatan Strait, 640 fms.; Florida Strait, 339 
fms. 


39 BULLETIN 38 69 


Genus GLYPHOSTGMA Gabb 
gabbi Dall, Bull Me Co Zui) puto. plan ah et Alas Oat SOOe 
B27 SUL SON IMG DA OO. yOl abut HAC ISAM oa oOE 


Distribution.—Gulf of Mexico, Blake Station 36, 84 fms.; U. 
S.F.C.Sta. 2405, west Florida, 30-50 fms. Also off Barbados. 


gratula Dall, Bull. M. C. Z., 9, p. 64, Aug., 1881, (as Pleuro- 
LOMAVE TS De LTO pk M2 TOW Soweto.) Os) LOO molars 
Le TOMASO! 

incilts Watson, Jour. Linn. Soc., 15, p. 425, Oct., 1881; Rept. 

ChalliGastrijip. 2o04n) ple 2auhen Sigs 
Distribution.—Gulf of Mexico between Mississippi delta and 
Cedar Keys, 227 fms. Also Hast Florida and Antilles. 
Pliocene, Caloosahatchie beds. 


Genus MANGILIA Risso 


balteata Reeve, Conch., Icon., 3, pl., 8, f. 57, 1846; Tryon, Man. 
Conch 6) pi 2474 plai24 nes D alin stil i277 ies aaING 
M., p. 100, ’89; Dall & Simpson, Bull. U.S. Fish Com., 
Dis Zs,) LOOM: 
Distribution.—Hatteras to Barbados and West Florida. 
Pliocene to Recent. 

astricta Reeve, Pr. Z. S. p. 46,'46; Tryon, Man., 6,p.260, pl. 24, 
i260 784 Dall Bill a7 Wat SeeN EM VIES ip.) LOO; 30) 
Distribution.—West Florida to Florida Keys. 

biconica C. B. Adams, Conch. Contr., p. 65; Dall Bull. 37, p. 
100, ’89. 
Distribution.— West Florida to Jamaica. Unfigured. 

plicosa C. B. Adams, Contr. Conch., p. 54, 50, (as Pleurotoma) ; 
Dali Buall2 7. Us SNM Me pA MOON DIS @, ft) )l4.. Onsite 
ly, 4th Ann: Rept, Tex. Surv.;)p: 333,92; Johnson, Oce: 
Palsy Bost soc. IN. URES paulAlsp ymiOiis): 


Pleurotoma plicata C. B. Adams, Bost. Jour. N. H., 3, p. 318, 


70 MOLLUSCA OF THE GULF OF MEXICO 4O 


Pht G, 40%) Not of) Wamarck, 
plicatum Kurtz, 1860; drunnea Perkins, ’69. 


Clathurvella jewetti Stearns, Pr. A. N.S. Phila., fcr 1873, p. 
BAO Dally Pry Uo So NG MEG. pi 220)) T8833" 


Distribution. —Massachussetts to Florida Keys and Gulf 
coast at Cedar Keys, Fla.; Corpus Christi Bay, Texas, 0-5 
fms. Pleistocene, North Creek, Fla.; Pliocene, Caloosa- 
hat chie beds, Fla. 

rubella Kurtz and Stimpson, Pr. Bost, Soc. N. H., 4, p. 115,751; 
Dall Bull 27. USS. NeMi py 100,289: Tr Wagner Inst. 
SCZ) Pe 4190. 
Distribution.—Cape Lookout, N. C., to Charlotte Harbor, 
West Fla. Pliocene, Caloosahatchie beds. Pleistocene, 
Knapp’s No. 2 well, Terrebonne Parish, La., at 1800 feet. 


stellata Stearns, Pr. Bost. Soc., N. H., 15, p. 22, ’72; Dall, Pr, 
eon NE IME, 6.0 3280.87) bnyvont Man 6.) 240) pluiaa. 
MOAGon Dally Bal. 2730) pi TOOM SOs Mer aN GON Sin aus 
4I, ’90. 
Distribution.—Cedar Keys and Tampa to Key West, Fla., 
Pliocene, Caloosahatchie beds. 


athoswvlay Dali, Bull!) 37) WU: So Ne Me puino2 plan isa) aay 
1889. 
Distribution.—Hatteras to Barbados. Also West Florida, 
14-333 fms. 


l imonitella Dall Pe WS. ING MEO pi B2Om ple TOs) hy TOMS 83 = 
PPS oul 7 MP y1O2, Pl. j48, £:) 25°89. 


Distribution.—Cedar Keysto Tampa, Fla., o-5 fms. Pleis- 
tocene, Knapp’s No. 3 well, Terrebonne Parish, [a., at 
1150-1200 feet. Perhaps this species is a dwarfed form of 
atrostyla Dall. 


cerina Kurtz & Stimpson, Pr.Bost. Soc.,4, p. 115,1851;(as Pleuro- 
toma); Hilgard, Rept., Chief Engineers to Sec., War., p. 


4I BULLETIN 38 at 


Bos 1870; HOUSe OL Rep wah xn DOCAMu pian 2h Dal SS OMMor 
Dall Pr Uy So NAM Gh ne28. 82 ilnvonun Maniosnps 
Blow plwi22)) faa sae Mall eB illy ea Olas Nien Danilo 
plait. 16a, 180.) Vanattan ier. ALONE S: ieinllaiccs hon sie 
LOO3;: Johnson), Bost. ,ooc. IN. EA Ocesn Pan einen AG. 
IQI5. 

Distribution.—Massachusetts to Texas. Gulf coast: Cedar 
Keys, St. Joseph’s Bay, Crooked Island, Florida; Galveston 
and Corpus Christi, Texas. Pleistocene, New Orleans 
well of 1856; Lake Borgne borings; Grand Chenier; Knapp’s 
wells, Terrebonne Parish, No. 2 well at 1190-1439, No. 3 
at 1500-1525 feet. 


cerinella Dall, Bull. M. C. Z., 18, p. 113, 1889; Bull. 37, U. S. N. 
MDs KOZ WoO: 


Distribution.—Hatteras to West Florida and Galveston, 14- 
22 fms. Pleistocene, Knapp’s No. 3 well, Terrebonne 
Parish, La., at 1500-1525 feet; North Creek, Fla. 


quadrata Reeve, Pr. Zool. Soc., p. 114, 1845, (as Clathurella) ; 
Dnyonti Man. 6).p:/278 platen sen SAS malie Baller 
a SNE MES OS mO2 8 oO. 
Distribution.—Hatteras to Yucatan. Dall regards diminuta 
C. B. Adams (Conch. Contr., p. 63) asa variety ranging 
from Hatterasto Florida Keys and West Fla. 


melanitica Dall, Bull. 37, U. S. N. M., p. 102, 1889. 
Distribution.—Hatteras to Haiti and West Florida. 


antonia Dall, Bull. M. C. Z., 9, p. 59, 1881; 18, p. 116, pl. 10, f. 
Ay planta if 11. So. iBull ey ane oMING VLD. OZ) hp leuimo spt 
AvOlMMntte ATL SSO. 

Distribution.—Yucatan Strait, 640 fms. Also Guadeloupe. 


peripla Dall, Bull. M. C. Z., 9, p. 68, Aug., 1881; (as Drzllza) ; 


It) [Oy SES OL MIM Eas MyMac/y ) oxoy>.JeibUUL, Ziaial hoy. Oe OLlANcrg, st) 107), 
1889. 


73 MOLLUSCA OF THE GULF OF MEXICO 42 


chyta Watson, Jour. Linn. Soc., 15, p. 466, Nov., 1881 (as 
Defrancia); Rept. Chall. Gastr., p. 358, pl. 18, f. 4, 1885. 


Distribution.—Vucatan Strait, 640 fms. 


elusiva Dall, Bull. M. C. Z., 9, p. p. 69, Aug., 1881, (as Drillia); 
TOUS ply P27 SOU 27 OU Sa NM Ds TO2, 
leant 7. SO) 
Distribution.—Yucatan Strait, 640 fms. Perhaps identical 
with Defrancia perpauxilla Watson. 


bandella Dall, Bull. M. C. Z., 9, p. 59, 1881; Verrill, Tr. Conn. 
Neada OND 250) 10457 Dalle BullvomMin CH eZsSh ype Glo mepll. 
Romig Son Bulla. Us oo Ney MEY pa ro2t) plano sien p ls 
Govt 73.80. 
Pleurotomella diomede@ Verrill and Smith, Tr. Conn. Acad., 
Opals 2, pli zi ht.) 5, 5a boda. 


Distribution.—East coast United States and Gulf of Mexico, 
Blake Station 47, at 321 fms, Greatest depth 2100 fms. 


comatotropis Dall, Bull. M.C. Z., 9, p. 58, 1881; Verrill, Tr. 
Conme Acad. p.452, 17°82; Dall sBallMe C2) 75.18) ps1 bo, 
ples 8o7 NB ull) 27) Un Sa Ne ME p. 102, ply nity L2, 
Plait. 8. plO1mi i 77) (89: Johnson, Occ! Pal) Bost: Sec:, 
NEVE 7. (P- 1405, TOLS. 

tiara Watson, Jour. Linn. Soc., p. 440, 1881; Chall. Gastr., 
Pp. 347, pl. 21, f. 7, °85. 

Taranis pulchella Verrill, ’82 and 84. 
Distribution.—Massachussetts to Barbados, 50-1075 fms. 
Also Yucatan Strait, 640 fms. 

Helagiaywal Bull MNCZ. 9, p 1Or, Lost, 1S,1p. (uly, Plv nL, 1. 
Oso Bally 277-14 So NEN pO 2esple tn falaynt SO): 


Distribution.—Gulf of Mexico, Blake Station 44, in 539 fms., 
bottom temperature 39° F. Also Antillean. 


43 BULLETIN 38 73 


exsculpita Watson, Jour. Linn. Soc., p. 247, 1882, (as Drillia); 
Chall 'Gastri py amy Pin aqetbe Nisin (asiG@/orela) Dall 
Bulli Me CY Zr Shon plains on Sor Maula ah mos 
PLY TS VON oo. 
Distribution.—Vucatan Strait, 640 fms. Also Santa Cruz, 
Wie 

subsida’ Dall, Bull) M:C..Z., 0, p. 62; 18815 18) py 108, pliin2.t 
3h (SOs bBulliage) Uli S uN Men pato2 plu i2nit ena inSOE 


Distribution.—Gulf of Mexico, West of Florida, 339 fms. 
Also Cuba. 

para Dall Balle Ni CZ on) oa WSS Sil: eS) Nope i sols si pke una 
1889. 
Distribution.—VYucatan Strait, 640 fms. Also Martinique, 
502 fms, 

Genus CYTHARELLA Monterosato 

cymeila Dall, Bull. M. C. Z., 18, pl. 12, f. 4, 1889, (as Cythara) ; 

Bolla 7 Wii S.. IN ME pu oot plain A So. 


Distribution.—Gulf of Mexico, 220 fms. Also Antillean. 
galvestonensis Harris, Bull. Am. Pal., vol. 1, p. 100, 1895. 


Distridbution.—Upper Miocene, Galveston well, at 2236- 
2871 feet. 
Genus DAPHNELLA Hinds 
jeucophlegma Dall, Bull. M. C. Z., 9, p. 70, 1881; 18, p. 102, pl. 
Cito wis sou 
Distribution.—Gulf of Mexico, Blake Station 2, in 805 fms. 
(Eubela) limacina Dall, Bull. M. C. Z., 9, p. 55, Aug., 1881; Ver- 
Tl ire Conn: VA caddis Nip uuias eMs2. NON Dl 205 O4 mall: 
Ball Mi CC.) 22.118) p./106;plion inion So.) Bulle) 27.V Oa osuINe 
Val LOOMUD LO it ON sop 
hormophora Watson, Jour. Linn. Soc., 15, p. 457, Nov. 1881; 
Challis Gastra. (pis 5a) polyoma nines: 


74 MOoLLUSCA OF THE GULF OF MEXICO 44 


Distribution.—Massachussetts to Pernambuco. Gulf of 
Mexico, Yucatan Strait, 640 fms. and Blake Station 2, west 
of Florida, at 895 fms. 


Genus CANCELLARIA Lamarck 


reticulata Linnzeus, Syst. Nat., ed. XII, p. 1190, 1767, (as 
Voluta); Sowerby, Thesaurus, 2, p. 442, pl. 92, f. 17, 1848; 
iieyonn Wan.))7,p. 69, pl.) 2))t,.25, 26,853, Dall) Bull: Me: 
C48 sp. 129,\ 89.) Vanatta; hr. AYN |S.) Piila.) 155, p: 
757; 1903. 
Distribution.—Hatteras to Guadeloupe, 5-30 fms. Gulf 
coast: Crooked Isl., Charlotte Harbor and Sarasota Bay, 
West Fla.; Chandeleurs, La.; Galveston, Tex. Pleistocene, 
New Orleans Pumping Station No, 7. 


Note.—The fossil species, C. conradiana Dall, (Tr. Wagner 
Inst., 3, p. 42, 1890), is closely allied to C. 7efculata. 

galvestonensis Harris, Bull. Am. Pal., vol. 1, p. 100, pl. 3, f. 11, 
1895. 

Distribution.—Upper Miocene, Galveston well at 2552-2600 
feet. 

(Norona) clavatula Sowerby, Pr. Zool. Soc,, p. 52, 1832; Conch. 
Meee 2h a2-wlinyon) i Mian si 7 pale 75a ipl -hAs uy tenOds Vso: 
Inlanris.) Bulls (Am) (Paly vol!) tepeoon os; Wall) Bn U.S. 
INN 3375 (200, LO LO. 


Distribution.—Recent, Panama to Paita, Peru. Upper Mio- 
cene, Galveston well at 2552-2600 feet, (Harris). Notin 
the recent Atlantic fauna. 


(Trigonostoma) agassizi Dall, Bull. M. C Z., 18, p. 130, pl. 35, 
ME MMSSO Mt o7eW. Ss IN MEY pu monmipley ai5iih.) auc 89: 


Distribution.—Cape Lookout, N.C., 18 fms. Gulf of Mex- 
ico between Mississippi delta and Cedar Keys, 25 fms. 


(Trigonostoma) tenera Philippi, Zeitsch. fur. Mal., 5. p. 24, °48; 


A5 BULLETIN 38 73 


Dall, Bulla 7 Us Sane Ny Mom onsnS ori dlie NN ile OE a Eine 


43, 1890. 
stimpsont Calkins, Pr. Dav. A. S., 2, p. 250, pl. 8,£. 4, 5, 1878: 


Distribution.—Gulf of Mexico, Yucatan region. Pliocene, 
Caloosahatchie beds, Fla. 


Genus ABDMETE Kroyer 


microscopica Dall, Bull. M. C. Z., 18, p. 131, 1889,(as 7rigon- 
ostoma sd). Bull a7. U S) IN Mes patos 1 OOnwe i Oo neNe 
NEV 24 pA sO4.) plii2o ut {Anat @@2) 
Distribution.—Campeche Bank, 200 fms. Also off Fernan- 
dina and Cuba. 
Genus OLIVA Martyn 


circinata Marrat, Sowerby’s Thes. Conch., 4, Oliva, p. 21, pl. 17, 
f. 277, 1880; Johnson, Nautilus, 24, p. 123, Mar., 1911. 


litterata Lamarck, Ann. du Mus., 16, p. 315, 1810; Not of Bol- 
tem 1708. | Hilgard) House) or Repyik x MDoc) 1) pts 2M. 
S8ori78)) Dall wBulls 27. Uae SupiNeaev lenin oO planation 
$a)) 89; Singley, 4th) Ann) dReptamexa GS.) p. sacung2: 
Wanatta,)Pr. A. Ni S) Phila og or sloos. 

carvolimensts Conrad,’ Pr. A. Ne Swieiila 4s ip, 563,203: 


Distribution. —Hatteras to the West Indies, 0-2 fms. Gulf 
coast: Franklin and Calhoun Counties, Fla., Chandeleurs, 
Point on Fer, Cameron, La.; Galveston, Corpus Christi, 
Matagorda, Tex. Pleistocene, New Orleans artesian well of 
1856, Lake Borgne borings, Knapp’s wells, Terrebonne 
Parish, No. 1 well at 2000-2450, No. 2 at 1780-1800, No. 3 
at 1150-2029 feet. Pliocene, Caloosahatchie beds, Fla. 


liodes Dall, Tr. Wagner Inst. Sci., 3, p. 1576, pl. 58, 1903. 


Distribution.—Lower Miocene, Chipola marl, West Florida; 
Bascom No. 2 well, Mobile, Alabama, at 1241 feet, Chipola 
horizon. 


76 MOLLUSCA OF THE GULF OF MEXICO 46 


Genus OLIVELLA Swainson 


mutica Say, Jour. A. N.S. Phila., 2, p. 228, 1822; Hilgard, Rept. 
Chief Engineers to Sec. War, p. 358, ’70, House of Rep., 
EomWOCle pt 2) pi Sho, 78; Wally Pry Wiis. NG Me, p: 
B24 Se) Bull e327) UL Se Ne ME pp. \1o6)) pl ee ee OS OF 
eV AMeN So0.4\5,)) 290%) Vanattay) Pr, A. ING) S. Pinta. 55), 
Pavi571 1903; Deussen, U2) S: Gs S:) Water- eine es 
INOW ZC5) (D:77; Told. 
Distribution.—North Carolina to Trinidad Isl. Gulf coast: 
Recent, Cedar Keys, St. Mark’s and St. Joseph’s Bay, Fla.; 
. Galveston and Corpus Christi, Tex. Pleistocene, New Or- 
leans artesian well of 1856, Lake Borgne borings; Knapp’s 
wells, Terrebonne Parish, No. 1 at 1600-1700, No. 2 at 1050- 
1842, No. 3 at 1150-1839 feet; North Creek, Manatee and 
Labelle, Fla. Pliocene, Caloosahatchie beds, Fla., and New 
Orleans Gymnasium well at 1200 feet. Miocene, Gilbert 
well No. 10, Bateson, Tex. at 323 feet. 
pusilla Marrat, Thes. Conch., t. 21, f. 356-358, 71; Tryon, Man. 
Wonchipes.op.1645) ply gent. 45) he Seni Vanattas bi ANG 
Seaelailane SS. 171575) 203. 
Disiribution.—Cited by Vanatta from St. Joseph’s Bay and 
Crooked Isl., West Fla. 
nivea Gmelin, Linn. Syst. Nat., ed. XIEI5-p. 3442, 1792; Reeve, 
Sonchaylcony, it 640a, 150; Tryon, Manis) pe 67) pl. 15, f 
fences Dall Bill a7) UW. 3. Ns (Meaiipe roo, So. Dall Sz 
Simpson, Bull. U.S. Kish Com., 1, p.) 393) or. 
Distribution.—Sarasota, West Fla. to Haiti, 27-805 fms. 
jaspidea Gmelin, Syst. Nat.,ed. 13,p. 3442, 1792; Reeve, Conch. 
icone 6, p:)22, 0. 58,1850: Dall & Simpson, Bull, WU), S.) Fish 
Comes) ph 202, ‘Toor. 
Distribution.—Hatteras to Brazil and West Florida, 27-805 
fms. 
floralia Duclos, in Chenu, Ill. Conch., p.6; Tryon, Man., 5, p. 


47 BULLETIN 38 77 


OS plein.) fh: / 2. an ba SS a aallllmin Stilley) aru aS upIN emy os Do 
106, ’8o. 
Distribution.—Hatteras to Tortola and West Florida. 
eulacta) Dall, Tr.) Wagner) Inst.) Seis) 3). 5 76)) pola ison tage 
£903. 
Distribution.—Lower Miocene, Chipola marl, Fla, and Mo- 
bile Oil Co’s No. 2 Bascom well near Mobile, Alabama, at 
1241 feet, Chipola horizon. 
galvestonensis Harris, Bull. Am. Pal., vol. 1, p. roo, pl. 3, f. 13, 
1895. 
Distribution.—Upper Miocene, Galveston well at 2410-2871 
feet! 
subtexana Harris, Bull. Am. Pal., vol. 1, p. 101, pl. 4, f. 1, 
1895. 
Distribution.—Upper Miocene, Galveston well at 2552-2871 
feet 


Genus MARGINELLA Lamarck 


cared otorer, Jour. Bost))\Soc) .NEiT ps AG5, plionit 2A 
1837; Sowerby, Thes., f. 103;) Tryon, Man., 5, p. 33, pl. 10, 
P86; 1882.) Dall: Bull srw OE SNe Vien paytOoN SO: 


Distribution.—Charlotte Harbor, West Fla., Florida Keys 
and Antilles. 


storeria Couthouy, Jour. Bost. Soc. N. H., 1, p. 440, pl. 9,f. 1,2, 
18373) Dall Ball 37. SUL Sain ven: OOS 89: 


Distribution.—Gulf of Mexico, West Fla., to Colon. 
cassis Dall, Bull) (M,C. Zin Sexpime7in ply 35, \\\t../ 8 1889. ule 
BTM TOO. Dl 2 Shit owas 


Distribution.—Gulf of Mexico, Lat.25° N., Lon. 84° W., at 
tol fms. Also off Cuba. 


apicinca Menke, Syn. Meth. Moll., p. 87, 1828; Tryon, Man., 5, 
DA BS,ipl. 10, ef. So) 28a. yDalleiB all se S46 ING Mo aTOos 


78 MoLLUSCA OF THE GULF OF MEXICO 48 


"89; Singley, ath Ann. Rept. Tex., p. 333, 92; Vanatta, 
PrivA. N.S) Phila’, 55, p. 757, 1903; Vaughan, 2nd) Ann. 
Rept. Fla..Geol. Surv., p. 148, 1909. 


conoitdalis Reeve, Conch. Icon., 15, pl. 18, f. 87. 
Distribution.—Hatteras to Jamaica, Gulf coast: West Fla. 
and Texas, Pleistocene, Manatee, Orient and Labelle, West 
Florida. 

pellucida Pfeiffer, Wiegmann’s Archiv; 1, p. 258, 1840; Tryon, 
Mansi pl aa pl.) TO fon og mDall bull ian. ss. 
ME pa LOOM NSO: 
Distribution.—Sarasota, West Fla. and West Indies. 

watsoni Dall, Bull. M. C. Z., 9, p, 71, 1881; 18, p. 137, pl. 19, f. 
Bplansss by 2486" Bull: 27. Wis wINen Mey pu OG spleToiHt: 
Bullen 28h, fein 18 Oe 
Distribution.—Yucatan Strait, 640 fms. Also off Havana, 
220-805 fms. 

virginiana Conrad, Am. Jour. Conch., 4, p. 67, pl. 5, f. 4, 1868, 
(as Prunum); Dall, Bull. 37, U. S. N. M., p. 106, ’89; Tr. 
NV Salen Seizes 43.) 190} 
Distribution.—Recent, North Carolina to Yucatan, 10-294 
fms. Pliocene, Caloosahatchie beds, Fla. Miocene, James 
River, Va. 

denticulata Conrad, Jour. A. N. S. Phila., 6, p. 225, pl. 9, f. 21, 
u83o0.) | @Not of Late, /73); Dall) Bullsiay iW! S.)) N° IMs, p: 
OSM SOs: WV LS. (35 Pp. Sisuplsnsi ty ommrooo: 
Distribution.—Recent, Hatteras to Barbados. Also West 
Florida, 5-294 fms. Pliocene, Caloosahatchie beds, Upper 
Miocene, Maryland and Virginia. 

opalina Stearns, Pr. Bost. Soc. N. H., 15, p. 21, 1872; Dall, Pr. 
CN SwNE MS 6ap els 24. 183) Bullea7ipe TOS." 8Q,Cas\ acre 
ticulata var. opalina). 


Distribution.—Hatteras to Barbados. Recent, Tampa (type 


49 BULLETIN 38 7S 


locality) and Cedar Keys, 14 fms. Also Barbados, 100 fms. 


aureocincta Stearns, Pr. Bost. Soc. N. H., 15, p. 22, 1872; Dall, 
Balle 27.1. SON Mie apr rsh iSOe arly Sea ONS 2 
1890. 


Distribution.—Virginia to West Florida, 3-44 fms. Pliocene, 
Caloosahatchie beds. 


minuta Pfeiffer, Wiegmann’s Archiv. fur Naturg.1, p. 259, 1840; 
Dryon, Mans 5.) p. 43), pl. 12h i oowmmeano Dalla bull 27 mr 
SHINGO. O81 SO! 


Distribution.—Mediterranean, West Indies, East and West 
Florida, 5 -294 fms. Miocene to Recent. Also Pleistocene, 
southern Europe and of North Creek, West Fla. 


minima Guilding, Sowerby, Thes. Conch., 1, p. 388, pl. 78, f. 
220u18a6- Wall Pre U. /Si Naive toMpa 22464) 182i tayore 
Mans .p: 44, pl. 12) £./62 8a nDallienBullea en sS: 0 Ni vier 
p. 108,’ 80. 

Distribution.—Cape Lookout, N. C., to Haiti. Also West 
Florida at Cedar Keys, 0-22 fms. 


Note.—Tryon placed this species in synonymy of mzz2uta 
Pfr. According to Moerch mznzma is asynonym of lavalleana 
d’Orbigny, (Moll. Cuba, 2, p. rot, pl. 20, f. 36-38, 1842). 

succinea Conrad, Pr. A. N.S) Philavi26.) pl. 1, f- 17, 1840; 
(iryon. Man. 5,'p. 34, pli iromnmogmcss, Dall) Bull: 27 -aUs 
SAONEYMES ps 108, ple no, fs 64 soy 


Distribution.—Fernandina, East Fla., to Sombrero, W. I. 
Gulf coast at Tampa, 70-1002 fms. Placed by Tryon in 
synonymy with zztzda Hinds. 

fusinayDall, Bull. M. C. Zo p72 ir88r: 18. p. 138. pla toy t: 
ANS: Mati. 27)\p. (TOON PION n SO, 


Distribution.—Yucatan Strait, 640 fms. Also off Fernan- 
dina, Fla. 


Bo MOLLUSCA OF THE GULF OF MEXICO xe 


yucatecana Dall, Bull. M. C. Z., 9, p. 72, 1881; 18, p. 138, pl. 
NOM USHE SOV BUlla OK S20 NV py mOoN pI TOR fis, 
1889. 


Distribution.—VYucatan Strait, 640 fms. Also Sand Key, 
Rilaen 25) bas: 


seminula Dall, Bull. M. C. ZO puT Za) USSU LSe iP) HO, (Oke LO), 
Hen So Bull. 27) U.S. Neve py los. pl Loli, 25.789. 


Distribution.—VYucatan Strait, 640 fms. Also Antillean. 


rostrata Redfield, Cat. Marg., Am. Jour. Conch., 6, p. 246, 
@iotelwirs 70. Dall Bull Me CZ. 1 8)ypuitia7 | 805 re) Wie 
SA ahs 5O,k SOO: 
oblonga Sowerby, Thesaurus, pl. 76, f. 106, 108. j0N 
oblonga of Swainson. 


Distribution.—Gulf of Mexico, Lat. 23° N., Lon. 89° W., 
in 84 fms. Pliocene, Caloosahatchie beds. 


(Volvarina) avena Valenciennes, in Kiener, Coq. Viv. arginella, 
PW plow tl. 24.) 1841: Sowerby, WaeswConchyi1) p, sor, 
DEO te1Zo0, 49>) Dall) Bull) 27) Ws Suen Me ps) 108; 180: 
Dall & Simpson, Bull. U. S. Fish Com., 1, p. 394, 1901; 
Wauchans Publ. No. 133, Carn. inst pez 1910: 


Distribution.— Key West to Colon. Also Texas region, 
Gulf of Mexico. Pliocene to Recent. Pleistocene, La- 
bella, West Fla. 


(Volutella) ovuliformis d’Orbigny, Moll. Cuba. 2, p. 101, pl. 20, 
feasts. 6845 Dall Bull 37.) Usa Say NGmMiNs {ip TOS... 289; 
Dallesa Simpson Bull Wes) hishh) Comets n)31'395,) Look: 
Vaughan, 2nd Ann. Rept. Fla. G. 5., p. 148, I910. 


Distribution.—Cape Fear, N. C., to Guadeloupe Isl. Also 
West Florida. Pleistocene, Manatee, West Fla. 


(Volutella) hadria Dall, Bull. M. C. Z., 18, p. 142, 1889; Bull.37, 
Oe SONG MEN pastoS.))7380, 


SI BULLETIN 38 $x 


lacrimula Dall, Pr. U. S. N. M., 6, p. 324, 1883. Not lacrimula 
Goulds) Pr. Bost. Soe Ni El) Si pi 2dn L8O2: 


Distribution.—Cedar Keys, West Fla. Collected by Hemp- 
hill on mud flats. Also Charlotte Harbor, West Fla. 


Genus VOLUTA Linnzeus 


virescens Solander, Dillw. Desc. Cat., 1, p. 562, Sowerby, Thes. 
Lplis2) fH 78 Ey Ony Mane Aa PanOqwph 2a uiiaa sn ieSZe 
Dally Balli37. Us So ING Me panos Son 
Distribution.—Texas to Cartagena, Colombia. 


Genus SCAPHELLA Swainson 
junonia Hwass, Chemnitz, Conch. Cab. 11, p. 16, pl. 177, f. 1703, 
L7 Oar pip wr 7Os5s Dall Bal VCH ZT Sak. AS. pl aann te 
5ae-en SO; Bulls 27. (Ul S NN A Men Onno sla gau sts Cem 
1889. 
Distribution.—North Carolina to Nassau. Also Gulf of 
Mexico!) Wat! 25° Ni. Lonij82-. Wat 26) fms: 


Genus AURINIA H. and A. Adams 


dubia Broderip, Zool) \Jour.,) gipe wemupliai ft.) 1, 1828)).(as 
Voluta); Reeve, Conch. Icon., Voluta, pl. 22, f. 59, 1849; 
Tryon, Man., 4, p. 90, pl. 27, f. 77, 81, 1882; Dohrn, Jahrb. 
Mal! Ges.;)'6,,p: 150, pk. ait eaeai 7G Dall) Bull. Mi CZ 
CS Ps USL. SOs BU 27. KU Eisai NNO. TTO;) Sore ra Wase 
NE 24D. Soa.) pli zoutiinramoo2) 


Frusus tesellatus Schubert & Wagner, Suppl. Bd. Mart. und 
Chemn., Conch. Cab., 1829; Kiener and Reeve; but not of 
Zekeli and Pictet, 1852. 

mutabilis ‘Tuomey & Holmes, 1856. Not of Conrad, 1841. 
Distribution.—Hatteras to Cape Florida. Gulf of Mexico 
between Mississippi delta and Cedar Keys, at 111 fms. 

ropusta Dall Bally My CZ yrs apt 53, ples, f) 24)’ so. Bulle 
BT De LLO Mp.) 250 Lua sOE 


82 MOLLUSCA OF THE GULF OF MEXICO 52 


Distribution.—Gulf of Mexico, Lat. 28° N., Lon. 86° W., 
at 280 fms. Also off Cuba, 119 fms. 


Genus MITRA Lamarck 

gemmata Sowerby, Thes. Conch., sp. 334, 1874; Tryon, Man., 
AMM Too Plu 5S) toss ross Oa Bully, 277 Ui SiN My 
p. 110, ’89; (as var. of hkanleyz Dohrn): Dall & Simpson, 
Balle WS) ish Com\, 0,’ pi 396) reer. 
Distribution.—Charlotte Harbor, West Florida, to Jamaica. 

straminea A. Adams, Pr. Zool. Soc., p. 132, 1851; Sowerby, 
dthess Conch. t..561; Tryon, Many, 4. wae) pl. ar, ft. 183, 
HSS OA sBull 7. Ui. 95) Nao Meio hO: .8Or 
Distribution.—Gulf of Mexico, west of Florida. 

wandoensis Holmes, Post-Plioc. Fos. S. Car., p. 77, pl. 12, f. 10, 

10a, 1860, (as Volutomitya); Dall. Bull. 37, U. S. N. M.,p. 
LOMAS OMT NY olla)... 34) 9! O21 LSGe, 
Distribution.—Cape Hatteras to Florida Straits and West 
Florida, 14-80 fms. Very closely allied to and probably 
identical with J7. rusht Dall, (Conch. Exch., 2. p. 9, 1887; 
Bull. M. C. Z., 18, p. 160, ’89) from the Gulf of Mexico, 27 
fms., U. S. Fish Com. Station 2372. 

SiviiaeWallk Bull Mi CiiZ.. 18, Pp. 150) plenmse eon 1880 Bull, 
BRO ONG Me ips Tro) plu Tes dOmses 
Distribution.—Gulf of Mexico, west of Florida, 19 fms. 
Also Antillean, 

(Conomitra) blakeana Dall, Bull. M.C. Z., 18, p. 163, 1889; Bull. 
By On Sa Nine Mere Ds Isto}, 80) 

Distribution.—Recent, Yucatan Strait, 640 fms. Closely 
related to the fossil species, stamzzea Conrad and fusoides 
Wear 

Genus MITROMORPHA A. Adams 


pygmaea Dall, Tr. Wagner Inst. Sci., 3, p. 95, pl. 10, f. 3, 1890. 


53 BULLETIN 38 83 


Distribution.—Type from Caloosahatchie marl, Fla., Plio- 
cene. Also Bascom No.1 well, near Mobile, Alabama, at 
1500-1556 feet. Miocene. Not in the Recent fauna. 


Genus FASCISLARIA Lamarck 


gigantea Kiener. Icon. Coq. Viv., p. 5. pl. 10, 11, 1840; Tryon, 
Man 2. p75; (pls GO. tama Slam atl eb unl Ieee sy IN eave 
PMT? Sora NV LS.) 2p mon OoOu time leven tiny Acmns 
Rept: Tex. ’G.'S.;)p. 234,92; Walliéz/Simpson, Bull) U27s: 
Fish: Com.))1,\p)207,)) Teo Waughaniiadi Ann. Rept.vElas 
G. 3., p. 149, 1909; Johnson, Nautilus, p. 44, Oct., 1919. 


Distribution.—Hatteras to Brazil. o-1Io fms. Gulf coast; 
West Florida; Point au Fer. La.; Corpus Christi and Mat- 
agorda Isl., Tex. Pleistocene, New Orleans pumping Sta- 
tion No. 7, North Creek and Manatee, West Fla. Pliocene, 
Caloosahatchie River marl. 


tulipa Linneeus, Syst. Nat. ed X, p. 754, 1758, (as Murex); 
Kienenr. Cog) Vive, ps2) pla mz s Ao.) dvnyon\) Manis eau: 
FAN DIAS ON te hon) ron) Oa MeN iS oe si. SOTA IUN Alte 
iGO; Nanatta, SPraial uN e Samaalan nis 5) Duis Tal OOgr 
Johnson, Nautilus, p. 46, Oct., 1919. 


Distribution.—North Carolina to Colombia and Antilles.o-10 
fms. Gulf coast: Cedar Keys, St. Joseph’s Bay and St. 
Marks, West Fla., Texas coast. Pleistocene, North Creek, 
Fla. Pliocene, Caloosahatchie beds. 


distans Lamarck, An. s. Vert., 7, p.119, 1822; Tuomey & Holmes, 
PLoS SCH ph 150 plesowena tos 7) iiloardy) Houselok 
Reps ix) MOC.) 1, peep esomenonn Dall, lic Well Seaicnmny 
102, pl. 7, f. 10, ’90; Singley, 4th Ann. Rept. Geol. Surv. 
Tex.,'p. 334, ’92; Mitchell,List. Tex. Sh., p. 16; Vaughan, 
Publ. No. 133, Carn. Inst., p. 171, I910; Johnson, Nautilus, 
DN47 NOCE TOTO) 
Distribution.—Hatteras to Yucatan o-54 fms. Gulf coast: 
Cedar Keys, St, Marks, Fla.; Cameron and Chandeleurs, 


54 MOLLUSCA OF THE GULF OF MEXICO 54 


La.; Galveston, Matagorda, Corpus Christi, and Mustang 
Isl., Texas. Pleistocene, New Orleans well of 1856, and 
pumping station No.7; Orient and Labelle, West Fla. 
‘The Pliocene shell of the Caloosahatchie beds is the related, 
but slenderer, /. apicina Dall. 


(Mesorhytis) meekiana Dall, Bull. M. C. Z., 18, p. 172, pl. 36, f. 
Fin WSSOS) EUG oe Oo IN IE soy, si shzA iol aXe). aha ale cXG), 
Distribution.—Gulf of Mexico, Blake Stations 16 and 20, at 
292 and 220 fms. Also off Cuba, 400 fms. ‘The first living 
species of Mesorhytis known, the type being the Cretaceous 
species, gvacilentts Meek. 

Genus FUSINUS Rafinesque 

timessus Dall, Bull. M. C. Z., 18, p. 166, 1889; (as Fuses); Bull. 

COE NEE ep 112.02 89. 


Distribution.—Gulf of Mexico, between Mississippi delta 
and Cedar Keys, 60 fms.; between Cedar Keys and Dry 
Tortugas, 27 fms. Also near Cuba. 

eucosmius Dall Bull MeeC. Z)) 18) pit 67 plans ,it.) 5, 18805 
IBLUDLS B75 \Oy sha FON eae Ns eluls cshay, 
Distribution.—Gulf of Mexico, west of Florida. Also Bar- 
bados, 27-111 fms. 

couel Petit, Journ de Conch, 4, pr 2torsplly 8) fim) 18535) Dall, 
Balle Ne CZ e168) ps 167, 789) 
Distribution.—Gulf of Mexico, between Tampa and Dry 
Tortugas, 26 fms. 

alcimus Dall, Bull. M. C. Z., 18, p. 170, 1889. 
Distribution.—Gulf of Mexico, 100 miles north of Yucatan, 
g5 fms. 

amiantus Dall, Bull. M. C. Z., 18, p. 169, pl. 15, f. 11, 1889; Bull. 
Boy NUIG Sio AN OM GAN ove esi zae Wo) Me ell/adids Uhksyay, 


Distribution.—Gulf of Mexico, Blake station 2, at 805 fms. 


O15) BULLETIN 38 85 


aepynotus Dall, Bull. M. C. Z., 18, p. 169, 1889. 
Distribution.—Gulf of Mexico, 84 fms. Also Antillean. 
amphiurgus Dall, Bull. M. C. Z., 18, p. 171, 1889. 
Distribution.—Gulf of Mexico, Blake station 45, at ror fms. 
Genus LEUCOZONIA Gray 


cingulifera Lamarck, 7, p. 107, 1822, (as Zurbinella); Reeve, 
Conch. Icon., 4, pl. 3, f. 17, 1847; Dall and Simpson, Buil. 
We) Ss) hish? Come. 1) 7p: 208) r901 


Distribution.—Sarasota, Fla., to Texas and Brazil. Also 
Antillean. 


ocellata Gmelin, Syst. Nat., p. 3488, 1792, (as Buccinum) ;Reeve, 
Conch: Icon.; 4, pl. 8,°f 28, 18475) Dall and Simpson; Bull? 
Weis wish Com:;) 1; p.208, gor 


Distribution.—Cedar Keys, Fla., to Guadeloupe, W. I. 
Genus BUSYCON Bolten 


pyrum Dillwyn, Desc. Cat. Rec. Sh., 1, p. 485, 1817; Dall, Bull. 
Bre Wise ON Mp) 2 Sonia bei Md Sil c2\ Tse TOO Or 
Singley, 4th Ann. Rept. G. Sex) \p. 334,92; Vanatta, 
Pre VAL ING S,) Phila p. 757 roos i aughan: : ulblamnacs 
Carn, Inst., p. 171, 19Io. 


Distribution.—Hatteras to Gulf of Mexico. Upper Miocene 
to Recent. Gulf coast: Cedar Keys, Fort Barranca, St. 
Mark’s, Fla.; Cameron, &c. La.; Galveston, Matagorda, 
Corpus Christi, &c., Texas. Pleistocene of Louisiana at 
Grand Chenier and New Orleans pumping station No. 7. 
A high spired variety isfound in abundance at Cedar Keys, 
Point au Fer, Cameron and Galveston, and in the Pleistocene 
of Grand Chenier, La., North Creek, Labelle, Manatee and 
Orient, West Fla. 


canaliculatum Linnzeus, Syst. Nat., p. 753, 1758, (as Wurex); 
iolmes,Post- Pl) Hos) Si) Chip) /66.;npluann his) 18 5oe ale 


86 


MOLLUSCA OF THE GULF OF MEXICO 56 


gaid, House of Rep., Ex. Doe., 1, pt. 2, p, 886, 78; Dall, 
Balke OL Sua Meni pnt? pla 7 akin toon Clark Plers- 
tocene Md., p. 180,pls. 46, 47, 48, 1906; Johnson, Occ. Pa. 
BOSH SOC NE Ee i7,.) Dal sOQninOiliss 


Distribution.—Cape Cod to West Florida and Texas. 
Pliocene (Carolinas). Pleistocene, New Orleans well of © 
1856 (Hilgard). 


perversum Linnzeus, Syst. Nat., p. 1222, 1766, (as Murex); 


ditionmey,&) Elolmes, Pl. Kosi 'SCiipy 145; pl. 204 f: 3.1856; 
Hilgard, House of Rep. Ex. Doc. 1, pt. 2, p. 886, ’78; Dall, 
ree S38 0)) 116,\\/o07)) Vanatta wer ay NES. bhilas, 
SDaTsie Loos N auchant PubitinegatCarnainst sn pi ij, 
IQIO. 


Distribution.—Hatteras to Cuba. Miocene (Carolinas) to Re- 
cent, o-3 fms. Gulf coast: Cedar Keys, St. Marks, &c., 
Fla.; Point au Fer, Cameron, La.; Galveston and Corpus 
Christi, Tex. Exceedingly abundant in the Pleistocene of 
Grand Chenier, La., and found by Hilgard in the Lake Bor- 
gue borings; also of Manatee, Orient, and Labelle, West Fla. ; 
Pliocene, Caloosahatchie beds, Fla. 


carica Gmelin, Syst. Nat., p. 3545, 1792, (as Wurex); Holmes, 


RostsPiihos, 5). Cy p65, pli 11, Pemaso. Dall Bullies, 
NEE Ma con 2) olla. £1.) 7897). Clarkebletstocene ss Mid= jp: 
179, pls. 43-45, 1906; Johnson-—Oec. Pa. Bost. Soc. N. H., 
7) P- 139, 1915. 

Distribution.—Massachussetts to the Antilles. o-10 fms. 
Gulf coast, West Florida and Galveston, Tex. 


eliceans Montfort, Conch. Syst. 303; Dall, Bull. 37, U. S. N. M., 


Os, BE, SO) 
Distribution.—South Carolina to Campeche. o-5 fms. 
Genus MELONGENA Schumacher 


corena Gmelin, Syst. Nat., p. 3552, 1768; Tryon, Man., 3, p. 


LOSMAD OI 1OD-2O2 18Sry Dalla Mla Seige 12 TOO); 


57 BULLETIN 38 87 


Vanatta,Pr Au iN) SS) Rinilal iss). 7757.) ko033) Vatenany 2k 
Ann. Rept. G. S. Fla., p. 148, I909. 


Distribution.—Florida Keys to Yucatan and the Antilles. 
Gulf coast: Cedar Keys, St. Marks, St. Joseph Bay, Fla.; 
La.; and Tex. Pleistocene, North Creek, Orient and La- 
belle, West Fla. 


melongena Linnzeus, Syst. Nat. ed XII., p. 1220; Tryon, Man., 
Bu PLO7s Platt. LOZ. TOS moon Dallumb illus 7A: Sie Nte 
Me peirr2)). 805) Harris Bulli AqM ealeavole i lipsnt Or nOuys 


Distribution.—Florida to Grenada. o-50 fms. Gulf of 
Mexico, Texas region. Also Upper Miocene, Galveston 
well at 2448-2465 feet (Harris). 


Genus CANTHARUS Bolten 


tinctus Conrad, Pr. A. N. S. Phila., p. 25, pl. 1, f. 9, 1846, (as 
Pola) Dall) Bull. 27 Us SaNeRMeNn point oy So) Case fizz 
tonidea); Dall and Simpson, Bull. U. S. Fish Com., 1, p. 
399, 190T. 


Distribution.—Hatteras to Vera Cruz. Gulf coast: Cedar 
Keys, Fla., and Matagorda, Texas. Pleistocene, North 
Creek, West Fla. 


cancellaria Conrad, Pr. A. N.S. Phila., p. 25, pl. 1, f. 12, 1846; 
Dalry WL0S. IN) Mover comige mess omeley.” athiAcmr, 
Reptyvbexs G.'S.) p. /334wozsanaialbvaset. An) Ni...) elder 
Soup 574.1903; Vauchany 2daAnm Rept... bla. (GA Se iis: 
148, 1909. 
Distribution.—Florida to Isthmus of Panama. Gulf coast: 
Cedar Keys, Fla.; Horn Isl., Miss.; Cameron, Point au Fer, 
Chandeleurs, La.; Galveston, Tex. Pleistocene, Grand 


Chenier and New Orleans pumping station No. 7, and of 
Orient, West Fla. 


orbignyi Payraudeau, Cat. Moll. de Corse, p. 159, pl. 8, f. 4-6, 
To20\ (as Buccnun)|:) Dally Bulle 7 iiss Nis HVE spa menGs 


MoLLUSCA OF THE GULF OF MEXICO 58 


os) 
es) 


"89; Dall and Simpson, Bnll. U.S. Fish Com., 1. p. 4oo, 
IQOL. 
Distribution.—Florida to Yucatan. 22-25 fms. Also Medi- 
terranean. 

limbatus Philippi, Abbild, 1, 3, pl. 1, f. 9, Tryon, Man., 3, p. 
HO MPU M257 TOSd Dall wll ar UE Sen INi WOW i 
TOMS Oe 
Distribution.—Gulf of Mexico to Cuba. 

Genus PHOS Montfort 


eandeil d’Orbigny, Moll. Cuba, 2, p. 129, pl. 21, f. 23-25, 1845, 

(as Cancellaria); Arango, Fauna Mal. Cub., p. 201, 1878; 
Dall, Bull. 37, p. 116, 1889; Dall and Simpson, Bull. U. S. 
Fish Com., I, p. 401, 1goI. 
Distribution.—Hatteras to Colon and the Antilles. Gulf 
of Mexico, between Mississippi delta and Cedar Keys, at 88 
fms. Synonyms, fide Dall, are antillarum Petit, 1853 and 
grateloupensts Petit, 1853. 

parvus C. B. Adams, Contr. Conch., p. 59, 1850, (as 7rzton) ; 
Dall, Bull. 37, U. S. N. M,. p. 116; 1880. 

LLUGICALUS WA IPT IU S.IN. MES Gye 225Miply TO mio os. 
Distribution.—Charlotte Harbor, West Fla., and Texas. 
Also Antillean. 

galvestonensis Harris, Bull. Am. Pal. vol. 1, p. 102, pl. 4,f. 4, 5, 
1895. 

Distribution.—Upper Miocene, Galveston well at 2158-2871 
feet: 


Genus NASSARINA Dall 
columbellata Dall, Bull. M. C. Z., 18, p. 182, 1889. 
Distribution.—Cape Catoche, Yucatan. 


grayi Dall, Bull. M. C. Z., 18, p. 183, pl. 32, f. 12a, 1889; Bull. 
STO Se Ne MEE Di TIO) pln aoh aha i QeNee On 


59 BULLETIN 38 89 


Distribution.—Arrowsmith Bank, Yucatan, 130 fms. Also 
Antillean. 


bushi Dall, Bull MCA 1s. 182. oluusniayn 2) Uneso. 


Distribution.—Gulf of Mexico, Blake station 5, 152-229 fms. 
Also off Sand Key, Fia. and Barbados. 


Genus ALECTRION Montfort 
vibex Say, Jour. A. N. S. Phila., (1), 2, p. 231, 1822, (as WVassa); 
Ann} Coneh. pli 57, 10/25), Dalla ull iar aes Se eINMEe Go 
116, pl. 50, ¢; 8, 1889; Singley,athWwaAna Rept, Mex! GaSe 
Ps335, Fso2: Johnson, Occ: PayBosty socsiNi inl 7. ps nade 
IQI5. 

Winetens7s Perkins, Pr) Bost.) SoC. 1 3upy L741 SOO. 
Distribution.—Cape Cod to Colon. Gulf coast: Cedar 
Keys and St. Mark’s, Fla., Point au Fer, La., Galveston, 
Corpus Christi, Tex. Pleistocene, New Orleans pumping 
station No. 7; Manatee, Orient and Labelle, West Fla. 
Pliocene, Caloosahatchie. 

ambiguus Montagu, Test. Brit., pl. 9, f. 7, 1803, (as Buccinum); 

Dall, Bull. M. C. Z., 18, p. 184, ’89; Not amdigua Dunker 
(West African), which is zzcrassata Strom. 

alba Say, 1826; candidissima C. B. Adams, Krebs Cat., p. 32. 


Distribution.—North Carolina to West Florida and Barbados. 
Yucatan Strait, 640 fms. Variety antillarum d’Orb., Ca- 
loosahatchie Pliocene. 


acutus Say, Jour. A. N.S. Phila., 2. p. 234, 1822, (as Vassa); 
Holmes, -Post-Pl. Kos SiC mow 2hplen 12) 01.) 3. ut SOOM Cas 
Buccinum); Hilgard, House of Rep. Ex. Doc. 1, pt. 2, p. 
886.4757) Dall Bulli27 Wes umNn ME i pe TLON oom S™ 
Bully pA, | Pails voll pmo2nOsn wanattay peta aeiN wp 
Phila., 55, p. 757, 1903; Deussen, Water Supply Pa. 335, p. 
77, 1914. 
Distribution. —Carolinas to Barbados. Gulf coast: Recent, 
St. Joseph’s Bay and Crooked Isl., West Fla.;Point au Fer, 


go MOLLUSCA OF THE GULF OF MEXICO 60 


Cameron, La.; Galveston and Corpus Christi, Tex. Pleisto- 
cene, Knapp’s wells, Terrebonne Parish, No. 1 at 1600-2443 
feet, No. 2, 1050-1842, No. 3, 58-1839 feet, New Orleans 
ar.esian well of 1856, Lake Borgne borings. Pliocene, New 
Orleans Gymnasium Club well at 1200 feet. Upper Mio- 
cene, Galveston well, 2871 feet. Miocene, Gilbert well No. 
10, Bateson, Texas at 323 feet. 


consensus Ravenel, Pr. A. N.S. Phila., p. 43, 1861; Dall, Bull. 
VEC Z iS DATS 5s: Sor 


Distribution.—North Carolina to Florida. 8-49 fms. Gulf 
coast 20 miles off Charlotte Harbor, W. Fla., at 13 fms. 


hotessieri d’Orbigny, Moll. Cuba, 2, p. 142, 1845, Atlas, pl. 21, 
f. 40-42, (as Vassa); Dall, Bull. M. C. Z., 18, p. 185, 1889; 
Dally s) Simpson) Bull. U.S) Fish) Com) p. 402) 1901r: 


Distribution.— Gulf of Mexico, Blake station 36, at 84 fms. 
Also Hatteras and Sombrero, 30-85 fms. 


galvestonensis Harris, Bull. Am. Pal., vol. 1, p. ror, pl. 4, f. 2, 
1895. 


Distribution.—Upper Miocene, Galveston well at 2410-2871 
feet. 


trigalvestusiilaris: bull) Am?! Pal. volume) py mony iplaia. sp: 
3, 1895. 


Distribution.—Upper Miocene, Galveston well at 2158-2871 
feet. 


obsoletus Say, Jour. A. N. S. Phila., (1) 2, p, 232, 1822, (as 
Wassa) Gould, Inv: \Mass., pl. 15, f))210;), Dall) Bull: 37, 
UA Se ONGEMe pr nO) ply So. f you mesonmoluson\ Oce! Pax 
BOSts SOCHNE EE 74) Ds TA Shir Or ss Wal buble rere WSN). 
Mis en Oss LOZ. 


Distribution.—Nova Scotia to Tampa, West Fla. Also San 
Francisco Bay, (introduced with oysters from the Atlantic). 


61 BULLETIN 38 Or 


beaumontensis Aldrich, Bull. No. 1, Univ. Texas, July, 1901; 
Nautilus, p. 74 and figs., Nov. 1901, (as Wassa). 


Distribution.—Pleistocene. Spindle Top Hill oil well, near 
Beaumont, Texas, at 390 feet. 


Genus STROMBINA Moerch 


gibberuia var. galvestonensis Harris, Bull. Am. Pal,, vol. 1, p. 
103; pli iauifi6. 1895. 


Distribution.—Upper Miocene, Galveston well at 2410-2871 
feet 


Note—The typical form of gtbberula Sowerby, (Pr. Zool, 
Soc., p. 115, 1832), is recent on the Pacific coast of Central 
America southward to Paiti, Peru. 


Genus ANACHIS H. and A. Adams 


avara Say, Jour. A. N.S. Phila., 2, p. 230, 1882, (as Columbella) ; 
uomey and Holmes) Pl. skosiS.4@ap. 139) iphi28. i 12 
1357) Dall Bull. 37. Ui. Sa iNe wep sore) ple son ty mons o: 
NManatta, Pr.) (ALUN. S20 Philaswssenpsi574 £6035, Oonnsons 
OeciPat Bost, Soc.) Nye ipa 122M Toms) 


Distribution.—Recent and Pleistocene. Massachussetts to 
Florida Keys. (Gulf coast: Crooked Isl., West Fla., and 
Texas. Pleistocene, Lake Borgne borings, La., and North 
Creek, West Fla. 


avara var. semiplicata Stearns, Pr. A. N. S. Phila., p. 344, 
1873, (as Columbella); Tryon, Man., 5, p. 159, pl. 55, f. 70, 
7iewos: Dall Bulls 275 Ue S.eNeME Moe nT Od 8o. 


Distribution.—Cedar Keys and Charlotte Harbor, West Fla.; 
Louisiana; Corpus Christi, Texas. 


avara var. similis Ravenel, Pr. A. N. S. Phila., p. 41, 1861, (as 
Columbella; ‘Tryon, Man., 5, p. 159, pl. 55, f. 69, ’83; Dall, 
Bull 7h1 4) TOs); Sor 


QZ MOLLUSCA OF THE GULF OF MEXICO 62 


Distribution.—Carolinas to West Florida, Galveston and 
Yucatan. 

avara var. translirataRavenel, Pr. A. N.S. Phila., p. 42, 1861; 
dinyon,Mani5, p. 159. 783; Wally Bulliaz7 pp.) 1L6,.)'so. 


Distribution.—New York to West Florida, Louisiana and 
Yucatan. 


catenata Sowerby, Pr. Zool. Soc., p. 52, 1844; Conch. Icon., 11, 
DMe2nety IO, 158") Deyouy Mani suis m7, a ipley 5S.) ss) Sis 
i824) Dall and Simpson, Bull) UW. Ss) Pish\\Com:) 1, p. 404, 
IQOl. 


Distribution.—Jamaica to Porto Rico. A shell closely re- 
sembling this species, and probably identical with it, is in 
our collection. As far as I know, this is the first time that 
catenata has been found on the Gulf coast. 


obesa C. B. Adams, Pr. Bost. Soc. N. H., 2, p. 2, 1845, (as 
Buccnum) iryon, Man, 5, p. HOON piniszh\tai7 (aud 20, (33. 
Dalle uil s7 i p.ar18, 280: Singley,.q4th Anne) Rept. dex: 
Cacompr 335.925 Evartis, Bulli Amy Pal ivoliam ip: O25, 0)5); 
Wanatta=Pr. A. N. Si Phila., 55, p: 758, 1903: 


ornmata Ravenel, cancellata Gaskoin. <A. ostretcola Melvell is a 
dark brown or black variety. 


Distribution.—Hatteras to St. Thomas, W. I. and Vera 
Cruz. Pliocene to Recent. Gulf-coast: St. Joseph's Bay, 
Crooked Isl., West Fla.; Point au Fer, Cameron and Chan- 
deleurs, La.; Galveston and Corpus Christi, Texas. Pleisto- 
cene, Grand Chenier, Teche No. 1 well, Jennings, La., at 
1158-1199 feet, Knapp's well, Terrebonne Parish, No. 1 at 
1050-1800 feet, No. 3 at 1150-1440 feet, Galveston well at 
440-446 feet. Pliocene, New Orleans Gymnasium Club well 
at 1200 feet. 


amphisella Dall, Bull, M. C. Z., 9, p. 91, 1881; 18, p. 188, pl. 
1OM Me LOG SOs Bull 2 72 0 S..N weep aero ple  TOnpia LOG, 
89. 


Ore BULLETIN 38 03 


Distribution.—Yucatan Strait 43—640 fms. 


hotessieriana d’Orbigny, Moll. Cuba, 2, p. 138, pl. 21, f, 37-39; 
‘Tryon, Man), 5) p.lrgasnpl sods 2unneses Dall «pitti aE 
POLLS. OO: 


Distribution.—Tampa, West Fla., to Guadeloupe, W. I. 
Very closely allied to the following species. 


pretri Duclos, Chenu, Conch: Ill., pl, 16, f. 7, 8; Tryon, Man., 
Rep A MAAN DL (52a ONTOS 
Note.—C. B. Adams’ eldella [Conch. Contr., p. 63, 1850, 
(as Pleurotoma); Dall Bull. U.S. N. M., p. 118, 1889] an 
unfigured species, is 7#de Dall and Simpson (1901) merely a 
mutation of prefrz, as shown by cotypes from Adams. 


Distribution.—Variety albella, Cape Fear, N. C., to West 
Florida and Jamaica. 


Genus COLUMBELLA Lamarck 


mercatoria Linnzeus, Syst. Nat. ed. X, p. 730, 1758, (as Voluta); 
Kiener, Coq. Viv., Columbella, p. 23, pl. 5, f. 1, 1841; Dall, 
Bull. 37. p. 116,89; Dall and Simpson, Bull: U.S. Fish 
Comat, p4'403) 1901. 
Distribution.—Cape Lookout, N. C., to Barbados and West 
Florida. Pleistocene of Costa Rica. 

rusticoides Heilprin, Tr. Wagner Inst. Sci., 1, p. 81, pl. 8, f. 9, 
1887. Pr VAC ONS. Philaliip is 2mploia. f..20)' 87.) Dally ain 
WET SM ia) ip. 125; oouNauohan i eubl. Nose. tC@artsae 
nSty pl 7 Ti TOO; 
Distribution.—Cedar Keys and St. Marks, West Florida to 
Cuba. Pleistocene, Labelle, West Fla. Pliocene, Caloosa- 
hatchie beds, West Fla. 

Genus ASTYRIS H. and A. Adams 


lunata Say, Jour. A. N.S. Phila., 5. p. 213, 1826, (as Massa ) 
Tuomey aud Holmes, Pl. Fos. S. C. p. 136, pl. 28, f. 6, 


O4 MOLLUSCA OF THE GULF OF MEXICO 64 


1857, (as Buccinum); Hilgard, House of Rep. Ex. Doc., 1, 
PEN) Ps S8ou1s73> Walle Bull27) 5 Ue SAIN MES (ps 118, pl: 
SOM TOME SOs OMnMsom. Occ: Pai BostanSocun Np bln) 7.0. 
133, I915. 

Bnecinum zonalis Linsley, Am. Jour. Sci., (1); 48, p. 285, 
1845. 
Distribution.—Canada to Florida Keys. 0-12 fms. Gulf 
coast: Cedar Keys, Fla.; Chandeleurs, La.; Galveston and 
Corpus Christi, Tex. Pleistocene. Lake Borgne borings, 
La., Knapp’s wells, Terrebonne Parish, No. 2 at 1050-1790, 
No. 3 at 258-1375 feet. Also of North Creek, Manatee and 
Labelle, West Fla. Pliocene of the Caloosahatchie, Fla. 


duclosiana d’Orbigny, Moll. Cuba, 2 p. 136, pl. 21, f. 31-33, °45. 
(Not of Sowerby, 1847, which is a Javan shell); Dall, Bull 
37, U.S.N.M.,p. 118, ’89, (as dunata var.); Dall and Simp- 
son, Bull. U. S. Fish Com., I, p. 405, 1901. 
Distribution.—Hatteras to Barbados and Tampa, West Fla. 
0-63 fms. 

diaphana Verrill, Tr. Conn. Acad., 5, p. 513, pl. 58, f. 2, 1882; 
Dalla ME CHZ ITs p. TOL plenaisinewloun So; “bth 27° 
Oe SHUN MES Da mrs ple 25. fi. Op SOE 
Distribution.—Rhode Isl. to Gulf of Mexico between Mis- 
sissippi delta and Cedar Keys, 64-487 fms. 


Genus NiTIDELLA Swainson 


parvula Dunker, Zeitschr, Mal., p, 64, 1847, (as Buccinum); 
Mnyvony Man Si. p.022.\ pl. 48a te Sa Wall Dialling, 
We Sa NG Mes pe 183) 30: 


Distribution.—Gulf of Mexico, Texas region, to Barbados. 
Genus AESOPUS Gould 


stearnsi Tryon, Man. Conch., 5. p. 179, 1883, (as Semznella) ; 
Dah pBulliMeyC.\ Zi 18) pi) 194. ply Zou tisn SOs mba 3 7, 
PALLIS Hi le Lous 52 SO: 


65 BULLETIN 38 


a 
ea] 


jilosa Stearns, Pr. AUN S) Bhilai pais asics mig ime 7a4 (as 
Nitidella). Not Aesopus tilosus Angas, 1867, cf. Columbella 
peculiarvts Guppy, Geol. Mag., London, pl. 18, f. 20, 1874. 


Distribution.—Type locality, Tampa Bay, Fla. Also off 
Cape Fear, N. C. 15-17 fms. Pliocene of the Caloosahat- 
chie, Fla. 


Genus MUREX Linnzeus 


beaui Fischer and Bernardi, Jour. de Conch., 5, p. 295, pl. 8, f. 
TiTos5os Dall Bulls Mo CZ a mee wemnO sien Soi a llama, 
We SING pS) 78O: 


Distribution.—Gulf of Mexico between Mississippi delta and 
Cedar Keys, at 111 fms. Also Florida Reefs and Guade- 
loupe. 82-183 fms. 


cabriti Bernardi, Jour. de Conch., 7, p. 301, pl. 10, f. 3, 1858; 
Dall Biull 13700. 9S. NE UMES iospene m7 So. 


Distribution.—Hatteras to Barbados, Gulf of Mexico, Blake 
station 36, at 84 fms. 


messorius Sowerby, Pr. Zool. Soc., p. 137, 1840; Conch. Illust., 
Murex, £. 93, 1839; Reeve, Conch. Icon., Murex, f. 90, 45; 
Dalit W. Li S., 3; pwineo. 160. Dalland) Simpson) Bull 
UMSirish Com, 1, \p:) 407) 19e1 


Distribution.—Gulf coast, Cedar Keys, Fla. and Texas, to 
Colon. 2-30 fms. Pleistocene, New Orleans pumping 
station No. 7. Pliocene of the Caloosahatchie River, Fla. 


(Chicoreus) rufus Lamarck, Anim. s. Vert., ed. I, 7, p. 162,1882; 
Apyonuk) Man) 2, yp) oo) uplimmsnatent4s) 1.30.) (CAttenuNeceyculsy 
Avo) Dally Bull 27, Up SaNe Vern Si SO: 


sexcostata Emmons, Geol. N. Car., p. 248, f. 106, 1858. 


Distribution.—Cape Fear, N. C., to Cartagena, Colombia, 
and Antilles. Also West Fla. 5-30 fms. Pliocene Ca- 
loosahatchie beds, Fla. 


96 MoLLUSCA OF THE GULF OF MEXICO 66 


(Chicoreus) brevifrons Lamarck, Anim. s. Vert., ed. I, 7, p. 161, 
1822; Reeve, Conch. Icon., 3, pl. 3, f.13, 1845: Tryon, Man., 
PPOs tb 72) Toso; Dalle sBull 27. Ue Sa NEMO. Os. 01S, 
"89; Tr. W. I.S., 3, p, 140,’90; Dall and Simpson, Bull. U. 
Sadish Com: 15.p..407, LOL: 


Distribution.—South Carolina to Cartagena, Colombia. Gulf 
coast, West Fla. Pliocene, Caloosahatchie beds. 

(Phyllonotus) pomum Gmelin, Syst. Nat., p. 3527, 1792; Reeve, 
Gonchs eon: 3) pl. 9, f.-35; 1845: sbryons) Many-2.. p. 97, 
piezo. f. 132, 80; Dall) Bull. 27, U2 Ss NaMe, pp 120, pl 16, 
fee? e280" br. Wi. I. S., 3, p: 42), 007 Dalljand) Simpson; 
Bull. U. S. Fish Com., I, p. 408, I90t. 

oculatus Reeve, 1845; asperrimus d’Orbigny, 1853; mextcanus 

Petit, 1852; zmperialts Swainson; globosa Kmmons, 1858. 


Distribution.—North Carolina to Venezuela. Gulf coast at 
various localities, West Fla.; Cameron and Chandeleurs, 
La., and Texas. Pleistocene, New Orleans pumping station 
No. 7. Pliocene, Caloosahatchie River, Fla. Lower Mio- 
cene, Antilles. 

(Phyllonotus) fulvescens Sowerby, Conch. Illust., f. 30; Kobelt, 
Nalin Mal. (Gesell), p.155,. 77; Dalit Bullaa7. ULo5 Ne ME, 
perzo7s8o7) Vanatta, Pr: A. NaS Phila 55 apa 75011903: 


Distribution.—Hatteras and the Gulf coast at Indian Pass, 
West Fla.; Point au Fer, Chandeleurs, La.; Galveston, 
Corpus Christi and Quintana, Texas. Pleistocene, Grand 
Chenier, La., Knapp’s wells, Terrebonne Parish, No. 1 at 
2000-2150 feet, No. 3 at 790-830 feet. 
Genus FAVARTIA Jousseaume 

cellulosa Conrad, Pr. A. N.S. Phila., 3, p, 25, 1846, (as AZurex); 
DalleBull Me Ce: Zo18, p. 210, pli monteuns Go (as Oczzclra):. 
Bully.3 eae SNe Ms ps 120, pla Tomine SO ambi WEL noe, 
Beant 50s 90: 

nuceus Moerch, Cat. Kierulf,p. 14, pl. 1,f. 9, ’50; Cat. Yoldi, p. 
95, 1852, (as Ocinebra). 


67 BULLETIN 38 O7 


Distribution.—Cape Lookout, N. C., to the Antilles and 
Wera Cruz. Gulf coast, Tampa Bay (type locality), Cedar 
Keys and Texas. Pliocene of the Caloosahatchie. 


cellulosa var. levicula Dall, Bull. M. C. Z., 18, p, 211, 1889; Bull. 
27, Py 120% (89. (as Ocweora): 


Distribution.—Cape Lookout, N.C., to West Florida and 
Yucatan. 22-37 fms. 


intermedia C. B. Adams, Contr. Conch., p. 60, 1850, (as Murex); 
Dall Bull) 27, U.S) NOM. pait2onSoii(as) Oczeb7a) Dally 
and Simpsou, Bull. U. S. Fish Com., 1, p, 408, 1901. 


Distribution.—Key West to St Thomas and Texas region of 
the Gulf. Variety a/ta Dall, Caloosahatchie Pliocene. 


Genus UROSALPINX Stimpson 


cinereus Say, Jour. A. N.S. Phila., 2, p. 236, 1822, (as Fusus); 
Amer./(Conch., 3, pl. 209, 1831; Gould, Inv. Mass:,p.'370)) £: 
637, (as Buccinum); Tryon, Man., 2, p."152, pl. 39, f. 3,80; 
Dall Pr iOS SiN MeO np eiseOumnsse btu 27500 an suiNE 
M., p. 120, pl. 50, f. 6, ’89; Johnson, Occ. Pa. Bost. Soc. N. 
AYipyire2) Tor5:) Dall. Bulleri2 Wen SsoNii Mi. p.TOo MN TO2. 


Distribution.—Nova Scotia to Florida. Gulf coast, Cedar 
Keys (Dall); Pleistocene, Lake Borgne borings, La., (Hil- 
gard). Also on the Californian coast (introduced with 
oysters). 

perrugatus Conrad, Amer. Jour. Sci., p. 397, 1846, (as Fusus); 
DallBall Mi Ce OZ isi py 2mwimson Bull) 27 pei 2ONWso. 
Vaughan\2d' Ann. Rept. /HlanG.iS.))p. 148. 909. 


Distribution.—West Florida from Cedar Keys to Key West. 
Type locality, Manatee River, West Fla. Pleistocene, 
North Creek and Orient, West Fla. 


tampaensis Conrad, Pr. A.N.S. Phila., 3, p. 25, 1846, (asMurex); 
Dall Bull. Me iC. Z. 18) pw 214 8on sBull37,) Ui) Si Noe, 
DIE 2 SO; 


g8 MOLLUSCA OF THE GuLF OF MEXICO 68 


Distribution.—West Florida from Cedar Keys to Sarasota. 
Pleistocene, North Creek, West Fla. 


Genus MURICIDEA Swainson 


multangula Philippi, Zeitschr. fur Malak, 5, p. 25, 1849, (as 
uss) eX pid: ind: Beschr) (33) pel Lay plan 244i.) O18 50% 
DAB aN a7 UL SON Mi pent 20) Olan 7ONton tense Oats | Ve 
SMa E49) T890%)) Pry Ua Oo INN 2AR DE SO5 i Bl) 20) 
ite) 1G) CKO. 
hegeprele Dall. Pro Us SiN .\M.) 6; pelza7i 1833) 
Distribution.—Cape Fear, N. C., to Yucatan and the North- 


ern Antilles, Gulf coast, Cedar Keys and Texas. o-g5 fms. 
Pliocene, Caloosahatchie beds. 


ostrearum Conrad, Pr. A. N.S. Phila., 3, p. 25, 1846, (asAZurex); 
Dalley Oi SNe ME. 240.) 505.0 pla goin Z LOO. 


Muyjomdana Conrad, Am, jour. Conch.) )5)) pi 206, (pl. 12, £2145 
T8oo9u (as) Crosalpznx):\Dall, Pr. Us S MNAME ow p. 326) 183. 
lle CZ TS oni 20 Bon ra Wilh Sait DW AON)) GO} 
Vaughan, 2d Ann. Rept. Fla. G. S., p. 148, 1909. 
Distribution.—East Florida south of St. Augustine and Gulf 
of Mexico, Cedar Keys, Fla., to Yucatan. 0-13 fms. Rocky 
shores. Pleistocene, Manatee and Orient, West Fla. Plio- 
cene, Caloosahatchie beds, Fla. Resembles Uvosalpinx 
perrugatus Conrad, but is slenderer and has a very different 
operculum. 

hexagona Lamarck, Anim. s. Vert., 7, p. 169, 1822, (as Murex); 
Reever Conch Micont)(2.)\pl4) 27 tuiTzonmaqs Mable iil 27. 
We So Ne OM. p20, 789; Dall & Simpsonm/Bully U0. SiFish 
Com., I, p. 409, I9OI. 

Distribution.—Gulf of Mexico, Texas region to St. Thomas, 
W.1I1.25 fms. Genotype. 

philippiana Dall, Bull. M. C. Z., 18, p. 213, 1889; Bull. 37, U. S. 
ING OM ape 20% SOs) ert. Ole.) UNE) Meh 2ani mn S@unnmle Doi fic) 
1902. 


69 BULLETIN 38 99 


Distribution.—Off Cape Catoche, Yucatan, and also Key 
West, Fla. 20-25 fms. Coral sand. 


Genus EUPLEURA H. and A. Adams 


caudata Say, Jour. A. N.S. Phila., 2, p. 236, 1822, (as Ranella); 
Stimpson, Am: Jour!) Conch. ai.) 158.) ple 8s te Sun oO: 
iilsard, Mouse of Rep. Ex. Docr 1, pt-2) sp..886. 47.33) Dalle 
BU VS) AN Mi Oli pass 2On Hn San sbi tac mila SemiNen MeN moe 
TZOM Pl.) 50 te LT, SOx} Ounson) Oces Par aBost Soc iN Eley 
Fey, (Oui KO EO) Hse 
Distribution.—Massachussetts to Florida. 1-3 fms. Gulf 
coast: Cedar Keys, St. Mark’s, Charlotte Harbor, Fla.; 
Chandeleurs, La. Pleistocene, Lake Borgne borings and 
New Orleans well of 1856 (Hilgard), and of Labelle, West 
Fla. 


caudata var. sulcidentata Dall, Tr. W. I. S., 3, p. 144, 1890. 


Ranella clathrata ‘‘Gray’’ Calkins, Pr. Dav. Acad. Sci., p. 
234) 1878. 

muriciyormis Viyon, | Man.) 2.) palnss.. (nm part) Noor 
Broderip. 


Distribution.—Gulf coast of Florida, Pleistocene of South- 
west Fla. 


Genus TROPHON Montfort 


(Boreotrophon) lacunellus Dall, Bull. M. C. Z., 18, p. 205, pl. 15, 
iq rsso; }bull).37.) Ue Sa Ne Meeps c 20. (poll 1 Satis Or 


Distribution.—Cape Fear, N. C., to Dominica. Also West 
Florida. 227-769 fms. 


Genus THAIS Bolten 


patula Linnzeus, Syst. Nat., ed. X, p. 739, 1758, (as Buccinum) ; 
Reeve, Conch. Icon., 3, pl. 1, f. 3, 1846; Tryon, Man., 2, p. 
150, Pls) £4) 19-22) 807m all nB alle 7 Amis. Ne vines 


OO 


MOLLUSCA OF THE GULF OF MEXICO 70 


M22 SOM (AS A UUpuUnd) wall Pr Wa Sum Ne vie 3751s 221, 
IQIO. 

Distribution.—East and West Florida to Texas, the An- 
tilles and Brazil. Also Pacific, Gulf of California to Pana- 


ma and Peru. 


haemastoma Linnzus Syst. Nat., ed. XII, p. 1202; Reeve, 


Conehyicon., 3, f. 21, 1846; Tryon, Man.\)2) :p.)167,) pl. 40, 
Paso, 83 1880; Dall, /Bull.)327 0 U2ssy Ne Mes p22 .\pliaa: 
is. 4, pl. 46, Ta-2b, 89°) Singley ‘athyAnn. Rept, dex.) G: 
Se pessow oor WVanatta,) Pir) AVENE Sei Phila sic pag 5o, 
1903. (All as Purpura). 


Distribution.—Cape Hatteras to Trinidad Isl. Gulf coast 
Cedar Keys and Ft. Barranca, Fla.; Horn Isl., Miss.; Point 
au Fer, La.; Galveston, Corpus Christi, Tex. Pleistocene, 
Grand Chenier, La., New Orleans pumping station No. 7, 
Knapp’s No. 3 well, Terrebonne Parish, at 1400-1440 feet. 


haemastoma var. floridana Conrad, Jour. A. N.S. Phila., 7, p. 


Z2OceapnZOwta 2h. Iker ry On, Miadei2 epsnno7a splerd@u ft: 
85, pl. 50, f. 86; Dall and Simpson, Bull. U. S. Fish Com., 
InP: Gul, Loon.) CAllias Purpura): 

Distribution.— Florida to the Antilles, Gulf coast, West Fla., 
Cameron, Chandeleurs, La.; Matagorda Bay, Tex. Pleisto- 
cene of Grand Chenier, La., (very abundant and showing 
the same mutations as the living shells). 


deltoidea Lamarck, Anim. s. Vert., 7, p. 247, 1822; Reeve,Conch. 


liconsusn plan fis, 1S4o> Muster mC onchaCabet pla 2.) fh 
De Mirror, MEL Boy mda gollan Mya is Us. lacctsroe! IDEN Is i000 |. 
BE OM ol UNE) Men pe 122) So. Dally andtSimpsony Bully U): 
SwhishiComyyis py 411, Toor. | CAlWasw aarp na. 


Distribution.—Florida to St. Vincent, W. I., Gulf coast, 
West Fla. and Tex. 


Has BULLETIN 38 LOL 


Genus SISTRUM Montfort 


nodulosum C. B. Adams, Pr. Bost. Soc. N. H., p. 2, 1845, (as 
Purpura) rvons Nani (2) pi TOOn Dien 5 One n27 Saelooor 
Dall and Simpson, Bull. U. S. Fish Com., 1, p. 411, Igor. 
Distribution.—Cape Romano, southwest Florida, to Colon. 
Also Antillean. 

roseum Reeve, Conch. Icon., sp. 46, 1846; Dall, Bull. 37, U. 5. 
IN VMs p.)/ 122.) 1830: 
Distribution.—Gulf of Mexico, Texas region, to Barbados. 
The rosy color is thought to be caused by an encrusting hy- 
drozoon. 


Genus CORALLIOPH!{LA H. and A. Adams 


abbreviata Lamarck, Enc. Me’th., pl. 435, f. 2a-b, 1795, (as 
Haynie) AX mans.) Vientuni emule on ene 22) 
galea Chemnitz. 
Distribution.—Cape Fear, N. C., to the Antilles. Gulf 
coast, West Florida. Pliocene, Caloosahatchie beds. 


lactuca\; Dall, | Bull. M. iC. Z.\18))p1)220, pl! 16,/f. 6, 1880: 


Distribution.—Gulf of Mexico, Blake station 5, in 152-229 
fms. Also off Fernandina, Fla. 


deburghiae Reeve, Pr. Zool. Soc., p. 208, pl. 38, f. 3, 1857, (as 
Riizochilus)\:\ Mnyon, Mane 2iinla Gon) fs) 283.) TS soOuncas 
Watzax7s)i2)\ Dall, Bull WA SNe Men p22. pla TONE Li siiso: 
Distribution.—Cape Hatteras to Barbados. Also West Fla. 
56-878 fms. 

Genus EPITOMIUM Bolten 

angulatum Say, Amer. Conch., No. 3, pl. 27, (two upper figs) 
1831, (as Scalaria clathrus var.); Hilgard, House of Rep. 
Bx DOC. WT)" Pts) 2) (Passive cso ne oO allen Dall 27. Oks kINe 
M:)) (py) 122,89; Singleya Ath Pano iRept Mex CaiSii pe 
236,792; Johnson) \\Occ mea Bosts SOC4 UN). al...) 7.npIntOs" 


IQI5. 


LOZ MOLLUSCA OF THE GULF OF MEXICO 72 


Distribution. —Connecticut to West Fla., Point au Fer., La., 
and Galveston, Tex. Pleistocene, Lake Borgne, La., bor- 
ings. 

sayanum Dall, Bull. M.C. Z., 18, p. 313, 1889, (as Scala); Bull. 
SPO SUN VEN Dana pl.) SO,) Mutou Sonmebt WN lS. 3, 
Piss) 90; Singley, 4th) Ann.) Repty Tex. G.'S.))p. 336, 
1892. 

cdathrum Say (in part). 


Distribution.— Virginia to Florida Keys, West Florida, Cam- 
eron, La., Corpus Christi and Galveston, Texas. Pleisto- 
cene, Grand Chenier, La. Pliocene, Caloosahatchie River, 
Florida. 


tenue Sowerby, Pr. Zool. Soc., p. 110, 1844, (as Scala); Dall, 
Bulle a 7e Wi SiN iM. pi 522) t8so: 


Distribution.—Gulf of Mexico, Texas and Cameron (?) La., 
tole sLnomas, WT. 


nitidellum Dall, Bull. M. C. Z., 18, p. 314, 1889, (as Scala); Bull. 
TORRONE: Wen pink). 8: Er O en oamiNn Mest 245) 6 SO, 
DI BOK ta.8.) 1902: 
Distribution.—Hatteras to Cedar Keys, Fla. 32-63 fms. 


Seinen Wall Bull..May/C./Z.) 18) p. 210 n88q,n bull 375) Uso. N 
NRT PY 12407 BOs) (aS Sala) 5) (REAM E ROU MING IME N24: Ds N5O6) 
PLE ZOR TOM TOO2: 


Distribution.—Hatteras, N. C., to Texas and Vera Cruz, 
Mexico. Io-16 fms. Pink with white varices. 


denticulatum Sowerby, Thes. Conch., p. 87, pl. 32, f. 25, 26, °47; 
miyon Mani. Os pls) 13. f. 100, (e875) DallyyBuhlhy i237 (Us) Ss 
INE py n2qn Soe 
Distribution.—Hatteras to West Florida and the Bahamas. 


lineatum Say, Jour. A. N. S, Phila., 2, p. 242, 1822, (as Sca- 
laria); Hilgard, House of Rep. Ex. Doc. 1, pt. 2, p. 887, °78; 
Mnvonw Mani Ope 79. ply 16, cf kmorisnu Oa lair Nis be se. 


73 BULLETIN 38 103 


20 pe 5S, QO; JohnsonaiOce wha OSes SOC. IN auelna aap: 
HOPE Akay els 

Distribution.—Massachussetts to Charlotte Harbor and 
Cedar Keys, West Fla.; Cameron, Point au Fer, La.; Gal- 
veston and Corpus Christi, Tex. Pleistocene. Grand 
Chenier, La., and Knapp’s No. 2 well Terrebonne Parish, 


at 1550-1570, 1731-1739 feet. Pliocene Caloosahatchie 
beds, Fla. 


turriculum Sowerby, Thes. Conch., 1, p. 92, pl. 33, f. 61, pl. 34, 
f. 88, 1847; Singley, 4th Ann. Rept. Tex. G. 5S., p. 336, 
1892. 


Distribution.—Cape Hatteras to Haiti. 16-32 fms. Gulf 
coast at Galveston (Singley). Pliocene of the Caloosahatchie 
River, Fla. 

mitchelli Dall, Nautilus, 9, p. 112, Feb. 1896; Pr. U.S. N. M., 
24, p. 506, pl. 30, f. 3, 4, 1902. 
Distribution.—Matagorda Island, Texas. Collected by J. D. 
Mitchell. 

cochlea Sowerby, Thes. Conch., Scalaria, p. 103, pl. 35, f. 142 
(only) 1847;Moerch, Jour. A. N. S. Phila., 8, p. 205, 1876; 


Mayon Man), (9. 81, :pln7 ink 2400 8o7 Walls PEW. Saunt 
MONZA D500. ply; 30, taj, oo2r 


Distribution.—Hatteras, N. C., to Tortola. W.I. Gulf of 
Mexico near Cedar Keys, Fla., at 25 fms. Sowerby’s refer- 
ence to its occurrence at Loando, Portuguese West Africa, 
is questionable. 


galvestonense Harris, Bull. Am. Pal., vol. 1, p. TO3) Wplena iter e7pe 
1895. 


Distribution.—Upper Miocene, Galveston well at 2552-2871 
feet. 


Note.—The following species of Epztomium have been re- 
ported by Singley from the Texas coast:—sericifila Dall, 


104 


MOLLUSCA OF THE GULF OF MEXICO 74 


(Bull. M. C. Z., 18, p. 313, 1889); multistviatum Say (Jour. 
ONES ve hilay nse pe 20S" 1520 DallveBulliisr We SauN. Mie. 
pl. 50, f. 5, 1889) ;contorquatum Dall, (Bull. M. C. Z., 18, p. 
318, pl. 18, f. 9, 1889); and modestum C. B. Adams (Jamaica 
Shells). These species are not cited, as far as I am aware, 
by other authors from Gulf coast. They are not in our col- 
lections from the Gulf and are noted here solely fide Singley, 
(Fourth Ann. Rept. Tex. Geol. Surv., p. 336, 1892). 


Genus JANTHINA Bolten 


janthina Linneeus, Syst. Nat., ed. X., p. 772, 1758, (Helix); 


d.Oxrbigny, Voy. Am: Me’r); p.1413,\ pl: 61, f) 8-10), 13841; 
Dal Wer Wyss Ne M327. p. 2220 neo sn) olnson. Occ. Pal 
IBOSta SOCNE Een 7.) p. TOT Tor 5: 


fragilis Lamarck, An. s. Vert., p. 89, 1801; Tryon, Man., 9, 


p. 36, pl. 9, f. 4, 1887; Singley, 4th Ann. Rept. G. S. Tex- 
ASD son 1892. bartsch, Bull. orgwUaySe Ney Mie. ‘py (65; 
IQI5. 

Distribution.—Nantucket, Mass., to Colon; and West Fla. 
to Quintana and Galveston, Texas. Pelagic. Also Pacific 
Ocean, Lat.42°N.to 36° S.,and Indian Ocean, Port Alfred, 8. 
E. Africa. 


pallida Harvey, Thompson’s Ann. Phil. and Nat. Hist., 5, p. 96, 


Dieta Sat nbyon.. Man. Opec ep lmTOntN tS) .T S870 


Distribution.—Europe, the West Indiesand Gulf of Mexico, 
Cameron, La. (washed ashore). Also tropical waters of the 
Pacific Ocean. Pelagic, 


globosa Swainson, Zool. Illust. (1), 11, pl. 85, 1822; Tryon, 


Mano) ple ton tb2,)87-) singley Athy Ann Rept mMexa) p. 
336, ’92; Mitchell, List Texas Shells; Bartsch, Bull. 91, 
Se Ne Me pos sro T's: 


Distribution.— Gulf Stream, Pelagic. Washed ashore at 
Quintana, Tex. and West Fla.Also Port Alfred, S.E. Africa. 


communis Lamarck, An. s. Vert., ed, Desh., IX, p. 4; Tryon, 


BULLETIN 38 Io5 


= 
Cr 


Manitou plio, ti ooumesre Dall Butiia 7.) MUL S.. iNet VME os 
126, 80; Mitchell, List. Tex. Sh.; Bartsch, Bull. 91, U.S. 
INS pL 05.) LOLS: 

Distribution.—Nantucket to Coion, West Florida, Matagorda 
Bay and Mustang Isl., Tex. Pelagic. Also Indian Ocean, 
Port Alfred, S... Africa. 


Genus MELANELLA Bowdich 


(Eulima) conoidea Kurtz and Stimpson, Pr. Bost. Soc., N. H., 4, 
Pulses all wr WSs IN IM Nowe 22ONTsS2.) Miao 
Mani Sipe 27i2ipl. (40 Nao oo moally Mira Vieni 2. hpe 
159, pl. 5, f. 11, 1890; Singley, 4th Ann. Rept. Tex., p. 336, 
192: )OonsSon, Occ.'Pa: Bost: sSocwiNetrts 7.) pol TO 15. 


Distribution.—Hatteras to the Autilles. Gulf coast: Cedar 
Keys, Fla.; Galveston and Corpus Christi, Tex. Pliocene, 
Caloosahatchie beds, Fla. 


(Eulima) gracilis C. B. Adams,Contr. Conch., p. 110, 1850; Dall, 
Balla OL SiN Mo) pitE26. 780% 


Distribution.—Hatteras to St. Thomas, W. I., West Fla. 
and Yucatan. Pliocene, of the Caloosahatchie. 


(Eulima) intermedia Cantraine, Diagn, Moll., Bull. Ac. Brux., 
Pata HL8353) dnvon, Mamiisinmz74, ple hoo, £43.80; Dally 
Bully 270 Ul) S-Ni MoS puoi 28h ia) So. i 


Distribution.—England to the Canary Isls., New England 
to West Florida and Barbados. 


(Eulima) jamaicensis C. B. Adams, Pr. Bost. Soc., N. H., 2, p. 
Gi ns453/\Dryon (Mans ipii2 73s pli Go. tS vain) i807) Dalit 
Balle Me Cy IZ i) raMipal 227i oSo! 


Distribution.—Gulf of Mexico, Blake station 20, in 220 fms. 
off West Fla. Also Barbados. 


(Eulima) subcarinata d’Orbigny, Moll. Cuba, p. 217, pl. 16, f. 4- 


6. Dryon Man.) 18) \p.) 273. 4plst6o) t447)/1886;) Dall) Bulltizin: 
ONS SNM ip IziG ear 


106 MOLLUSCA OF THE GULF OF MEXICO 76 


Distribution.— West Florida to Haiti. 

(Eulima) gibba De Folin, Les Me’le’agrines, p. 64, pl. 6, f. 4,’67; 
sbi OnwViane nS pv27 20 plow tiacwe 7 ballyn Bulla 7) 
Ws Sa INIA WEY Fo AIS Key 


Distribution.—Cape Hatteras to Campeche. 


(Liostraca) bilineata Alder, Moll. Northumberland and Durham; 
sirevonti Man .8) p.i279, (oli ZO. Wge-7a Tees Dal Ball, 
BRD E 20%) SQ. 

Distribution.—Norway to Mediterranean, and West Florida 
to Haiti. 


(Liostraca) hemphilli Dall, Pr. U. S. N. M., 6, p. 330, pl. to, f. 
Perso BU 3 7S. IN: | Mapai 2onplcaSs of. Tn. SO. 


Distribution.—Cedar Keys to Marco, West Fla. 


(Liostraca) fusus Dall, Bull.M. C. Z., 18, p. 329, pl. 19, f. 11b, 
SOR IBiCldls Aq NOheioy | ONO Cais ope caLS)y sFelliadeyy seyiieueloye Meso) 


Distribution.—Fernandina, East Fla., to St. Kitts, W. I. 
Gulf of Mexico, Yucatan Strait, 640 fms. Also off Hava- 
na, 400 fms. 


Genus NiISO Risso 


splendidula Sowerby, Pr. Zool. Soc., p. 6, 1834, (Zudimia); A. 
Indams) viness(Conch 7/250) Pp. Som NOMAzanplamt7Os mS) 2 54% 
siyon Man iS. Pal2o7) Dl. 7 Le emoOs a ball Bill i2i7) 
We SeeN yee ose 80) SO: Pr. MU EEO sN EN aie Tila N 2215 
IQIO. 


Distribution.—Cape Fear, N. C. to Grenada, and Gulf of 
Mexico, between Mississippi delta and Cedar Keys in 111 
fms. Also Pacific Ocean, Sta. Elena, Guayaquil Bay, Ecua- 
dor (Type locality), in 6-8 fms. (Cuming). 


aeglees Bush, Rept. U. S. Fish Com., p. 83, 1883-'85; Dall, Bull. 
BOM Os NG Me plas si ple var fai TOa NS On CaS iZalenrupra 
Sowerby var. ). 


aa BULLETIN 38 lo7 


Distribution.—Hatteras to Tampa, West Fla., 7-32 fms. 


Pleistocene, Knapp’s No. 3 well, Terrebonne Parish, La., 
at 1150-1200 feet. 


Genus PYRAMIDELLA Lamarck 


delabrata Linnzeus, Gmelin, Syst. Nat. p. 3585, No. 113, (as 
Trochus): (Lryon, ) Maney8\ ps (200 pl agen i772 alll 
Bull 276 WSs IN Me oan 2. 8 eS or 


Distribution.—Sarasota, West Fla. to Barbados. 


(Longchaeus) crenulata Holmes, Post-Pl. Fos. S. C., p. 88, pl. 
13, tw Alaa, 850!) (asiOve/esers) ew allie vill NI) Ce Zanersy 
10 AL eSKstOe Ue AWE SL io. arin ome Wewoigeyy exe, Ja. 
Ne SA ehila sp. 75S. loose 
Obeliscus tesellatus Dall, Pr. U.S. N. M., 6, p. 330, 1883. Not 
of Adams, 1855. For full synonymy see Dall, 1889. 


Distribution.—South Carolina to St. Thomas, W. I. Gulf 
coast, Cedar Keys, Tampa Bay. Fla., and Galveston, Texas. 
Pleistocene, North Creek, West Fla., and of the Lake 


Borgne borings, La. Habitat, grassy or muddy flats to 2 
fms. 


(Longcaaeus) candida (Meuschen) Moerch, Mal. Blatt., 22, p. 158, 


(as Obelescus)):) Dall,” Biullaie7eOe. sae Ne OMe pil t2 Sens oor 
ie Wei Sa 3 py 247, neece 


Distribution.—Hatteras to Barbados and West Florida. 
Pleistocene of Texas. 


(Syrnola) smithi Verrill, Pr. U.S. N. M., 3, p. 380, 1881,(as Zur- 
bonella)-) hr Conn: Acadasuap isco. pl 58, yf. 1 emuss: 
Bartschs Pr.) Bost. (Soc: Ne WEleneae ps) gil. | pl. 1. Lenni: 


1909; Johnson, (Occ!) Ra Bost) soc Nive, 7. 92,0) 15 NOt 
of Tryon. 


Eulimella unifasciata Dall (pars), Bull. M. C. Z., 18, p. 338, 
PLMO, ft.) 11) 1889) Bulle enw See NLM spt 20. p leno yte 
IIc, 89. Not of Forbes (Rept. Aegean Inv., p. 188, 1843) 
which is a European shell. 


x08 MOLLUSCA OF THE GULF OF MExICco 78 


Distribution.— Massachussetts to Barbados and West Fla. 


(Syrnoia) thelma Dall, Pr. U. S.N. M., 46, p. 254, pl. 20, f. 3, 
IQi4. 
Distribution.—Upper Miocene or Pliocene. Well near Al- 
exandria, La., at 49 feet. 


Genus TURBCGNILLA Risso 


exilis C. B. Adams, Contr. to Conch., p. 74, 1850, (as Chemntt- 
Ea ee Dalal 3700 Ss NG Maia 28h Soro. Wi. Meio. 
Bee 250, 02: 
Distribution.—Hatteras to Haiti and West Florida, 3-63 fms. 
Pliocene of Caloosahatchie and Shell Creek, Fla. 


belotheca Dall, Bull. M. C. Z., 18, p. 335, pl. 26, f. 7d, ’80; Bull. 
CHAO OE NE OMee pr 28%) ol. 26) tad SOs 


Distribution.—Gulf of Mexico, Blake station 32, at 95 fms., 
west of Florida, 50fms. Also Barbados, i100 fms. 


interrupta Totten, Am. Jour. Sci., (1), 28, p. 352, f. 7, 1855, (as 
engoicla Dalles) 27): VU: Sy INGiMeen py in28, pl.) 26h 
Za Dessowm lr WWE le S43. pl 25oumo2umBanusch bia) Bost: 
SOCwN A EEN 24h ps7 pl. 12) ft ToM2etTrooo +) johnson, Occ: 
ae OSt 7 SOCe NN Ble 759 .) O15) OMG 

wmaana Mal Pr. 10S: Ni: M), 62-0 i232 1882) 

Distribution.—Nova Scotia to Barbados. 2-107 fms. Gulf 
coast: Cedar Keys, Fla.; Galveston and Corpus Christi, Tex. 
Pleistocene, Lake Borgne borings, La., Knapp’s wells, Ter- 
rebonne Parish, No. 2 at 1050-1790; No. 3 at 1150-1839 feet, 
also of North Creek, West Fila. Pliocene of the Caloosahat- 
chie beds. 

textilis Kurtz, Cat. Rec. Mar. Shells, N. and S. Car, p. 8, ’60, 
(ash@hemnzizza)\ Dall, ir. We) IeSeet pa 257" 1S O2- 
Distribution.—Charleston, S. C., to Marco, West Fla. 
Pliocene, Caloosahatchie beds. 


79 BULLETIN 38 109 


obeliscus C. B. Adams, Contr. Conch., p. 72, 1850, (as Chemnit- 
Ze) NOM M SHUN eva IOL a SWINE AVIS op iqeken staan Bren INI A Ile Sic 4 
BNO ZOTe Oa" 


Distribution.—Hatteras to Jamaica. 12-63 fms. Gulf coast, 
West Fla. and Galveston, Tex. Pliocene, Caloosahatchie 
beds, Fla. 


Virga; Dall; Pry Ue SiN. MAG Osipsz2Q ns Sar iBall 27. Unis sine 
Mie OuI2 5) 7 8o. 


Distribution.—Hatteras to Florida Keys and Cedar Keys, 
Fla. 2-15 fms. Pliocene, Caloosahatchie River. 


punicea Dall iPr, SiN aM IG ips agioms8arlin.) Wi lene: 
Bean Zon ESO2h 


Distribution.—North Carolina to the Bahamas. 2-31 fms. 
Gulf coast, Cedar Keys, West Fla. Pliocene, Caloosa- 
hatchie beds. 


curta Dall, Bull, M. C. Z., 18, p, 337, pl. 26, f. 7c, 1889; Bull. 
WET SONG Mp. 28. pl 26h ies, 


Distribution.—Yucatan Strait, 640 fms. Also off Cape Hat- 
teras and Antilles. 


speira Ravenel, Pr. Elliott Soc. Nat. Hist, 1, p. 280, 1859, (as 
Chemnitzia) Tolmes:) Post kiiHossnou ©. p./82, pliant 
LAWLOsSor Mal k ar. WW ise wDe25S.) TOO2. 


Distribution.—Pleistocene, S. Carolina, and “de Hilgard, of 
the Lake Borgne, borings, La. Pliocene, Coloosahatchie 
beds (?). Not reported in the recent fauna. 


reticulata C. B. Adams, Contr. Conch., p. 75, 1850; Dall, Tr. 
WE Seip: 2OOn 1O2. 


Distribution.—North Carolina to Jamaica, o-60 fms. Re- 
ported by Singley from Galveston, Tex. Pliocene, Caloosa- 
hatchie beds, West Florida. 


conradi Bush, Vanatta, Pr. A. N. S. Phila., 55, p. 758, 1903. 


{IO MOLLUSCA OF THE GULF OF MEXICO 80 


Distribution.—Cited by Vanatta from Calhoun Co., North- 
west Florida. 


(Parthenia) cedrosa Dall, Pr. U.S. N. M., 6, 1883; Bull. 37, 
WES NGM Dai ZO pl Asi fa) 4m Sao. 


Distribution.—Cedar Keys, West Fla. 


(Careliopsis?) styliformis Moerch, Malak,Blatt., 22, p. 169, 1874, 
DAS we MN COZ.) 18. pi i338.) TSsoq Bulle aan ss MIN: 
Wile. Oe IeeXON \eloy. 


Distribution.—Hatteras to Sarasota Bay, West Fla. Also 
St. Thomas, W.I. 2-52 fms. 


Genus ODOSTOM!IA Fleming 
acutidens Dall, Pr. U.S. Nat. Mus., 9, p. 331, 1883. 


Distribution.—Hatteras to Cedar Keys, West Florida, 2-107 
fms. 


Note.—In 1892 (Tr. W. I. S., 3, p. 250), Dall regarded this 
shell as.a variety of the European Miocene to Recent species, 
O. conoidea Brocchi (Conch. Foss. Subapp.,2, p. 659,pl. 16,f. 
2, 1814). ‘The latter species is recorded by Harris from the 
Pleistocene of the Galveston well at 440-458 feet. 


impressa Say, Jour. A. N. S. Phila., 2,p. 244, 1822; Dall, Pr. 
WeeSUNGEMe.) 6. pH 2am! T882) Bulbas MWe Sa Ni iMEn D130) 
PeNS2 it bso Sineley, 4th pAnnivRepty lex.) Gi Si)p. 
BOT RO2 i banrtsch ahr Bost. SOCAN nen azn Ml 20it oS 
LOO: oOnnsSonh Occ! Pay Bost. SocwNe Mee Te Dp osn LOLs: 


Distribution.—Massachussetts to Florida. Gulf coast: 
Tampa and Cedar Keys, Fla., Corpus Christi, Tex. Plio- 
cene, Caloosahatchie beds, Fla. 


seminuda C. B. Adams, Jour. Bost. Soc. N. H., 2, p. 280, pl. 4, 
Tana S Zo (asi /azzenza) > DallsBuallai aa Wo SsuNi) Mes p 
MZOM pS 25) £410,994) Bartsch, Pry Bost Soe. Ni ME 24.\p 


T} 
Leon} 


BULLETIN 38 Tit 


97, pl. 13, f. 45, 48, 1909; Johnson, Occ. Pa. Bost. Soc. N. 
H., 7, p. 97, 1915. 


Distribution.—Massachussetts to Florida Keys and West 
Florida. 


Genus PERISTICHIA Dali 


toreta Dall, Bull. M: C. Z., 18, p. 340, 1884- Bull. 27, U.. SY Ne 
Mei piso; ply 42 hOnsa) 


Distribution.—Cape Lookout, N. C., to Key West and 
Charlotte Harbor, West Fla. 2-22 fms. 


agnia) Dall) Bull. M. (C. Z.)\-18) op avowrsso- Bulli27, U.S, Ne 
MEN pst ZON Bo. 


Distribution.—Marco, West Fla. o-2 fms. Also Key West 
and off Hatteras. 


Genus ATLANTA Lesueur 


peroni Lesueur, Jour. de Physique, 85, p. 390, pl. 2, f. I, 2, 
isi7 Gd ,Onbisny,) Voy: Amey Metis pr) 171, Pl.) t2.ite reise 
nSZon Wall “Bally 27 (Us OMeNenIMe api zoOn plana a maaaian 
Ple166. it) MItoa, 1889; (RrAOM Sanwa 637.) 1. U2 2 54uno nor 
Buller 1 2, Wi S. Nev pags toon. 


Distribution.—West Atlantic, Lat. 42° N. to Tropics. Gulf 
of Mexico, West Fla. and Texas. Also eastern Pacific. 
Pelagic. 


Genus OXYGYRUS Benson 


keraudreni Lesueur, Jour. de Physique, 85, p. 391, pl. 2, 1817; 
H. and A. Adams, Genera Rec. Moll., 2, p: 92, 1858, 3, pl. 
Gout, 6. 6b: Vern) \Transy Conny yAcad.) (6, "p14 20) 18 85¢ 
Dally Bull: 27. WU. Ss NM ep raoucso. johnson.) Occysrar 
Bost Soc uN: Eas 7. pu argoxmmoms: 


Distribution.—West Atlantic, Lat. 39° N.to Tropics. Gulf 
of Mexico, West Florida and Texas. Pelagic. 


X12 MOLLUSCA OF THE GULF OF MEXICO 82 


Genus SIMNIA Risso 


(Neosimnia) intermedia Sowerby, Spec. Conch., No. 1, p. 9, f. 32, 
33, (as Ovulum), Thes. Conch., Ovulum, No. 40, p. 479, pl. 
HOO HM. O02, 1848") Reeve, Couch Neon Ovrlane pl. 7.)t. 
Bet OWensoS lity on Man. 57) 7))i py \25Iq\ oleae nee S5-S0 as 
Ovnla); Dall, Bull. M. C. Z., 18, p. 235, 1889; Bull. 37, U. 
SePNee Me ps 134), '8o. 


Distribution.—Hatteras to Brazil. Gulf coast, Vera Cruz. 
15-170 fms. 

(Neosimnia) uniplicata Sowerby, Thes. Conch., p. 478, pl. roo, f. 
30-32, 1848,(as Ovulum); Reeve, Conch. Icon., Ovulum, pl. 
OMA Loos dinvon,) Mani hon 2isah polls.) cb. ao) Ao, 
coo WalieBulleiz7 py 134, 89: Bullemra Oe SN. Meo: 
LZOm OT. 


Ovulum subrostratum Sowerby, 1848, antillarum and arcua- 
tum Reeve, 1865. carolinensts Moerch, 1877. 


Amphiperas canadiensts Moerch, 1882 (typ. err. for carolinen- 
SAO) e 


Distribution.—Hatteras to Barbados and West Florida. 12- 
121 fms. Also Pacific coast, San Pedro, Cal. to Lower Cal- 
ifornia. 


Genus CYPHOMA Bolten 


gibbosa Linnzeus, Syst. Nat., ed. X, p. 726 1758, (as Bulla); 
Incevies Conch!) eons, )15, pl.) 7.) fangeni sos. (as Ovzz/e72):- 
igor Many 7) 01 250, ple 3, t-Sl, S2se5en Dally Bulla 27, 
U. S.N. M.,p. 134, ’89; Dall and Simpson, Bull. U. S. Fish 
Com., I, p. 420,1901, (as Ulizmus). 


Distribution.—Georgia to Trinidad, 15-50 fms. Also West 
Florida. 


83 BULLETIN 38 Itz 


Genus PEDICULARIA Swainson 


decussata Gould, Pr. Bost. Soc., N. H., 5, p. 127, 1855; Dall, 


Bolle Me Cy 2008. p 237. h plo vite Oa pus Soruis tt llaiai7. 
We SaN MIS pe ra. pling. tabs! 


alozda) Dall Bully MOC) Z. on piso mos: 


Distribution.—Georgia to Barbados, and Yucatan Strait 
640 fms. 


¥ 


Genus CYPRAEA Linnzeus 


spurca Linnzeus, Syst, Nat., ed. X, p. 724, 1758; Reeve, Conch. 
liconi 2) splot,” £68. 18415 :) dnyonee Manin 9. .0O'5 1) ple mon 
BELOW T 7.) T9e5>, Dall Bulle 27 Ura aiNe MeN D136) 780) 


Distribution. —Cedar Keys, West Fla., to Barbados, 0-25 
fms. 


exanthema Linnzus Syst. Nat., ed. XII, p. 1172, 1767; Reeve, 
Conch. Icon., 3, pl. 5, f. 16,/1848;) Tryon, Man., 7, p: 164, 
pl. 1, f.4, 5, 1885; Dalland Simpson, Bull. U. S. Fish Com., 
Hea 20%) LOOK. 


Distribution. Hatteras to Colon, West Florida and Texas. 
0-10 fms. 


cinerea Gmelin, Syst. Nat., p. 3404, 1792; Reeve, Conch. Icon., 
Biiply220t) 124) 1846:) Pryous Nan 7,)) 106, pl \/2)) ters. 
ROW 85: Dall Ball a7) Ue Sune iNn pst 20) 30: 


Distribution.—Hatteras to Guadeloupe, West Florida and 
Texas. 0-163 fms. 


Genus TRIVIA Gray 
pediculus Linnzeus, Syst. Nat., ed. X, p. 724, 1758,(as Cypraca); 
Reeve, Conch. Icon., 3, pl. 23, f, 131, 1846; Tryon, Man., 7, 


Py2or pl! 20/6. (94,95, Moose Mali Ta Ve MIU Sais DE 
168, ’90. 


IY4 MOLLUSCA OF THE GULF OF MEXICO 84 


Distribution.—St Augustine, Fla., to Barbados and West 
Fla. Pliocene, Caloosahatchie beds. 

suffusa Gray, Descr. Cat. Cypr. p. 16, 1832; Tryon, Man., 7, p. 
ZOT MMU 2T se Tao TSS sy Dalle Vi Cu ZS. 24.0% 
1880. 
Distribution.—Cedar Keys, West Fla., to Barbados. 1-3 
fms., on reefs. Pliocene, Caloosahatchie beds. 


candidula Gaskoin, Pr. Zool. Soc., p. 200, 1835; Tryon, Man., 
PED 203) Die) 2218) 23) 2a) 785 DalliBall ME CZ 8, 
ZAQwIOO: Billy 372 Ul S. IN. Moy Diz Ooo. 
olorina Duclos, approximans Beck. 
Distribution.—Hatteras to Barbados. 18-140 fms. Gulf of 
Mexico, Yucatan Strait, 640 fms. 


globosa Gray, Descr. Cat. Cypr., p. 14, 1832; Tryon, Man., 7, 
OMEZOON ple ama tay O2i F920 85;)) Dall iBall MeN CZs. 0p. 
2S Or wll 27 Oe Sa ON.) Mp E20 eOr 

pilula Kiener, sphaerula Mighels. 


Distribution. —Gulf of Mexico, U. S. Fish Com. station 2373, 
west of Fla., at 23 fms. Also Key Westand Antilles. 
Pliocene, Caloosahatchie beds. 


Genus ERATO Risso 


maugeriae Gray, In Sowerby, Conch. Illustr., p. 17,pl. 7, f. 47, 
Eee2ueryon Many.) 5.) p. (9). pli /asuty aawiaaw Sac alll. ili. 
WW Ala S24 3 Da LOS LOOO: 


laevis Emmons, N. Car. Geol. Rept., p. 262, f. 139, 1858. 
Not of Donovan. 


Distribution.—Hatteras to Antilles and Colon; West Fla. 
and Vera Cruz. 0-63 fms, 


Genus BURSA Bolten 
aifinis Broderip, Pr. Zool. Soc., p. 179, 1832, (as Ranella); 


85 


BULLETIN 38 LY) 


Mnyon. (Mani 2.) pudom rola anita a Senn oS i) ally ie aillemezn 
WSUS NS ME pine 24 y Son (asi Gy renee) 
Distribution.—Hatteras to the Antilles. Varietv cubaniana 


d’Orbigny, dredged on Arrowsmith bank, Yucatan, also off 
Key West and Cuba. 


Genus DISTORTIO Bolten 


reticulatus Link, Beschr. Rostock Samml., 3, p. 122, 1807, (as 


Wistorines)); Mall Bully 27) (0 Sean vik 122.) So. Dall 
and Simpson, Bull. U, S. Fish Com., 1, p. 416, 1901; Mitch- 
elinniist mlesc! Sells. cm arm 


Distribution.—Hatteras to Barbados, West Florida and 
Mustang Island, Texas, 22-154 fms. 


reticulatus var. clathratus Lamarck, An. s. Vert., 7, p. 186,1822,, 


Inc. Me’th., pl. 413, f. 4, (as Z7ztov); Dall and Simpson, 
BullesUs Sy) ish Com! pya16, noon: 
Distribution.—Porto Rico and Gulf of Mexico near Key 
West (dredged by U.S. Fish Com.), also at Chandeleurs, 
La. Pleistocene, New Orleans pnmping station No. 7. 


Genus COLUBRAR!A Schumacher 


lanceolata Menke, Synopsis, p. 87, 1828, (as Ranella); Reeve, 


Conch: Leon)! 2,) plus Wi opens 4m n(asi 77722072) Kad Stee 
Conch Cabs; spl.65,) £08) myonuneViamet 2) pL 27h plemuomtes 
1o2, Test: Dall Bully s7.(UemowNn vic "Pp: 132.0080 Meal 
and Simpson, Bull. U.S. Fish Com., 1, p. 416, 1901. 


Distribution.—Hatteras to Barbados and Vera Cruz on the 
Gulf coast. Genotype 


testacea Moerch, Yoldi Cat., 107, (asZ77zton); Mal. Blatt., 24, p. 


25 lnyon, Man:i(2 ol) ijn eesrssr. Dall Bulle sar Oe 
SH NEEM pe 24 "SoMa isu Nile S25. 35s LOS OO- 


Distribution.—Hatteras to Sombrero, and Texas region of 
the Gulf of Mexico. Pliocene, Caloosahatchie River, West 
Florida. 


1:26 MOLLUSCA OF THE GULF OF MEXICO 86 


Genus CYMATIUM Bolten 


olearium Linnzeus, Syst. Nat., ed. XII, p. 1216, (77zton); Reeve, 
Conchilicon aiuig2 yt sa4cMrvon Vlaneii aN Osim. DIAS) h. 
27a Sie Dal oBalleya7 OS. IN Mey pale 2h SO (asia ane- 
QuUsia) ww pareschs, Bully Ue Ss: Ne Me, py O4 i 1oms. 


Distribution.—Hatteras to Cartagena, Colombia, and the 
Texas region of the Gulf of Mexico. Also Indian Ocean, 
Port Alfred, South Africa. 


chlorostomum Lamarck, An. sans Vert., 7, p. 185, 1822, (as 777- 
tomeKeiener,. icon, Coq. -Viv., p;) 19), pl.) 12) 4.123, Pryon, 
Mammen ie pla Teka 48 Heer Dalle Wales 7 WES. 
INGEN ps132) 895) Dall’ andi Simpson Bully \Us (S) Bish 
Com., 1, p. 417, 1901,) as Lampusza). 


Triton pulchellus C. B. Adams, Contr. Conch., p. 60, 1850. 


Distribution.—East Florida to Barbados and Gulf of Mexico 
along the Mexican coast. 


graci'e Reeve, Conch. Icon., 77cton f. 58, 1845; Tryon, Man., 3, 
DiEuee-o7 4 1SSi Dall. Bull. MCHA ae Teihpy227, plei2oy te 
PSs ules UL S\N). Mi. Tae Mole aoivater 26nS ou (as 
Lampusia). 

Distribution.—Cape Catoche, Yucatan, 24 fms., U. 5. Fish 
Com. Also Colon and Barbados. —Max. depth Ioo fms. 


cynocephalum Lamarck, An. sans Vert., 7, p. 184, 1822, (as 777- 
ton); Reeve, Conch, Icon., 2, pl. 8, f. 26, 1844; Tryon, Man., 
Be {Os WO, ile iiny 0 On ole uscpe igs. oes ION ie rodblkM eye Ole 
SPNCOMei pr ii22)?89-) Dall and) Simpson, Bull. o.bishi 
Com), 1, p. 417, 1901, (as Lampusza): 
Distribution.—Florida Straits to Margarita Isl. Texas re- 
gion of the Gulf. 


Genus NYCTILOGCHUS Gistel 
femorale Linnzeus, Syst. Nat., ed. X, p. 749, 1758, (as Murex); 


87 BULLETIN 38 1l7 


Reeve, (Conch. i lcom ian olay ite 22 tensa ae (asmminetos)): 
Aryon,; Many iasip. Towiplnon Zon nse Dalla Billa iw 
S. N. M., p. 132, ’89; Dall and Simpson, Bull. U.S. Fish 
Comin i) p47", Toe), (as) Zotoz.21270))): 


Distribution.—Cedar Keys, West Fla., to Guadeloupe. 


Genus ASPELLA Moerch 


scalarioides Blainville, Faune Francaise, p. 131, pl. 5,f. 5, 6,1826, 
(as Wurex); Dall and Simpson, Bull. U. S. Fish Com., 1,p. 
410, IQIO. 


Distribution.—European and Antillean. 


scalarioides var. paupercula C. B, Adams, Contr. Conch., p. 
604 1850,) (as J7uvex) Wall Bali Zi 8. . (2ZOSsnsg: 
Billa e TU Se IN) Mee oat 2Ou aoe 


Triton cantrainet Recluz, Jour. de Conch., 4, pp, 246, 418, pl. 
8, f. 10, 1853; Kobelt, Jahrb. Mal. Gesell., 4, p, 244, 1877, 
(as Ocinebra). 


Distribution.—West Florida, at 50 fms.; Texas and the 
Antilles. 


scalarioides var. obeliscus A. Adams, Pr. Zool. Soc., p. 260, 
HeGIn iryon, Man. 2ipylmzotmple a8.f)\/467)) t88Os. Dall 
Buallkys7 US Si. UNM Spm Ou So! 


Distribution.—Vera Cruz to St. Thomas. Texas region of 
the Gulf. 


Genus OOCORYS Fischer 


abyssorum Verrill and Smith, Tr. Conn. Acad. Sci., 6, p. 177, 
(not the fig, which is O. sulcata), 1884, (as Benthodolium); 
Dall Balls MI COZ Ts pez ou eso pull 27.7 Un nse IN, 
JOE Ga em tidroxeyy 
Distribution.—Northern part of the Gulf of Mexico, U. 5. 
Fish Com. station, No. 2400, in 169 fms. Also off Caro- 
lina. Max. depth 2221 fms. 


MOoLLUSCA OF THE GULF OF MEXICO 88 


Genus CASSIS Lamarck 


cameo Stimpson, Silliman’s Am. Jour. Sci., p. 443, 1860; Tryon, 


Mame hor 27 olen vten SOs 3 Sau aliM alana) Use IN: 
Met Da lsan SO: 
Distribution.—Hatteras to Barbados and West Fla. 


tuberosa Linnzeus,Syst. Nat., ed. X, p. 735, 1758, (as Buccinunz) ; 


Reeve Conch, Icon.) 5) pli 3) 7 f) 74848; “iryon, Man. 
Conch 7 (pe. 27 Pl. 2h f)50.0 795 moat Bulk seo. IN 
M., p. 134, ’89; Dall and Simpson, Bull. U. S. Fish Com., 
PP Ww4Loy LOOL. 


Distribution.—Hatteras to Barbados and West Fla. 


(Cypraecassis) testiculus Linnzeus, Syst. Nat., ed. X, p. 736, 


MAG SUAS OeccLTUMe\\ Neeve. | ConchuwMliconme V5) plyiants TO} 
HOS wLEVvon | Manis 7p) 272. ple 2iwteoa pls Anite O25) (85) 
Dall, Bull. 37, U.S. N. M., p. 134, ’89; Dall and Simpson, 
Bulle WS) Mish Coma) p: 418) Tro]8n 


Distribution.—Hatteras to Trinidad, W. I., and the Texas 
region of the Gulf. 


(Semicassis) inflata Shaw, Nat. Misc., 5, p. 22, pl. 959, 1812; 


ivony Man 7 pura pla ihe OS es Dall Bulli 37. U: 
S. N. M.,p. 134, ’89; Dall and Simpson, Bull. U.S. Fish 
Com., I, p. 418, tg0I; Mitchell, List Tex. Shells, p. 6; Van- 
atta Ao NaSa Phila: |) 55...) 758 0rOO3: 


Distribution.—Hatteras to Brazil, Gulf coast: Calhoun Co. 
West Fla.; Point au Fer, Chandeleurs, Lost Island, La.; 
Galveston, Tex. Pleistocene, New Orleans pumping stze- 
' tion No. 7,Knapp’s No. 2 well, Terrebonne Parish, at 1791- 
1842 feet. 


Genus ONISC {DIA Swainson 


dennisoni Reeve, Pr. Zool. Soc., p. 21, 1842, (as Onzscia); Tryon, 


Manne p 252, plato: i 20;17 85-4 Dally) Bulls MeiC. Ze yrs, 
Pa Zensoe 


D 


BULLETIN 8 Es) 


w 
OQ 


Os 


Distribution.—Arrowsmith Bank, Yucatan, at 130 fms. 
Also Guadelonpe Island. (Type locality). 


Genus TONNA Brunnich 


galea Linnzeus, Syst. Nat., ed. X, p. 734, 1758, (as Buccinum); 
Reeve, Conch. Icon., 5, pl. 1, 1848; Tryon, Man., 7, p. 261, 
pl. 1, f. 3, 7°85; Dall and Simpson, Bull. U. S. Fish Com., 1, 


Pp. 419, 1901. (Allas Dolzzm). 


Distribution.— Hatteras to Trinidad, West Florida, Louisia- 
na, Galveston and Matagorda, Texas. Also Mediterranean. 


perdix Linnzeus, Syst. Nat., ed. - S 734, asBucciuune) ; 
Reeve, Conch. Icon., 5. pl. 6, , 1849, (es Dok cm); ‘Tryon, 
Man., 7, p. 264, pl. 3. £8 15, ie wa 23-25, 785; Dall, Balk: 
s7.0Ul: SN. M., p. 134, "So: 


SG 


Distribution.—West Florida to Brazil. 


Genus PYRULA Lamarck 

papyratia Say, Jour. A. N. S. Phila., 2, p. 238, 182 

Man... 7, p..266, pl. 6, i 35565 > mall) Tr. W_ 2. 
£63, °90; Vanatta, Pr. A. N. S. Phila... 55, p-. 7 


on 

YD 

fed ; 
Oo 

(@) 

Oo wo) 


Distribution.—North Carolina to the West Indies. Gulf 
coast, Calhoun Co., Fla. and Texas. Pleistocene, New Or- 
leans pumping station No. 7 and of Fla. Pliocene, Caloos- 


ahatchie River. 


Genus STROMBUS Linneus 


pugilis Linnzeus, Syst. Nat., ed. X, p. 744, 1758; Reeve, Conch. 
Icon., 6, pl. 16, f. 39-41, 1851; Tryon, Man., 7, p. 109, pl. 2 
f. 13-15, °85; Dall, Tr. W. I. S., 3, p- 177, °90; Singley, 4th 
Ann. Rept. Tex., p. ou "92; Dall ae Simpson, Bull. U. 
S. Fish Com., 1, p. 422, - Vanatta, Pr. A. N. S. Phila., 


L20 MoLLUSCA OF THE GULF OF MEXICO go 


DAS on 1903; Vanghan, Publ, No, 123) Carn. Inst.,)-p: 
sifu) Woe), 
Distribution.—Hatteras to Colon. Gulf coast; Cedar Keys 
and Calhoun Co., West Fla.; Point au Fer, Chandeleurs, 
Cameron, La., Galveston and Corpus Christi, Tex. Pleisto- 
cene, Grand Chenier, La., New Orleans pumping station 
No. 7; North Creek, and Labella, West Fla. Pliocene, Ca- 
loosahatchie River. 

costatus Gmelin, Syst. Nat., p. 3520, 1792; Tryon, Man., 7, p. 
Os 1So57 Dall, Bull. 37, U. So iNe Men ps 136; 307) Dall and 
Simpson, Bull. U. S. Fish €om., 1, p. 422, 1901. 

accipitrinus Reeve, Conch. Icon., 6, pl 7, f. 12, 1850; Tryon, 
Vie munp LAN Tents, 75. 
Distribution.—St. Augustine, East Fla. to Guadeloupe, W. 
I. Also West Fla. 
Genus SEGUENZIA Jeffreys 

monocingulata Seguenza, Boll. Comitato Geologico, 7, p. 188,’76; 
Watson, Chall. Rept. Gastr., p. 105, 86; Dall, Bull. M. C. 
Zon, US iO, Ay, SOP io Leos, UNG INC. Go, iMiay Fall Kors sic 
88, 89, 18869. 

formosa Jeffreys, Rept. Valorous Cruise, p. 200, 1876; Dall, 

SGN WL (CL AL) JO, Ale texe= 


Distribution.—Western North Atlantic and Bay of Biscay. 
Gulf of Mexico, West Florida and Texas regions. 100-2023 
fms. 

trispinosa Watson, Jour. Linn. Soc., 14, p.. 591; Chall. Rept. 
Cashes Plants 14s 01Ss0;” lnyont\ anny sO. uple 
Cai 7 Osoo, 1so7) Dally Bully 27a Uni Sm Ne Mien. 42) 
1880. 
Distribution.—Hatteras to Pernambuco, Brazil and the Tex 


region of the Gulf. 294-675 fms. 


No) 
i) 


BULLETIN 38 Tat 


iconica Watson, Jour. Linn. Soc., 14, p. 589, 1879; Chall. Rept. 
Gastr 15.) po io7. plaza i ats So7 ml nyonul Man yiouape 
Mohs sollyntswin dane lain crest rectyel SO ANOUN 1 BoUUls Wl) (Ce Aat ies, yoy. BOG). 
1880. 


Distribution.—Gulf of Mexico, Florida Strait region, Culebra 
Isl., W. I., and off the Azores. 390-1568 fms. 


Genus TRIPHOGRA Blainville 


perversa var. nigrocincta C. B. Adams, Jour. Bost. Soc. N. Jel Bs 
p. 286, pl. 4, f. 11, 1839, (as Cevzthzum); Gould, Inv. Mass., 
Pens23,ih- 1592, 013707 DalliPr Ue Sena MOM pag anoge 
Ballis7 WS. IN Me) pln 38, 78osnoimeley, 4th, Anni iepe. 
CUS Mext op. 238i oz Mallia NN ES Muis Dl 2OganO 2k 
Johnson: \Oces Pal Bost) Soc. MNAiHe 7) pe T2O. rons: 


Distribution.—Massachussetts to Barbados. o-30 fms. Gulf 
coast: Cedar Keys, Florida, Corpus Christi, Texas. Plio- 
cene, Caloosahatchie beds, West Fla. 


decorata C. B. Adams, Contr. Conch., p. 177, 1850, (as Cerith- 
Zee Dally Dr Wey iS ase a2 Om Aulsoe2e 


Distribution.—Gulf of Mexico, north and east shores, and 
the Antilles. Also Lower Miocene of the Chipola marl, 
West Florida. Variety oltvacea Dall, (Bull. M. C. Z., 18, 
Pw24AN TS8o7 Bull 275 USING ON ep hl 84) 8O)) i Gltlisot 
Mexico, west of Florida, in 50 fms. Also Key West and 
Antilles. 


colon Dall Bull). Mo CZ. vol RSGOMs ers ES) p. 247.) pli zou. 
T2NMS Oe Bulla 7 We So Ne Vien ane sen ple) 2Os) te 2 SOE 


Distribution.—Yucatan Strait, 640-1002 fms. Also off 
Havana, Cuba, 450fms. 


triserialis Dall, Bull. M. C. Z., 9, p. 84, 1881; 18, p. 246, pl. 20, 


122 MoLbLusca OF THE GULF OF MEXICO 92 


Mesa Oa 18897 (Balls U1 Ss New perasapl20, 6.054, 
6a, 1808. 
Distribution.—Yucatan Strait,640 fms. Also off Barbados, 
154 fms. 


bigemma Watson, Jour. Linn. Soc., 15, p. tot, 1880; Dall, Bull. 
Me CH Z anon ps Si.) 18er. Watson, Chall Repr.,Gastr. yp: 
Roermieageh. 6. 785". Dall Bully MiCa Zins) p.243,) 789. 


Distribution.—Vucatan Strait, 640 fms. Also off St. Thom- 
ase Wel) 200 fms; 


hmcus all Bull. M- C. Z., 9, p. $3, 18813) 18, p. 249, pl. 20, f. 
ISOM OU a7 Oi Sc Ne Mi ps T3Z8s pl 20%) fo iI SO, 
Distribution.—Yucatan Strait, 640 fms. Perhaps a variety 
of dzgemma Watson. 


abrupta Dall, Bull. M. C. Z., 9, p. 84, 1881; 18, p. 249, pl. 20, f. 
©, "BOs Iebly Sar Wk) See ING IME jon 1624S), Olly BO, 3s Cy evo 


Distribution.—VYucatan Strait, Cape San Antonio, 640 
fms. 


torticula Dall, Bull. M. C. Z., 9, p. 82, 1881; 18, p. 249, pl. 20, 
Pore so Biull aes 5. No NE pains se pl 20Mfin tb; 
1889. 

Distribution.—Yucatan Strait, 640 fms. 

inflata Watson, Jour. Linn. Soc,, 15, p. tor, 1880; Dall, Bull. 
MICA eno pi Si L881; Watson, \ChalliiRepey CGasth., ‘p: 
FO plow it 285: Dall, Bulli Mi CunZe ss nps i240, 
1889. 

Distribution.—Yucatan Strait, 640 fms. Also off Georgia, 
St. Augustine and Culebra Isl. 294-640 fms. 

inflata var. ibex Dall, Bull. M. C. Z., 9, p. 86, 1881; 18, p. 249, 

(Ol, BOS wig alo esky! 


Distribution.—Yucatan Strait, off Cape San Antonio, in 640 
fms. Also off Havana, 450 fms. 


93 BULLETIN 38 123 


cylindreila Dall, Bull. M. C. Z., 9, p. 83, 1881; 18, p. 250, pl. 20, 
PON. So.) Bulls 27 aU mou INE MEMO. MsSiunli aor talGai2oO. 


Distribution.—Cape San Antonio, Yucatan Strait, 640 
fms. 


Genus CERITHIOPSIS Forbes and Hanley 


greenei C. B. Adams, Jour. Bost. Soc. N. H., 2, p. 287, pl. 4, f. 
Ly 1820. (as) Gerzihzum)) Dallas mila Men on N on Evie aoe 
ESS pl 52 hl2</18Or) Blt NVA OMuen DA ZOO" 7O2- i) Ohl Soms 
OccewPaisBosts Soc. Ne ee .7.) pale On mols 


Distribution.—Prince Edward Island to Dominican Repub- 
lic and west of Cedar Keys, Fla., Corpus Christi and San 
Antonio, Texas. 3-10 fms. Pliocene, Caloosahatchie River, 
Florida. 


crystallina Dall, Bull, M. C. Z., 9, p, 89, 1881; 18, p. 254, pl. 20, 
fh SO; Hell 3 7 UL Se7 NEM ON tS. lin2Oumtin 2Ommea 
°89; Dall and Simpson, Bull. U.S. Fish Com., 1, p. 424, 
IQOl. 

Distribution.— Washed ashore dead, Cedar Keys, Florida, 
dredged alive west of Florida in 50 fms. Common in An- 
tillean dredgings. 50-805 fms. 


martensi Dall, Bull. M. C. Z., 18, p. 255, pl. 20, f. 2, ’89; Bull. 
27 UL SE INE ME pe 28 plea oust h iSO: 


Distribution.—Gulf of Mexico, west of Florida, at 1181 
fms. 


vanhyningi Bartsch, Pr. Biol. Soc. Washington, 31, p. 135, 
1918. 
Distribution.—Tampa Bay, West Fla. 

burkevillensis Dall, Pr. U. S. N. M., 46, p. 231, pl. 22, f. 5, 
IQI4. 
Di stribution.—Upper Miocene or Lower Pliocene, Burke- 
ville, Texas. 


124 MOLLUSCA OF THE GULF OF Mexico 94 


(Eumeta) subulata Montagu, Test. Brit. Suppl., p. 115, p!. 30, 
PVG WTSOSMiCasidu7ex) Dall Balle MAC i Zar Bi) ps) 252i: 
ZONA SO UM a 7n MOL Sa Ne Me De bao PLY 2x ti. )i4) 
PINS 2 hei Sor jonnsou,\|Ocel Pa Bost) SOC IN.) El.) 7), p- 
LZ0 VOWS: 

emersoni C. B. Adams, Jour. Bost. Soc. N. H., 2, p. 284, pl. 4, 
f. 10, 1838, (as Cerzthtum) Verrill, Inv. Vineyard Sound, p. 
OAs ple24nnt. 151.1873. 

punctatum Philippi, 1848. Not of Bruguiere, 1789. 
Distribution.—Massachussetts to Grenada, W.I. Gulf of 
Mexico, Cedar Keys, Texas and Yucatan. 2-15 fms.  Plio- 
cene of the Caloosahatchie. 


Genus SE/LA A. Adams 

adamsi H. C. Lea, Tr. Amer. Phil. Soc. 2d ser. 9, p.. 42, 1845, 
(Gs Cerethzm)< Dally: We TS. 2p i2o7.018o2)\)Vanatta 
Pi eae ONS. eladlasiis 5.) ps) (758, LOOgsmloumsons Occ.) Par 
BOSHESOCH ING (ela 7\ py 127. TOs: 

terebralis C. B. Adams, Jour. Bost. Soc. N. H., 3, p. 320, pl. 
aut. )74 1840 (as Gertthium): De) Kay, IN2)Y.) Moll) p. 130, 
OIoe ie ae. DalluBulls MIC. Ze isapaz5ou soul: 
SeoONE Mig pins 8) pole 52).°£.15,)) (SO NOt Cemsrerume icercunale. 
Lamarck. 

Distribution—Massachussetts to Samana Bay, Dominican Re- 
public. o-20fms. Gulf coast: Cedar Keys, Fla.; Chande- 
leurs, La.; Galveston and Corpus Christi, Texas, 0-20 fms. 
Pleistocene, Gulf coast. Pliocene, Caloosahatchie River, 
Fla. Lower Miocene, Chipola marl, West Fla. 


Genus CERITHIUM Bruguiere 
floridanum Moerch, Mal. Blatt:, 23, p. 114, 1876; Dall, Bull. 37, 
WAS TONG Me os 240) 18. Lr Welle Siete oS 2nn pla Tale ts 
ROW so2- 1 Vauoham Publ Now nagar last) pa) Tn 
IgIo. 
Distribution-—Hatteras to Cuba and West Fla. Pleistocene, 
Labella, West Fla. Pliocene, Calaosahatchie beds. 


95 BULLETIN 38 L25 


algicola C. B. Adams, Pr. Bost. Soc. N. H., 2, p. 5, ’48; Tryon, 
Mant9, pa l2o.pl22 i380) So. 8o- Dall ire Nia elaa sia ae 
Py2825));92. 


Distribution.—Tampa and Charlotte Harbor, West Florida 
to Jamaica. Between tides. Pliocene, Caloosahatchie 
beds. 


litteratum Born, Mus. Cees. Vind., p. 323, pl. 11, f. 14, 15, 1780; 
Dall and Simpson, Bull. U. S. Fish Com., 1, p. 425, 1901. 


Distribution.—East and West Florida, Antilles and 
Europe. 


muscarum Say, Amer. Conch., 5, pl. 49, f. 1, 1832; Tryon, Man., 
Ook ahead ener alee 3D ent. Iie NW Ih Sun iy) 0, Oe, 
Vaughan, 2d Ann. Rept. Fla. G. S., p. 148, 1909. 


Distribution.—Florida to Jamaica. Gulfcoast: Cedar Keys 
and St. Mark’s, Fla.; Chandeleurs, La. Pleistocene, Mana- 
tee, Labelle and North Creek, Fla. Pliocene, Caloosahat- 
chie beds. 


variable C. B. Adams, Pr. Bost. Soc. N. H.. 2, p. 5, 1848; Sow- 
ery, Conch! icon... 15 ply igiieronamso5: Dall) Bille yore 
U. S. N. M., p. 140, 1889; Mitchell, List Texas Shells. 


Distribution.—Tampa, Fla.; Galveston, Corpus Christi, 
Carancahua and Espiritu Santo Bays, Tex. South to the 
Antilles and Curacoa. 


minimum Gmelin, Syst. Nat., p. 3564, 1792, (as Murex); Tryon, 
Man., 9, p. 167, pl. 34, f. 9-11, 1887; Dall and Simpson, 
BallU.S. Fish Com: 174260 100%. 


Distribution.—Tampa, Fla., to Guadeloupe Isl. 


minimum var. nigrescens Menke, Synopsis, p. 85, 1828; Dall, 
Boll27. 00! SouN. Mee pamnao a So) 


Distribution.—Tampa to Venezuela. 


eburneum Bruguiere, Ency. Me’th., pl. 442, f. 1a, b; Tryon, 


126 MOLLUSCA OF THE GULF OF MEXICO 96 


Maite won pat Zou plole tn 7 lA 72 Nn noo 7m Dalla tila a7 sins. 
N. M., p. 140, 1889; Singley, 4th Ann. Rept. Texas Surv., 


PaissOn L892! 
Distribution.—Florida to the Swan Isls. Cited by Singley 
from Corpus Christi, Texas. 

galvestonense Harris, Bull. Amer. Pal, vol. 1, p. 104, pl. 4, f. 9, 
ga, 1895. 
Distribution.—Upper Miocene, Galveston well at 2236- 
2920 feet. 

Genus CLAVA Martyn 

chipolana Dall, Tr. Wagner Inst. Sci., 3, p. 290, pl. 22, f. 8, 

1892. 


Distribution.—Lower Miocene of the Chipola marl, Alum 
Bluff, Fla., and of the Bacom well, Mobile, Ala., at 1241 ft., 
Chipola horizon. 


Genus CERITHIDEA Swainson 
costata Wood, Mal. Blatt., 23, p. 90; Dall, Pr. U. S. N. M., 6, 
Pees Toos bw a7Ul 2 IN) MEM peniAoNTso: 
Distribution.—Tampa, West Fla., to St. Thomas and Ja- 
maica. 


scalariformis Say, Jour. A. N. S. Phila., p. 128, 1825; Tryon, 
Mansons tos) plage. t1. 73.) S70 Walilanby ull ras TNC 
MEF) Pad, So. 


Distribtion.— Georgia to Cedar Keys, West Fla. 


varicosa Sowerby, Genera Shells, No. 42, f. 5; Dall, List Came- 
ron Shells; Bull. M. C. Z., 18, p. 259,’89 


Distribution. —Texas, Breton Island, Chandeleurs, La., and 
south to Cuba and Jamaica. 


turrita Stearns, Pr. Bost. Soc. N. H., 15, p. 24, 1872; Tryon, 
Manving.cp. Tod ply sau hk 82.0787 Dalla dm We TenSss 20 pe 
290,1892. 


Oy BULLETIN 38 I27 


Distribution.—Cedar Keys, Fla., to the Bahamas. Pliocene 
of Shell Creek, Fla. 


hegewishi Philippi, Mal. Blatt., 23, p. ’89; Zeitschr. fur Malak., 
Pp) 19,1841; Singley, 4th Ann.) Rept!) Dex.) pi aso, \18o2e 
Mitchell, List Tex. Sh. 


Distribution.—Cited by Singley and Mitchell from Matagor- 
da, Carancahua and Corpus Christi Bays, Texas. 


Genus POTAM!IDES Brongniart 


matsoni) Dally) Pr. S, .Ne OM taonpayzennpl. 2a.) fo) Tonge 
1914. 


Distribution.— Well near Alexandria, La., at 49 feet; well 
at Pine Prairie, La., at 1540 feet; surface near Burkeville, 
Texas. 


Note.—The brackish water molluscan fauna of Burkeville, 
Texas; Alexandria and Pine Prairie, La., was referred in 
1914 by Dr. Dall to the Pliocene. Dr. W. D. Matthew thought 
the mammalian remains (tibia of a rhinoceros and upper 
molar of a horse, either Protohippus or Merychippus) indicat- 
ed a Late Miocene or Early Pliocene age. For discussion of 
the Burkeville beds (lower member of the Fleming clay) see 
Dumble, Univ. Tex. Bull. No. 1869,pp. 224-225, 1918. The 
stratigraphic relation of the Burkeville beds rather sug- 
gests an Upper Miocene horizon. 


matsoni var. gracilior Dall, Pr. U.S. N. M., 46, p. 231, 1914. 


Distribution.—Well near Alexandria, La., at 49 feet; well 
at Paririe Bluff, La.; near Burkeville, Tex. 


Genus ALABINA Dall 


adamsi Dall, Bull. M. C. Z., 18, p. 258, ’89; Bull. 37, U.S. N. 
IME pro, S77 MVM Oe namo Nr2/70.. 7.92) 1 (UAUllanas 
Bittium) . 


128 MoLLUSCA OF THE GULF OF MEXICO 98 


Distribution.—Hatteras to Haiti and West Florida, moder- 
ate depths. Pliocene. Caloosahatchie River and Shell 
Creek, West Fla. 


cerithioides Dall, Bull. M. C. Z., 18, p. 258, ’89; Bull. 37, U.S. 
INEM OIA: So: Nebr Wl S03 conn 765) Digimon su 7O2: 
Dall and Simpson, Bull. U. S. Fish Com., 1, p. 427, 1901. 
(All as Bztttum cerithioide). 


Distribution.—Cape Lookout, N. C.. to Samana Bay, Do- 
minican Republic. Singley lists with a question a shell from 
Corpus Christi, Tex. Pliocene, Caloosahatchie River, Fla. 


Genus BITTIUM (Leach) Gray 


varium Pfeiffer, Arch. fur Naturg., p. 256, No. 139, I840, (as 
Cerithium); Tryon,Man., 9, p. 152, pl. 29, £. 86, °87; Dall, 
a VS Te Seale. 27/4. 92:) Dall andi Simpson, Bulle Us, S: 
Fish Com., 1, p. 426, 1901. 


wnoocrulum C.. Bs Adams, Pr. Bost. SociNe (HY pis, 184.5; 
Sowerby’s Thes., p. 876, f. 210, 211, (as Cerzthium). 


Distribution.—Chesapeake Bay to St. Thomas, W. I. Gulf 
coast: Cedar Keys, Fla.; Chandeleurs, La.; Corpus Christi, 
Galveston and Laguna Madre, Tex. Pleistocene, North 
Beach. Fla. Pliocene, Caloosahatchie and Shell Creek, 
Florida. 


Note.—&ittium nigrum Totten, synonym of &. alternatum 
Say, has been erroneously reported from Cedar Keys, Fla. 
and Chandeleurs, La. These specimens were probably 
varium. ‘True alternatum, tide Dall, does not extend south- 
ward of New York Harbor. 


DoiplexDall, Tr Woale'S.))3, p.1275)) pls arta 92: 


Distribution.—Lower Miocene, Chipola marl, Fla., and of 


the Basom No. 1 well, Mobile, Ala. at 1500-1556 ft., Chipola 
horizon, Aldrich’s collection. A varietal form. 


99 BULLETIN 328 129 


galvestonense Harris, Bull. Amer. Pal.,vol. 1, p. 104, pl. 4, f. 8, 
1895. 
Distribution.—Upper Miocene, Galveston well at 2550-287¢ 
feet. 
Genus MODULUS Gray 


modulus Linnzeus, Syst. Nat.. ed. X, p. 757, 1758, (as Trochus); 
Dall, Bull. 37, U. S. IN. M:, p: 142,/1889; Singley, 4th Aun: 
Rept. Tex. G. S., p. 339, ’92; Dall and Simpson, Bull. U. 
SH ish Comp 4 27) ar ook 


lenticularis Chemnitz, Tryon, Man., 9, p. 261, pl. 48, f. 91,92, 
1887. 


Distribution.—Hatteras to Brazil and to Cartagena, Colom- 
bia. Gulf coast: West Florida, Chandeleurs,La.; Galveston, 
Corpus Christi, and Espiritu Santo Bay, Tex. Between 
tides. Pliocene, Caloosahatchie and Shell Creek, Fla. Up- 
per Miocene, Galveston well at 2552-2871 feet (identifica- 
tion doubtful). 


floridanus Conrad, Amer. Jour. Conch., 5, p. 107, pl. 12,f. 6, ’70; 
ulgard, House of Rep ex Woc.y 1 pt. 2))\p) S87. ase 
Dall Pr OS! IN NEO on ees oe. Bull’.37 Un SauNe 
MO pada 24( 789; Tr OW IS eae ip 205.) \92° Vaugdaanmy 
Bubly No. 133) Carn. Inst) joao no: 
Distribution.—Hatteras to St. Thomas, W.I. Gulf coast, 
Cedar Keys and St. Mark’s, West Fla. Pleistocene, Lake 
Borgne borings, La., and North Creek and Labelle, Fla. 
Pliocene, Caloosahatchie beds, Fla. and of the New Orleans 
Gymnasium club well at 1200 feet. 


Genus PACHYCHEILUS Lea 
anagrammatus Dall, Proc. U. S. N. M., 46, p. 232, pl. 21, f, 
SOLO LA. 


Distribution.—Upper Miocene or Pliocene. Well near 
Alexandria, La., at 49 feet. Also near Burkeville, T cxas, 


130 MOLLUSCA OF THE GULF OF MEXICO 100 


surface exposure. Brackish water formation. 

satillensis Aldrich, The Nautilus, 24, pt. 11, p. 132, pl. 8, f. 1, 
HA-CmMOl was eoramcaes)):) Wall ProvUis SN Mis p22) 
1914, 
Distribution.—Upper Miocene or Pliocene. Satilla River, 
Ga., and Burkeville, Tex. Brackish water formation. 


Suavismoale roc. We) S. NM: 46) \p.' 232). pl!) 2) £16) 9; 2614. 


Distribution.—Upper Miocene or Pliocene. Well near Al- 
exandria, La., at 49 feet, and well, Pine Prairie oil field, 
La., at 1540 feet. Also surface exposure, Burkeville, Tex. 


Genus CAECUM Fleming 


floridanum Stimpson, Pr. Bost. Soc. N. H., 4, p. 112, 1851; 
Mayon Man, /Se pi2n7)\ ply 66,4) 620786; Dally Bully a7. 
WHRSHEN Moura Boe dir Wet Omak mao sn 72. 


tmnegmlare de Kolin, Jour. de Conch.) 15))\p)) 46) pl.3,/.f. '6, 
1867. 


annulatum Emmons, Rept. Geol. N. Car., p. 274, f. 190,’ 58. 
Not of Brown, 1844. 


Distribution.—Hatteras to Bahia, Brazil. Also West Fla. 
2-18 fms. Pliocene, Caloosahatchie, Fla. 


instructum de Folin, Fonds de la Mer; Dall, Bull. 37, U.S. N. 
NV De aLhAZ SO: 


Distribution.—Hatteras to Tampa, West Fla. Pliocene to 


Recent. 
bipartitum de Folin, Fonds dela Mer; Dall, Bull. 37, U.S. N. M., 
Pi 142, 80. 


Distribution.—Hatteras and West Florida. 


carolinianum Dall, Trans. Wagner Inst. Sci., 3, p. 300, pl. 22, f. 
Zs rso2s itis 37. WY iS.) Ney Mee haa NES so. (maine 
only). 


IOI BULLETIN 38 13k 


Distribution.—Hatteras to Tortugas and to Egmont Key, 
West Fla. 2-63 fms. Pliocene, Caloosahatchie River, 
Florida, 


glabrum Montagu, Test. Brit., 2, p. 497,1803, (as Dentalium); 
‘Anyon,| Mani Sap.) 215.0 pliiGO, Foy rSSor mDallnu baller. 
WEE SOUND V4 2a So. 


Distribution.— North Carolina to Tampa, West Fla. Also 
European. Pliocene of the Caloosahatchie and of England. 
Miocene of Yorktown, Va. 


cooperi Sanderson Smith, Ann. & Lyc. N. Hist. N. Y., 7, pp- 
L544 LOS T8025..0,) (Py BOs ues 7 Oslin Oi. ania Sap: 
22TH DU O7, ik SA Sse alltguAerNN ein SiS. ZOOM MS OZ. 
Wanatta, Pro Al Ni S) imine igs. 7158.) 1903; Johnson, 
OceiPa. Best. Soc) INTE omet2 An he 15) 


costaium Neri) (Aa jour Setangs)i snips 253) pl: 0, tee 
1872. 


Distribution.— Massachussetts to the Antilles. Gulf coast: 
Crooked Island, Calhoun Co., West Fla. Pliocene of the 
Caloosahatchie and Shell Creek, West Fla. 


pulchellum Stimpson, Pr. Bost. Soc. N. H., 4, p. 112, 1851; Shells 
of New England, p. 36, pl. 2, f. 3; Hilgard, House of Rep. 
Bx Doc. \1,°pt:) 2) p.  S8oumerouy anyon, Man. 'Siaosmiam 
pi 661.63.) 786; Dalle mB allie anOretsy ON. Mi... wea orale 
KOwit. 122.) So: \Vanattan | bana Neon EP hilal scum mages 
1903; }ohnson, Occ) PaiBostasocweNey El.) (74). 2 ahnolsn 


Distribution.—Cape Cod, Mass., to the Tortugas. Gulf 
coast: Crooked Island, Calhoun Co., Fla. Pleistocene, Lake 
Borgne, La., borings. 


Genus MEIOCERAS Carpenter 


deshayesi de Folin, Annales Soc. Linn. Maine et Loire, 11, ’69, 
Dal B all 275) SOL SN leis nig 8 Oy 


Distribution.—Tampa, West Fla., to Jamaica, W. I. 


132 MoLLuscA OF THE GULF OF MEXICO 102 


undulosum de Folin, Ann. Soc. Linn. Maine et Loire, 11, 1869; 
Dall Balls Ueto NS VE pw 14.2030) 


Distribution.—Charlotte Harbor, West Fla. to Jamaica. 
Pliocene to Recent. 


nitidum Stimpson, Pr. Bost. Soc. N. H., 4. p. 112, 1851, (as 
Guecum)- Tryon, Man.. 8, p. 222) pl) 67, f. 73; 86; Wall, 
Miva S30 ps 3O2, 68922 Vanatta brs AUN] on ehilas 
55) P- 758, 1903. 
Distribution.— West Florida, at Tampa and Crooked Isl. 
(Calhoun Co. ), to Jamaica. 


Genus BIVONIA Gray 


exeriay Dall Bull. MM. C. Z:. Marvard Colli;o.-p. 39; 1881; 18, 
pazo4") pl 26, f..6, °8o; Bull. 37,U) SaNyM:, p.) 144) pl. 26, 
Te Ony OOr 
Distribution.—Cape Lookout, N. C., to Barbados. Gulf of 
Mexico, West Florida and Texas regions, deep water. 3I- 
1002 fms. 


Genus VERNICULARIA Lamarck 


spirata Philippi, Arch. fur Naturg. 2, pt. 1, 224, pl. 7, f. 1, 1836, 
(as Vermiculus); Tryon, Man., 8, p. 187, pl. 55, f. 99, 100, 
oor Wall Balla WS. Ne Iv pan ra pl Siete SOs 
MNES) sp) 204, 92; Singleya 4th Amiy Kept, Gi iS) 
slkexa 9.) 340,792) Johnson, Occ) Pal /Bost/SocsN.)H., 7, \p- 
122 1915. 

Distribution.— Massachussetts to the West Indies. Gulf 
coast: West Florida; Matagorda and Corpus Christi Bays, 
and Quintana, Tex. Pliocene of Shell Creek, &c, Fla. 


Note.—Moerch described as lV. melanosclera (Pr. Zool. Soc., 
p- 174, 1861) a form from Vera Cruz which Tryon regarded 
as a variety of sfzvafa, and on the same page of the Proceed- 
ings Moerch described V. guadrangularts from Yucatan. This 
was also regarded by Tryon as a variety of spzvata (See 


103 BULLETIN 38 133 


Tryon’s Manuel, 8, p. 187, 1886). Shells from the Gulf, 
Blake station 36, at 24 fms., have been referred by Dall to 
V. lumbricalis Vann. (See Bull, M.C. Z., 18, p. 261,/ 789) 
but the true Linnean /umbricalts is Oriental. 

nigricans Dall, Pr. U.S. N. M., 6, p. 334, 1883, (as /umbricalis 
Vat. Ball a7 So) Ni ME tos MAAN SOE 


Distribution.— West Florida and Texas. 2-14 fms. 


Genus PETALOCONCHUS Lea 
irregularis d’Orbigny, Moll, Cuba, 1, p. 235, pl. 17, f. 16, 18,’42, 
(as \Vermetus); Dall, sBullea7 iia su Ne Meo p. 144. 188o. 
Cra We S23 spaigonaiogr 
Distribution.—Gulf of Mexico between Mississippi delta and 
Cedar Keys, Fla., at 27 fms. Also Antilles to Guadeloupe. 
Pleistocene of Fla. Pliocene, Caloosahatchie beds. 


varians d’Orbigny, Voy. Ame’r Me'’rid., p. 456, pl. 54, f. 7-10, 
reas nyon! (Mani: Som 7Omi pli gon hy 22.) 220786 allle 
Cree Sas DAO smmoz. 


Distribution.—Southwest Florida to Rio de Janeiro. Plio- 
cene, Caloosahatchie beds. Oligocene, Tampa silex beds. 


erectus Dall, in Agassiz, Three Cruises of the Blake, 2, p. 71, f. 
Z2O7 SS) oulls Mei. Zins apyneo2znpla 2S st.) An SOM taille 
BUM So NEM p44 LSM ineEAN eG. 


Distribution.—Antilles and West Florida. 37-805 fms. 
Pliocene, Caloosahatchie River, Fla. 


Genus SIPHONIUM 


nebulosum Dillwyn, 2, p. 1076, No. 19; Tryon, Man., 8. p. 184, 
plisas it: S75 1886; Dall BullMia 7 Ue S) IN MAY py Ta an Sor 


Distribution.—East and West Florida to Tortola. 
Genus SILIQUARIA Bruguiere 
squamata Blainville, Dict. des Sci. Nat., 49, f. 213; Tryon, Man., 


134 MOLLUSCA OF THE GULF OF MEXICO 104 


SMPALOOMIpLY SS ai aG SOR Mall ys mile Mar MOMSEN en VIO) to: 
144, 89. 
Distribution.—Sarasota, West Florida, to Barbados. 2-163 
fms. 

modesta Dall, Bull. M. C. Z., Harv. Coll., 9, p. 39, ’81; Agas- 
siz, Three Cruises of the Blake, 2, p. 71, f£. 296, 788; Dall, 
PEEON LS IOS Oe AGH aokein Moy iPAetoyiy o) Us waXey) yea Lan PesKo\e | J8yal Ula Ne yriAl Bitsy 
INOS) Peord4. ple) 26. f)4 8o. 
Distribution.—Off Cedar Keys, West Florida, to Curacoa 
Isl. 94-805 fms. 

Genus TURRITELLA Lamarck 

(Haustator) variegata Linnzeus, Syst. Nat., ed. X, p. 767, 1758, 
(asi7z7lo) Reeve, ‘Conch: Icon!,; 5) plas.)f.)19)) 49" bryon, 
Mian Ss ps TOS. pl. Ol. tf. 5857 SOs Wall Bulla Os) 5.1 Ne 
M., p. 144, ’89. 
Distribution.—Matagorda and Espiritu Santo Bays, Texas 
to Cartagena, Colombia, and Antilles. Pleistocene Knapp’s 
wells, Terrebonne Parish, La., No. I at 1660-1700, 2250- 
2450 (?), No. 2 at 1434-1800, No. 3 at 1330-1375 feet. 


(Haustator) yucatecana Dall, Bull. M. C. Z., 9, p. 93, 1881; 18, 
D205, 0pl 20h 2 oo. bull. 27 UL om NGnMEN Dua Agy ple 
BORE ann OO. 

Distribution.—Gulf of Mexico, Yucatan Strait, 640 fms. 


(Toreula) exoleta Linnzeus, Syst. Nat., ed, X, p. 766, 1758, (as 
Turbo); Reeve, Conch. Icon., 5, pl. 6, f. 22, 1849; Tryon, 
Mani \3\ 9p. 205,) pluio8: ‘99, 18386; DullkiBullaa77 Us S. NE 
M., p. 144, ’89; Dall and Simpson, Bull. U.S. Fish Com., 
TDs AZOu TOOT 
Distribution.— West Florida, in 50 fms.; Texas coast. An- 
tilles to Barbados. 45-170 fms. 


(Torcula) acropora Dall, Bull. M. C. Z., 18, p. 264, 1889; Bull. 
BOR SHON: Means TAA SO. 


105 


BULLETIN 38 135 


Distribution.—Hatteras to Grenada Isl. Gulf of Mexico, 
West of Florida, at 14-50 fms., and the coast of Texas, Vera 
Cruz and Yucatan. 3-413 fms. The Pliocene, closely relat- 
ed form is 7. subannulata Heilprin of the Caloosahatchie 
marl. 


chipolana Dall, Trans. Wagner Inst. Sci., 3, p. 312, pl. 22, f. 24, 


1892. 


Distribution.—lLower Miocene of the Chipola beds, West 
Florida, and of the Bascom No. 2 well, Mobile, Ala., at 
1241 feet, Chipola horizon. Aldrich’s collection. 


subgrundifera Dall, Trans. Wagner Inst. Sci., 3. p. 313, pl. 22, f. 


22h TSO2. 


Distribution.—Lower Miocene, Chipola beds, Calhoun Co., 
West Fla., and of the Bascom No. 2 well, Mobile, Ala., at 
1241 feet, Aldrich’s collection. A varietal form was record- 
ed by Harris from the Upper Miocene, Galveston well, at 
2552-2920 feet. 


terebriformis Conrad, Cat. Mio. Foss., Pr. A. N. S. Phila., p. 


5034863; (Name only) Dall ir Wry le Ss.) 2enpa) amen 
1892. 


Distribution.—Lower Miocene, Chipola marl, Fla., and of 
the Bascon No. 1 well, Mobile, Ala., at 1500-1556 feet. 
Aldrich’s collection. Also Chesapeake Miocene of Easton, 
Md. (Conrad’s type locality). 


sajilla Dall, Proc: U.S) NM peieaa\pl 22) f 6.1914. 


Distribution.—Upper Miocene or Pliocene.. Well near Al- 
exandria, La., at 49 feet. Brackish water formation. 


Genus MATHILDA Semper 


yucatecana Dall, Bull, M. C. Z., 9,p. 90, 1881, (as Bzttzum); 18, 


p1/266, pl. 20, £217.77 807 nulla amon ING Mie iplen20wcts 
ae) rexOye 
Distribution.—Gulf of Mexico, Yucatan Strait, 640 fms. 


136 MoLLUSCA OF THE GULF OF MEXICO 106 


Also off Georgia and Fernandina, East Florida. Least 
depth, 294 fms. 


Genus LITTORINA Ferussac 


ziczac Gmelin, Syst. Nat., p. 3587, 1792, (as 7rochus); Reeve, 
Conchwlcon (ron) ply iii £) STS 572 yon viata 70)! 9, 
AT pAAS ie) 5 TSo7s) Dall Bullia7 Wl iS.) Ne M.D L460; 
’89; Dall and Simpson, Bull. U. S. Fish Com., 1, p. 429, 
Igol. 


Distribution.—Florida Keys west to Texas and South to 
Barbados. 


augulifera Lamarck, Anim. s. Vert., 7, p. 54, 1882, (as Phaszan- 
Cia) Dally Bull 37.40. (S) Ney Mei pa i464) 80;) Dall and 
Simpson, Bull. U. S. Fish Com., 1, p. 430, Igot. 


Distribution.— West Florida and Texas to the Antilles and 
Cartagena, Colombia. 


mespilum Muhlfeld, Verh, Berl. Ges. Nat., 1, p. 219, pl. 8, f. 8, 
no24ni(as: Aelia nyon,.. Man. Ovi ps 252s spl-4'5, 1h. 16) 
18S7- Dall Bulle 37. U.S. Ne MES pain4oy So.) Dally and 
Simpson, Bull. U.S. Fish Com., 1, p. 430, 190I. 


Distribution.—Texas to Barbados. 


inrorata. Say, Jour. A. N. S. Phila., (1); 2, p: 230; 1822, Tuo- 
mey and Holmes, Pl. Foss.czSCar:, p: i109, pl. 26, f. 5, 
L857); Bost.—Fl. Hoss. S, Car, por plait. 5, 1860; 
ayon, Man.9,p. 246, pl. 43) £48) 7375) Wally Bull 37, U. 
SUNCOM po 146 ole 69. fu Sonia Wen OH 2D. 13205 
’92; Singley, 4th Ann. Rept. G. S. Texas, p. 340, ’92: Van- 
atta ema ING Sa Plata: (9553) pa 75Sulnooss  jounson, Occ. 
Pan BOSta SOCeINGIEL 7. Pp. LZ LORS: 


Distribution.— Massachussetts to Jamaica. Culf coast: 
Cedar Keys, and Calhoun and Washington Cos., West Fla., 
Chandeleurs, Weeks Island, Point au Fer, La.; Matagorda, 
Espiritu Santo Bays and Galveston, Tex. Pleistocene, New 
Orleans pumping station No. 7. 


107 BULLETIN 38 137 


Fenus LITIOPA Rang 


bombix Kiener, Ann. des Sci. Nat.,p. 203, 1829; Ann. des Sci., 
BOM DA 222 NLA iT aan MMT SMa Sea uMultay: Olntnl MN lealkiey 
OQ PNZSL;) Pal sste 74s Mos WalluuB ull tac nSH NG yIN IGE! 
148, ’89; Singley, 4th Ann. Rept. Tex., p. 340, ’92; Dall, 
Bullen.) We Se Ne Mp Hinssui room lohnson, |Occy bas 
BOSE SOC WIN Mie 7 Mei LO Munon 5 


melanostoma (Rang) Sumner, Bull. U. S. Bur. Fish., 31, pt. 
Zep ZO LOL: 
Distribution.—New England to Brazil. Gulf coast at Cam- 


eron, La.; Corpus Christi and Galveston, Tex. Also Pacific, 
California coast. Pelagic on floating Sargassum. 


Genus TECTARIUS Valenciennes 


muricatus Linnzeus, Syst. Nat., ed X, p. 1758, (as 7urbo); Reeve, 
Conch Icon.,) 10) pla. aera aSs7e(as “ztior77a)- Tryon 
Man',9, p.258, pl. 48, £:°68,\1887>))Dall, Bull. 37; Ul S2iNe 
M., p. 146, ’89; Dall and Simpson, Bull. U. S. Fish Com., 


WP WABT LOOT: 
Distribution.—Hast Florida and West Florida and Antilles 
to Colon. 


Genus ECHINELLA Swainson 


nodulosa Pfeiffer, Tryon, Man., 9, p. 258, pl. 48, f. 75, 1887, 
(Es<cliisyn.): Dall Bulk WaiSiwiNe Wwe) po) 146.) ?8o: Dall: 
and Simpson, Bull, U. S. Fish Com., 1, p. 431, 1901. 


Distribution.— North Carolina to Barbados and Texas. 


Genus ISAPIS H. & A. Adams 


ebsoleta)) Dall (ProciyU) Sa uNiiiMneinA ON ol 222) umplliiao titi oe 
IQT4. 
Distribution.—Upper Miocene or Pliocene. Well near Al- 
exandria, La., at 49 feet. 


138 Mou.usca OF THE GULF OF MEXICO 108 


Genus ALASA A. Adams 


tervaricosa C. B. Adams, Pr. Bost. Soc., N. H., 2, p. 6, 1845, 
(as Rissoa); Moerch, Malak, Blatt., 23, p. 57, 1876; Dall, 
Bull. U. S. N. M., p. 146, ’89; Dall and Simpson, Bull. U. 


Swish Comet pyaar.) T9OT. 
Distribution.—Tampa, West Fla., to Haiti. 
Genus ARCHITECTONICA Bolten 
granulata Lamarck, Anim. s. Vert., 7, p. 3, 1822; Ency. Me'th., 
PMA. tf Sab om792)) (as Solazzum) Wall ire NV .5° 3,0) 
Beo) T8920) Bo Ss. INU My) 1374p) 232n nolo; Mlauny. Bull: 
mers Paley NO4)295) (ps1 Dl. 23, tsa onli 
perspectiva Tuomey and Holmes, Pl. Foss. S$. Car., p. 120, pl. 
26, 1857. Not of Linneeus nor Lamarck. 


Distribution.—Hatteras to the Antilles. Gulf coast at 
Chandeleurs, La., Matagorda and other localities, Tex. 
Also Pacific Ocean, Lower California to Peru. Pleistocene, 
New Orleans pumping station No. 7; Upper Miocene, Gal- 
veston well at 2158-2871 feet. Lower Miocene, Chipola 
marl, West Fla., and Dominican Republic. 


Genus TORINIA Gray 


cylindrica Gmelin, Syst. Nat., p. 3572, 1792, (as 7vochus); Tryon, 
Manion pai7pluisd. 82, 18875 Dalle BallniaynnWy is! iN. 
M., p. 148, ’89; Dall and Sampson, Bull. U. S. Fish Com., 


We pase, TOOL, 
Distribution. —Texas to St. Thomas, W. I. 

canalifera C. B. Adams. Dall, Bull. 37, U. S. N. M., p. 148, 
1889. 


Distribution.—Texas to Guadeloupe. 
Genus RISSOA Fre’minville 


nrecipitata Wall, Bull) Me C1207 18) psi27O ply tow f.) 1 e889: 
Bulle VU SNe OME pe 14S plerronp terion SO. 


Log BULLETIN 38 13G 


Distribution.—Gun Cay, East Fla., 498 ims., Yucatan 
Strait at 640 fms. 

acuticosiata Dall, Bull. M. C. Z., 18, p. 280, pl. 19, f. 10, 1889; 
Bell a7 US wNG ME pe SOM ply ToM tno MG Se: 


Distribution.— Yucatan Strait, 640 fms., and between Mis- 
sissippi delta and Cedar Keys, 32 fms. Also off Hatteras 
and Barbados. Probably a variety of R. xanthias Watson 
(Chall Gastr., p. 588, pl. 44, f. 5, 1885). 


Genus RISSOINA d’Orbigny 


decussata Montagu, Test. Brit., p. 399, 1803, (as TZurbo); 
Schwartz von Mohrenstern, Denkschr. d. Kais. Akad. d. 
Wissensch. Wien. 19, p. 80, f. 12, 1860; Tryon, Man., 9, p. 
ro HOMIE AU ain tore IORNIL IRONS AVON (Cu ZAR eS Sun io), DSi, 
1SOn MOL WIWis TS. 2p miaaan moo! 


Distribution.—Cape Fear, N. C., to Haiti and to West Fla. 
2-17 fms. Also Huropean. Variety plazata Dall, Caloosa- 
hatchie Pliocene. 


laevigata C. B. Adams, Contr. Conch., p. 14, 1850; Schwartz 
von Mohrenstern, Denk. Kais. Ak. Wien., 19, p. 111, f. 79, 
Hg60;)Tryou, Man.) 0) palig@ootmlunsouiy 943.0787 2) Dallaire 
Wel S208 ne aa2 nook 
Distribution.—Cape Lookout, N. C., to St. Thomas, W. I. 


Also West Florida. o-22 fms. Pliocene, Caloosahatchie 
beds. Lower Miocene, Chipola beds, West Fla. 


chesneli Michaud, Descr. des Cog. Nouv., p. 17, 1832, (as Rés- 
50g) bryon, Man. oul paiesomplamsOntn 7 3) S75 lDallenmerst 
WIS. 3p: 343) o2-otueleyanatiay Anim. Rept lexenaos 
BAM Mn O2: 
Distribution.—Hatteras to Guadeloupe, Gulf coast: Cedar 
Keys, Fla.; Corpus Christi, Tex. Also Mediterranean and 
Mauritius. Pliocene, Caloosahatchie beds, West Fla. 


browniana d’Orbigny, Moll, Cuba, 2, p. 28, pl. 12, f. 33, 35, ’42; 


r40 MOLLUSCA OF THE GULF OF MEXICO TIO 


nyouyy Manes oi ipisgo, pl.) Sour 45. « S7uaniattal, et Ie. 
ING Selatan ss. DMT Sou OOS. 

Distribution.— West Indies and Gulf coast, Crooked Island, 
Calhoun Co., West Fla. (Vanatta). 

elegantissima d’Orbigny, Moll. Cuba, 2, p. 26, pl. 12, f. 27-29, 
Hose Vl myo) Mata iG, 09s 274) pls 50. tai thy Loo 
Distribution-—West Indies and Gulf coast, Chandeleurs, 
ae Cally): 

Sagraiana d’Orbigny, Moll. Cuba, 2, p. 25, pl. 12, f. 4, 5, 1852; 
atnyout Man wionpanssa) pl. 578 ti SOn mosTe: Dalley Bully) 27%. 
Wao NMS Dp. 1/50; 780: 

Distribution.—Florida Strait, to Martinique. Also Gulf 
coast at Vera Cruz, Mex. 


Genus BENTHONELLA Dall 
nisonis, Wall, Bull’ Mi C.'Z., 18, p. 282, 18809; Bull) 27, U.S: 
NEMEC TDs SOMO} 
Distribution.—Gulf of Mexico between Mississippi delta and 
Cedar Keys at 940 fms. 
Gaza Dalal My Ci Z.. 18. p. 2825) 8o: Ballas aWians. IN. Me, 
Ds 2 SOn) jolly gay sen ee soy 
Distribution.—Georgia to Cuba,-West Florida and Texas 
regions of the Gulf. 6-463 fms. 
mschen Dall Bull, MaiCZ.)\ 18) 1282) So, Bulla Us S. NG 
INR JOy SAN exo), 
Distribution.—Gulf of Mexico between Mississippi delta and 
Cedar Keys at 940 ims., bottom temperature 39° F. Also 
off Cuba. 
Genus SKENEA Fleming 
planorbis Fabricius, Fauna Groenlandica, p. 394, 1780; Gould, 


ity AMiaSsie ps 206s) f4563) iS 7Onwo alu alla sau ShoNG 
Mi’, p. 150, pl. 52, f. 18, 1889; Bush, Tr. Conn, Acad. Sci., 


anit BULLETIN 38 TAL 


LO, \p. Too: pls 22) f:'5.'8. 09-1900; Johnson, Occ) Pay Bose: 
SOC INE JE 177 apis TO Me hose 


flelix depressus Montagu. 


Distribution.—Greenland (type locality) to Charlotte Har- 
bor, West Florida. This species is the genotype. 
Genus AMPULLARIA Lamarck 
depressa Say, Dall, Bull. 37, U.S. N. M., p. 150, ’89. 
Distribution.—Georgia to West Florida, Texas and Mexico. 
caliginosa Reeve, Conch. Icon., Mon. Ampullaria,{. 118, pl. 25, 


T8567) Mall, Procs) Us Ss NGWMeon ip. lah s.r Soy:  itliaia ze 
WET SEN Mei pet SOM ms Sor 


Distribution.—Cedar Keys, West Florida to Texas, Mexico 
and Nicaragua. In freshwater swamps. 
pinei Dall, The Nautilus, p. 75, Nov. 1898. 


Distribution.— Homosassa River, Fla. 


Genus PALUDESTRINA d’Orbigny 
aldrichi Dall, Proc. U.S. N. M., 46, p. 234, pl. 22, f. 7, 1914. 
Distribution.—Upper Miocene or Pliocene. Well near Al- 
exandria, La., at 49 feet. Brackish water formation. 
planawAddrich, Nantilus) 24) ptmen|ps) 220, pl: 8) 1) 2amonm: 
Dall wer U.S. (NM AGiipmezas Tora. 


Distribution.—Upper Miocene or Pliocene. Well near Al- 
exandria, La., at 49 feet. Also Satilla River, Ga., (type 
locality). 

curva, Dall. Proc. Uz) SyyNaMe 46.235, (pis 22,44 nomar 
Distribution.—Upper Miocene or Pliocene. Well near Al- 
exandria, Ga., at 49 feet. 

cingulata Dall, Proc. U. S. N. M., 46, p. 235, pl. 22, f. 1, 1914. 


Distribution.—Upper Miocene or Pliocene. Well near Al- 
exandria, La., at 49 feet. 


142 MoLLUSCA OF THE GULF OF MEXICO 12 


turricula Dall, Proc, U. S. N. M., 46, p. 235, pl. 22. f. 9, 1914. 


Distribution.—Upper Miocene or Pliocene. Well near Al- 
exandria, La., at 49 feet. Brackish water formation. 


milium Dall; Proc. U. S. N. M., 46, p. 235, pl. 22, f. 2, 1914. 


Distribution.—Upper Miocene or Pliocene.. Well near Al- 
exandria, La., at 49 feet. 
Genus ASSIMINEA Leach 

auberiana d’Orbigny, Moll. Cuba, 2, p. 8, pl. 10, f. 6, 7, 1842, 
(as\Paludestrina): Wall) Pr: U.S.) Ne WMEs) 6) p. 335, 1883; 
ables WS) INE Met.) TSO Son Miles nus" De SAT, 
1892. 
Distribution.—Cedar Keys, West Florida, and the Antilles. 
Pliocene, Caloosahatchie and Shell Creek, West Fla. 


Genus TRUNCATELLA Risso 


caribaeensis Sowerby (MSS.); Reeve, Conch. Syst. 2, pl. 182, f. 
7, 1842; Binney, Land & Fr-Water Shells, N. A. Smith. 
Mise. Coll., No. 144, pt. 3, p. 98, pl. 198, 1865; Dall, Bull. 
BRE) Ne NE pe ons52,8o;) Dall Sa Simpsont Bully U.S: 
BiShiCom! wn 436, ple 53), £475) LOOM 


Distribution.— West Florida, Alabama, Texas (?) and Mex- 
ico. Also Greater Antilles. 

bilabiata Pfeiffer, Wiegm. Arch., 1, p. 253, 1840; Binny, L. and 
Bie We Shu iN Al pt.) 3, p:/ 991, t)/L99%) L865 Dall, Bull, 37; 
We SeNe ME. wip 152), 89): 
Distribution.—Sarasota, West Florida, to Honduras and 
Cuba. 

pulchella Pfeiffer, Wiegm. Arch., 1, p. 356, 1839; Binney, L. and 


RmenWe Shi Ne AG pt) 3p) oot a2oommsog) Dall, Bully a7. 
Was IN Mvp. 152, So; Dall /éz)Simpsony) Bulls U.S. Fish 


Comin aps 4.304 LOOM: 
Distribution.—Florida to St. Thomas, W. I. Gulf coast: 


113 BULLETIN 38 I4 


i) 


Tampa, West Florida; Galveston, Texas. (Dr. Gurley). 


subeylindrica Linnzeus, Syst. Nat., ed. XII, p. 1248, 1767, (as 
Fleless);\ (Gray, in) Murtom, "Man p! 295) i472, 1185.7; pinueys 
Bandvand Er. Wess sine AGM pts 25 pr OO, am ahZOR noo be 
Dall and Simpson, Bull. U.S. Fish Com., p. 436, 1901. 


Distribution.—Tampa, West Fla., to St. Thomas, W. I. 
Genus CHEILEA Modeer 


equestris Linneeus, Syst. Nat., ed. X, p. 780, 1758, (as Patella); 
ayon (Man 8) paag7 pl.) ai hese 264 S38 oruDall lie. VE 
TAS.) 2419) 248) so2' as wideciarca) rile Gael SEWN» IVES aris 
Pal232. TOTO: 
Distribution.—Hatteras to Barbados, Gulf of Mexico, Blake 


station 2. Also Pacific Ocean, Mexico to Chile. Pliocene, 
Caloosahatchie beds, Fla. 


Genus CAPULUS Montfort 


intortus Lamarck, Anim. s. Vert., 6, pt. 2, p. 18, 1822: d’Orbig- 
mys Moll, (Cubal 1 (pilnS6 plea isd22) 2301842 idiom, 
Mane 8 ip. 13a ipl zoet aig San oor OAc mB willy 27) 1 Wena iN 
ME pal 545) (80: 
Distribution.—Gulf of Mexico, Blake station 5, at 229 fms. 
Also off Florida reefs, Hatteras and Bahamas. 


Genus CREPIDULA Lamarck 


fornicata Linnzeus, Syst. Nat., ed. XII, 1, p. 1257, 1767, (as 
Patella’. Wilgard, House of Rep! Ex. Doc., 1, pt. 2, p. 8387, 
19797 Dall,) Bull 37 (Un Sa aN iis 2) DliAS) ite ype 
BOE.) 235) 2A.) TSSO-) Mid Ee! Sean). 35 03 OZ eoleLeye 
ApwAnn. Rept. exp anime 2| WVanatta,: bt.) Aen Ss 
Bhilays5)) 7.58; noes wjolusonhOce. Pa. Bost. Soc). vrEen 
TPE LOO) LOLS: 
Distribution.—Granada to Cartagena, Colombia. Gulf coast: 
Cedar Keys, St. Mark’s, Ft. Barranca, &c., West Fla., La., 
Galveston and Corpus Christi, Tex. Pleistocene, Lake 


LA4. MOLLUSCA OF THE GULF OF MEXICO 114 


Borgne borings, New Orleans well of 1856, and New Orleans 
pumping station No. 7. Pliocene, Caloosahatchie beds. 
Lower Miocene Chipola beds, West Fla. 


plana Say, Jour. A. N.S. Phila., (1), 2, p. 226, 1822; Tuomey 
andmolmes)) PIN EKossS. (Car. p. Tima please ta 12.0857 
Dalia lee ze UM Ss Ne MeN piso) pl yAsaten ta lin 5Onn ts 
BOM SOn i WV lei Ss.) sh DYE 5 ol O22) Wanabbasiate Au VN) S) 
Phila., 55, p. 752, 1903; Johnson, Occ. Pa. Bost. Soc. N. H., 


Fy [Da Meaty vere) HiGy, 


Distribution.—Canada to Trinidad, W. I. Gulf coast: Cedar 
Keys, St. Mark’s, Fla.; Chandeleurs, La.; Galveston, Corpus 
Christi and Matagorda Bays, Texas. Pleistocene, New Or- 
leans pumping station No.7 and North Creek, Fla. 
Pliocene, New Orleans Gymnasium Club well at 1200 feet, 
and Caloosahatchie beds, Fla. Lower Miocene Chipola beds, 
West Fla. and of the Bascom No. 1 well, Mobile, Ala., at 
1500-1556 feet, Chipola horizon. 


aculeaita Gmelin, Syst. Nat., 7, p. 3693, 1792, (Patella); Tryon, 
Mamie Da 2oh pl. 2o)1t 61-62) 1886. DalllBull Mes .9Z, 
ES W260 1SSOn Vanatta,, Pr. ANS iehilaa S50 pel 758: 
noosy Walk Buallyr1 200). (S.NEMEs pro 2n oom 


Distribution.—Cape Lookout, N. C., to Barbados. Gulf 
coast: Indian Pass, West Fla.,-and Texas. Usually 0-25 
fms., but dredged in Gulf of Mexico, near Tortugas at 539 
fms. Dwarfed and white at that depth. Also Pacific Ocean, 
California to Chile. Pliocene, Caloosahatchie beds, West 
Florida. 

convexay say. Nour ANS. Philas (1) 402.) pl2o7iins22 J Dall, 
JEU 7h. Who SR INIS IME NG os aula mole Evo). w3t,/ is. sis O ea Ate EN, 
Ws Slop, Sy hOse Sven Ve 
Distribution.— Nova Scotia to Florida. Gulf coast: West 
Fla.; Chandeleurs, La.; Galveston, Tex. Pleistocene, North 
Creek, West Fla. 


115 BULLETIN 38 r45, 


Genus CRUCIBULUM Schumacher 


auricula Gmelin, Syst. Nat., p. 3694, 1780, (as Patella); Dil- 
ways Cat.) 2, tonya Lowi all Bulls Nien Cs 72am es ar 
234 /8O;buUlMia 7 UL ony INAIME OES 2 NmS oe 


planatum Schumacher, Essai, p. 182, 1817. 


scutellatum var. auriculatum (Chemn.) Tryon, Man., 8, p. 118, 
DI B2n/ fi 24) i258 Sox 
Distribution. —Cedar Keys, West Fla., and Texas, south to 
Barbados and Northern Brazil, 25-111 fms. Pliocene, Caloosa- 
hatchie beds, West Fla. The analogous Pacific species is 
C. spinosum. 


striatum Say, Jour. A. N. S, Phila., 5,:p. 216, 1826; (as 
Calypiraca); Tryon) Man.) (8) sp. 118; pli 33, £144, 45,-86: 
Dall Boils Mi Ca Zor pe 2 oA eo Ohnson.)) Oecumaar 
IBOSst-.S0ci INGE 7. oul OO NNO 5: 


Distribution.—Nova Scotia to Florida. Cited by Dr. Dall 
from the Gulf of Mexico at Vera Cruz. 


Genus CALYPTRAEA Lamarck 


candeana d’Orbigny, Moll. Cuba, 2, p. 190, pl. 24, f. 28, 29, 

1842, (as /nfundibulum); Dall, Pr. U.S. N. M., 6, p. 335, 

493, 1(as Galerus) Vlnyon Mane no.) pala. ply a Ak itaeon 

Tie rose: Malley Bull iowa one ies lie: 52h SOP 
parvulus Dunker,1875. 


Distribution.—Hatteras to Brazil. 6-52 fms. Gulf of Mex- 
ico, Cedar Keys, West Florida, and Yucatan Strait, 640 
fms., dead shell. 


Genus XENOPHORA Fischer de Waldheim 


conchyliophora Born, Index Mus. Ces., p. 333, 1778, (as Tvochus) ; 
snyon, Mant, 68) pO plo, te SOON oon Dalla walla. 
WAS OUN Mis p. 547 So lr Vial Sia Da1 30048 O2nIiole 
AP TAO, 0a. +.92) BiUlliGowOs SiN Vea Ose ape siusnntelhl, 


14.5 MOLLUSCA OF THE GULF OF MEXICO 116 


SH SeeMiatinven, BillcAmen Pals INOMiZOMhpNmEaaiipl.W2 3. ie 
7) IQIt7. 
agglutinans Lamarck. 

Distribution.—Hatteras to the Antilles. Gulf of Mexico, 
West Florida and dredged at various stations, 14-229 fms. 
Usually not so deep as X. carzdaca and more heavily loaded. 
Pliocene, Caloosahatchie beds, West Fla. Lower Miocene, 
Chipola marl, West Fla. Oligocene, Tampa silex beds, Fla. 
Upper Eocene, Jackson, Miss. Eocene, Wood’s Bluff, Ala. 
Upper Cretaceous, (Ripley) of Prairie Bluff, Ala. The 
most ancient species of the Gulf Mollusca. 


caribaea Petit, Jour. de Conch., 5, p. 248, pl. 10, f. 1, 2, 1856; 
iyo Vian. 8, pw162,.pl..474 f Go Soa Dalles Bull Merc 
ZS ps 261, 89° Dall) and Simpson, Bullanuey Ss Fish 
Comes 438.) TOOL. 

Distribution.—Hatteras to Barbados and Gulf of Mexico, . 
various stations, 14-274 fms. 
Genus NATICA Scopoli 

livida Pfeiffer, Wiegm. Arch., 6, p. 254, 1840; Moerch, Malak. 
Bate 24 pod gi) Dally Bulls Nie CZ knenn pn 20210780): 
Bll 7a Why Sa NMG pet's) 1480: 

jamaticensts C. B. Adams, Contr. Conch., p, 111, 1850. 


proxtma Sowerby, Thesaurus, atica, pl. 8, f. 111. Not of 
C. B. Adams. 


Distribution.—Hatteras to Barbados. Gulf of Mexico, be- 
tween the Mississippi delta and Cedar Keys, 26 fms. 


maroccana Chemnitz, Conch. Cab., 5, p. 270, pl. 188, f. 1905-10; 
Dillwyn Cat. Rec! Sh. 2) \p. 983) (Nonna sinsi7) Dall’) Bull: 
NM Cre Zen ieee 202. 188Os) Billi27 Wun SweN ey VCs Dil s 4. 
1889. 


Distribution.—Hatteras to Barbados and Gulf of Mexico, 
west of Florida. Also European and West African. 


Igy BULLETIN 38 , I47 


canrena Linnzeus (in part) Syst. Nat., ed. X, p. 775, 1758, (as 
Nerita); Reeve, Couch. Icon., 9, pl. 4, f. 14, 1855; Tuomey 
and) Ivolines, Pil Moss. (Ss Caren odin is wap ll i> Sanne an7 inns Suz 
DEVI SxeN ea ea OP INT IGA ou larcyaliieKoe: ACen Nin Ibs S4 
p304.7 025 (Maunye (Bull Amen pales Niow 2 ous ome Amine 
Ber.) 3s KOH KONIG 

pucaccila Conrad, Prva NS! Phila wipe s64.) soa. 

Distribution.—Hatteras to the Antilles, Pernambuco and 
Cartagena. Gulf coast, West Florida and Galveston. Plio- 
cene, Caloosahatchie beds, Upper Miocene, Galveston well 
at 2158-2920 feet. Lower Miocene, Bascom No. 2 well, Mo- 
bile, Ala., at 1241 feet, Chipola horizon. 


alticallosa Dall, Trans. Wagner Inst. Sci., 3, p. 365, pl. 22, f. 
Zon loo2e 


Distribution.—Lower Miocene, Chipola marl, Northwest 
Florida, and of the Bascom No. 2 well, Mobile, Ala. at 1241 
feet, Chipola horizon. 

Subgenus Crypionatica Dall 


pusillaSay, Jour. AlN) S.) Phila’, 24) p.257)) 1822: shialeard 
louse of Rep, Ex. Dochwpta2)) prssowers76., Wallaby 
B 7p On UN EM oe ESAS On meet oO cl SLnipl ea uie El 
Ann.) Rept.! lex. p:\ 240/92 anatear pen sy An Na Ss ieinilaas 
55s D207159s) LO0Ss MOhnson Ocean Boosts: Socy ING ween zea. 
TOS (TOLLS. 
Distribution.—Maine to Florida Keys. Gulf coast: St. 
Joseph’s Bay and Crooked Island, West Fla., Galveston, 
Tex. Pleistocene, New Orleans artesian well of 1866, Lake. 
Borgne borings, Knapp’s wells, Terrebonne Parish, La., No. 
2y/at) 1150-1200, 1330-1470, 1500-1525 feet. | Pliocene 
Caloosahatchie beds, and of the New Orleans Gymnasium 
Club well at 1200 feet. 


Genus POLINICES Montfort 
Subgenus Euspira Agassiz 


tenuis Recluz;) Jour. de! Conch) 1,\\p\)/388; pl: 12, £27, 1350; 


143 MOLLUSCA OF THE GULF OF MEXICO 118 


iyo Mane Sy pussy pl nav 254 88 aie stale Mie" Ce 
EES PK ZO5 SO olay 27) Uk) Sy Neen DS a Logs 
(All as Lunatia). 


Distribution.—Gulf of Mexico, Yucatan Strait, 640 fms. 
Also off Cape Florida, Cape Fear and Cuba, 84-640 fms. 
Also Pacific Ocean, Valparaiso, Chile, Recluz’s type lo- 
cality. 


lestalea Watson, Jour. Linn. Soc., 15, p. 261, I880; Dall, Bull. 
WER ZO pe Osh Sia Watson, | Challneot: | Gasth:.p. 
AAO 27.7. Ss Wall Ball Gv Cea inow pai 215). 0. 
(All as Lunatia). 


Distribution.—Yucatan Strait, 640 fms. Also off Fernandi- 
na, Fla., and Sombrero Isl,, 450 fms, 


inngillae Wall Bally Me CZ, 9, p. 03) 18807) 18a pe 2o5) pl: 27, 
fy) WO, SOs: LSDUlla ery LOS RON PIMC aL ialGyal soils aq ssauenuaa eexo). 
(All as Lunatia). 


Distribution-—Yucatan Strait, 640 fms. Also near Old 
Providence, 382 fms. JL. radiata Watson, may be identical 
with this species. 

hemicrypta, Gabb; Jour, A. N. S. Phila (2).4)p.375, pl. 
OVA 5.18003 Casi /Varzcea));) Dally Bullioqo Weis NE IM. p: 
HOG, Oe Oy Es Cy anoint 

eminwloiues Gabb,) jour, A. N.S.) Philas(2)18.p. 330,) pl: 
Aas) t. 4, 1875. 
Distribution.—Upper Miocene, Galveston well at 2465-2733 
feet, and of the Jennings-Heywood No. 29 well, Jennings, 
La., at 1960-1980 feet. Lower Miocene, Chipola marl, West 
Fla. Oligocene, Tampa silex beds. 
Subgenus NEVERITA Risso 


duplicata Say, Jour..A. N. S. Phila., 2, p. 247, 1822, (as Natica); 
Walls wlliy 7. We Se NOUN. ps) a54ae olsen i262 SO7 Clute 


11g BULLETIN 38 I49 


NUS OMS BE oinoa ors. (Ors) encima Jee. VAL IN, Sy Jelouilian se. 
D759, 1003; Johnson,/Oce; (Pa Bost soc: Ny Ey 74 los. 
IQI5. 


canwpechiensis Recluz, fossata Gould, texastana Roemer. 


Distribution.—Massachussetts to Mexico. Gulf coast: Cedar 
Keys, and many other localities, West Fla., Horn Island, 
Miss.; Point au Fer, Cameron, Chandeleurs, La.; Matagorda 
Bay and Galveston, Tex. Pleistocene, Grand Chenier, 
New Orleans well of 1856, New Orleans pumping station No. 
7, Zigler, No. 15, Jennings, La., at 1350-1481 feet, Jennings- 
Heywood No. 30 well at 1127-1169, 1169-1277 feet, Knapp’s 
wells, Terrebonne Parish, No. 2, at 1150-1200, 1330-1375, 
1443-1618, 1700-1839 feet. Pliocene Gymnasium Club well 
at 1200 feet. Chesapeake Miocene, Maryland to Florida. 
Upper Miocene, Galveston well at 2158-2920 feet. Miocene, 
Crowley No. 25 well, Jennings, La., at 2468-2500 feet. 


Subgenus Payraudeautia Bucq. Dautz. & Dollf. 


nubifa Dall, Bull. M. C. Z., 18, p. 2094, 1889, (as everzia); 
Dall and Simpson, Bull. U.S. Fish Com., 1. p. 439, pl. 58, 
ENO TOO 'G: 


Distribution.—Gulf of Mexico, Blake station 23, at 190 fms. 
Also off Barbados, 140 fms, and Porto Rico. 


Subgenus Mammilila Schumacher 


uberina d’Orbigny, Moll. Cubana, 2, p. 31, pl. 17, f. I9, 1842, 
(as Matica); Moerch, Malak. Blatt., 24, p. 60, 1877; Tryon, 
Man., 8, pl. 16, f. 57, ’86; Dall and Simpson, Bull, U. S. 
Bish) Com!) puidgou moor 


Distribution.—Gulf of Mexico, west coast of Florida, 14-40 
fms., living. Also off Sombrero 54-70 fms. and Porto 
Rico. 


factea Guilding, Tr. Linn. Soc., 17, p. 29, 733; (as Watcina); 
iiyou, Man \'3: 0p: 49 pl 16h ie sooo, allman ai. 


150 


MOLLUSCA OF THE GULF OF MEXICO {20 


U.S.N. M,, p. 156, ’89; Dall and Simpson, Bull. U. S. Fish 
Comyn y P4126 IGOL 


Distribution.—Florida Keys to Porto Rico and Brazil. Also 
Texas region of the Gulf of Mexico. West Fla. (?). 


brunnea Linck, Beschr. der Rostock Sammlung,2, p. 140, 1807; 


Dat ull 27) WS. Ni Ma py 156, 80) 


mamillarts Lamarck, Anim. s. Vert., 6, pt. 2, p. 197, ’22, (as 


Wage). inyon, Mani, (8) p. 43, pli iSite 74860, Dall and 
Simpson, Bull. U. S. Fish Com., I, p. 439, Igot. 


Distribution.—Florida Keys to Trinidad Island and Texas 
region of the Gulf of Mexico. 


Genus SINUNM Bolten 


prespectivum Say, Amer. Conch., p. 175, pl. 25, (as Szgaretus); 


Holmes, Post-Pl. Foss. S. Car., p. 81, pl. 12, f. 16, ’60;Tryon, 
Nine Se DNS 7 pleat Ol VO4u SO. Dalla iilema AOS. 
NEEM ps r56. "So; \Vanatta, Pr. AY Ne Sa Pihilaynsise p. 758: 
1903. 

Distribution.—New York to Martinique. Gulf coast: Cal- 
houn Co., Fla.; Chandeleurs, La.; Galveston, Mustang Isl. 
and Pass Cabello, Tex. Pleistocene, New Orleans pumping 
station No. 7, cited by Harris from Upper Miocene, Galves- 
ton well at 2252-2600 feet. 


maculatum Say, Amer. Conch., pacu7 On) plv25qmn@asiS7sarers) : 


bison, Man.( 8 aps i50y pl. 24,0157 5S nOOmmOale Btnll a aia: 
ese Ne ME pens On) SO: 


Distribution.—Hatteras to Guadeloupe and West Florida, 
shallow water. Closely related to martinzanum Phillipi, 
GAbb: a) Beschrin,) py 144)) plait ha sa) 1 Say) 


minor Dall, Bull. M. C. Z., 18, p. 297, ’89, (as Szgavetus); Bull. 


Bry ORS NM pat 5Ou SO. 


Distribution.—West Florida, Florida Keys and Sombrero 
Isl., W. I. 54-84 fms. 


12 BULLETIN 38 15x 


Genus EUNATICINA Fischer 


semisulcata Gray, Zool. Beechey’s Voyage, p. 137, (as Watica); 
@iyon) Man! 8. piiz2) plight. ga 86 DalluBraile Mey Cia, 
LS ON 200s) SOc Mir Helene snipe a SOnmozs 
fordiana and fordit Simpson. 


Distribution.—East Florida to Porto Rico. Gulf coast, 
Sarasota Bay, West Fla., between tides. 


Genus LAMELLARIA Montagu 


rangi Bergh, Mer., p. 94, No. 8, 753; Dall, Bull. 37,U. S. N. M., 
DAU5O. 1 SO! 


Distribution.—Texas region of the Gulf of Mexico. Ap- 
parently never figured. 


Genus MARSENINA Gray 


amplaVerrill,: Pr U.S) NGM miara) 80-0. Conn: 
NCA WATiS and Sei isi) pels usn plazh tae, (can So Dalle 
Bully 37, U.S. NM, p.1156,)8o.),ohnson: Occ. Pay Bost: 
SocwN He 7). TOO. KOs: 


Distribution.—Maine to Sarasota Bay, West Fla. 


Genus ACMAEA Eschscholtz 


candeana d’Orbigny, In de la Sagra’s, Hist. Pol. y Nat. Isla de 
Cuba; 2) p. 199) pl.25,) f) 1-3.) 1845,u(as-facela)): Dall Bull: 
BG US.) ONE EME pr SON Sorin oll Mamie) Taosiae. 
*91; Dall and Simpson, Bull. U. S. Fish Com., 1, p. 440, 
IQOl. 


Distribution. —Florida Strait to Tobago, W. I. Also West 
Florida and Texas. 


punctulata Gmelin, Syst. Nat., 13, pp. 3705, 3717, 1792, (as 
Ratela): PAlsbry,\eleny ota Mian nmieuiipAins7s) liDlis ature 


1891; Dall and Simpson, Bull. U.S. Fish Com., 1, p. 440, 
Igo!. 


152 MOLLUSCA OF THE GULF OF MEXICO 122 


Distribution.—Key West, Florida to Porto Rico, Guade- 
loupe, and the Gulf of Mexico at Vera Cruz, Mex. 


feucopleura Gmelin, Syst. Nat., 13, p. 3699, 1792, (as Patella); 
Eusbryhnyony Man) 13.7.) 40) plats firo-2n on) Dall & 
Simpson, Bull. U. S. Fish Com., 1, p. 440, 1901. 


melanoleuca Dall, Bull. 37, U. S. N. M., p. 156, 1889. Not of 
Gmelin. 


Genus LEPETELLA Verrill 


tubicola Verrill and Smith, Amer. Jour. Sci., (3), 20, p. 396, 
pon Verll iraConn Acad) Arts andeseluisnqp. 534) ple 
Some 2ou2qaue S25 Wall i Bull iM Ci Za arsip aan eile 2s 
VOM Soe ubln a7 Uli Ss. Ne Mi ph ng eegaplen 25e)t..6,) 70% 
Johnson, Occ. Pa. Bost. Soc. N. H., 7, p. 86, 1915. 
Distribution.— Martha’s Vineyard, Mass., at 130-388 fms., 
to Gulf of Mexico between Mississippi delta and Cedar Keys, 
at 324 fms. 

Genus PHASIANELLA Lamarck 

pulchelia C. B. Adams, Proc. Bost. Soc. Nat. Hist., 2, p. 7, 45, 
(asi Yio) Aryon, Man) 10, p: (264.383 Walle Ball’) Me. 
CIA re Os Sole oOs UBuUll raz.) sO Sa aiING Me Dae TSS. 
1889. 

brevis C. B. Adams. Not of d’Orbigny, 1842. 


Distribution.—Cedar Keys, Fla., to St. Thomas, W. I. Plio- 
cene of South Carolina. 


Genus TURBO Linnzeus 


castaneus Gmelin, Syst. Nat., p. 3595, 1792; Pilsbry, Man., 10, 
pazos, pligsy fi88-8o) 7838. Dall) Bull. i27esUM SNE Mey p: 
m5s- 780°) Dall) and Simpson! Bull Uy Ss-eishy)|Com:)\1,) p: 
A441, 1901. 

Distribution.—Cape Hatteras to Trinidad, W. I. Also West 
Florida in shallow water. 25-295 fms, Pliocene, Caloosa- 
hatchie and Shell Creek, Fla. 


123 BULLETIN 38 5 


a) 


crenulatus Gmelin, Syst. Nat., p. 3575, 1788; Pilsbry, Man., 10, 
DaZ2O2 pina si te on 7O8- Walle Bulla 7 Oe SlNeeN io: aa 
(S08 (Gy VE ME Sia 2e DHi aS ouiio2™ ‘ 


Distribution.—Hatteras to Barbados. 2-30 fms. Also West 


Florida at Crooked Island and St. Andrew’s Bay. Pliocene, 
of the Caloosahatchie, probably a variety cf crenwlatus. 


Genus LEPTGTHYRA (Carpenter) Pease 


induta Watson, Jour. Linn. Soc., 14, p. 715, 1879 (as Turbo): 
Challe Gastr.pi128) plo Oya t issn: 


abide Mall, Bull. Me CUZ. 18 Hp es2nuple 23. tO, SO. 


Distribution.—Hatteras to Martinique, 15-2805 fms. Yuca- 
tan Strait, 640 fms. 


Genus LIVONA Gray 
pica Linnzeus, Syst. Nat., 10, p. 763, 1758, (as Zzrbo); Pilsbry, 
Manta) pi 277.) olin pa 2a eo wiDaliane miles Morn Samet 
M., p. 160, ’89; Dall and Simpson, Bull. U.S. Fish Com., 
LD: 443, 1901. 
Distribution.—Charlotte Harbor, West Florida, to Colon. 
37-160 fms. 
Genus GAZA Watson 
superba Dall, Bull. M. C. Z., 9, p. 49, ’81; 18, p. 354, pl. 22, f. 
Aga, “895, Bulle g\ We SNe mea pA TOON Pls 22) iiayAletias 
1889. 
Distribution.—Gulf of Mexico between Mississippi delta and 
Cedar Keys, 324 fms. Also Barbados. 


fischeri Dall, Bull. Mus. Comp. Zool., Harv. Coll., 18, p. 355, 
Pa 7et. 16.) Sor Bulls ah oMeNe NEEDS OO wOl. i2y7. mt. 
1889. 

Distribution.—Gulf of Mexico, west of Florida and St. 
Lucia, W. 1., 423-426 fms. 


154 MOLLUSCA OF THE GULF OF MExIco 124 


(Callogaza) watsoni Dall, Bull. Mus. Comp. Zool., 9, p. 50, ’81; 
Hoss FO DeNe24atenenTani plana anh. | iaNmcu plan 2duniini 2. 2a 
18809. 

Margarita jilosa Dall, Bull, M. C. Z., 9, p. 42, ’81. 
Distribution.—YVucatan Strait, 640 fms. Also off Cuba and 
Barbados. 

Genus LIOTIA Gray 


gemmia Tuomey and Holmes, Pleioc. Fos. S. Carolina, p. 118, 
pli26,, tf. 4,156, (as 7vechus)-Dall and Simpson, Bull. U.S. 
Fish Com., 1, p. 445, 1901. 

tricarinata Stearns, Proc. Bost. Soc. Nat. Hist., p. 23, for ’72; 
(siavchiiecionica) Wall, Bull. M. CC) Aiars8s i380, 789; 
Bales 7 OW Su IN. Map. 166, 78: 

Distribution.—Hatteras to Porto Rico. Also off Tampa, 
West Fla., 15 fms. Pliocene, Caloosahatchie beds. 


Hanoi DWally Bulls Mi CZ. 18, (p. 280, pl. 334k Bi SO: 
Distribution-—Gulf of Mexico, Blake Station 2, at 805 fms. 


Also off Carolina, Key West and Havana. 


Genus CALLIOSTOMA Swainson 
euglyptum A. Adams, Proc. Zool. Soc., p. 38,54, (as Zizyphinus) ; 
Reeve, Moni Z7zy pi voll 2) t) 17.0085) Halles ule MeN Gs 174 
ROM PA GOS TOO wellibiva Man. napa eyiAiuo le tegtohen 7a sp). 
57, 1. 9, 89. 
Distribution.—North Carolina to Florida, Texas and Vera 


Cruz, Mexico. Erroneously referred by Reeve to Tasmania. 
3-32 fms. Pliocene ancestor is C. philanthropus Conrad. 


circumcinectum Dall, Bull. M. C. Z., 9, p. 44, ’81; 18, p. 364, pl. 
22 eR Baio Billy 270 oe NGM peo Dl 22 uh. nae 
3a, ’89. 
Distribution.—Florida Keys, Antilles and Yucatan Strait. 
640-805 fms. 


125 BULLETIN 38 155 


corbis Dall, Bull. Mus. Comp. Zool Harv. Coll., 18, p. 365, pl. 
Bhai SOs MBM) 277 OSH iING VE De LOD) Une mune omen 
18809. 
Distribution.—Gulf of Mexico, west of Florida, to Jamaica. 
220-450 fms. 


tiara Watson, Jour. Linn. Soc. 14, p. 696, ’79; Rept. Chall. 
Caste) ps 60,) plavOmt. vans suas Pivockiasyie Daly buleyarzs 
We Sane ol nO2ti SO: 


Distribution.—Gulf of Mexico, Texas region, to Dominica, 
W.I1. 220-780 fms. 


roseolum Dall, Bull. Mus. Comp. Zool, Harv. Coll., 9, p. 45, ’81; 
Ices OA CLO 8) ea by ote (OMI ovel. \” sOyae BION. ALON use ING Me INE 70). 
LOZ lI 2A.t.) GunGa aoe 


Distribution.—North Carolina to Yucatan. Also Florida 
Straits and Havana. 21-200 fms. 


pulcherum C. B. Adams, Contributions to Conch., No. 5, p. 69, 
n50;)\(as| Lvochus);\(Pilsbry,) Man) 11/4375) 89. (Malls Bull: 
BT UMS Na Mien sp MOMS O! 


Distribution.—Hatteras to St. Thomas, W. I. Texas region 
Gulf of Mexico. 15-63 fms. ‘Type locality, Jamaica. 


veliei Pilsbry, The Nautilus, p. 128, 1900. 
Distribution.—Caxambas Pass, Southwest Florida. 


(Euirechus) jujubinum Gmelin, Syst. Nat., p. 3570, 1792, (as 
ivockus) ;) Pilsbry;,)Man.\\ 0, psi4o4. ply 40... 16) 7899) Dall 
Bull 37, WU. S2Ne Mp a62 080; all and Simpson Bile 
WS) ish, Com), psjaaa, (toon, 


Distribution.—Hatteras to Cartagena, Colombia. Gulf coast, 
West Florida, Texas and Yucatan. Pliocene, Caloosahatchie 
beds, Fla. 


(Eutrochus) jujubinum var. perspectivum Koch, Philippi, Abbild. u. 


150 MOLLUSCA OF THE GULF OF MEXICO 126 


BESeAh yl, Pt.)2), i Lrocwes, pl. Tt. 15))) 1843+) Eilsbry,, Man, 
PP 4O5,) Dia OO 135,130,189) 

Up men sts \COntAC, WatiAe) ING os) bila: ape Zon pl 2 tua. 
Morn(asiiivockus)) Wally Bulliz7 Us Say Me ior 16260: 


Distribution.—Hatteras to Honduras. Also West Florida, 
shallow water. 


(Eutrochus) jujubinum var. rawsoni Dall, Bull. M. C. Z., 18, p. 
369, °89; Dall and Simpson, Bull. U. S. Fish Com., 1, p. 
444, 1gol. 

Distribution.—Cedar Keys, Fla. and the Antilles. 


(Eutrochus) yucateeanum Dall, Bull. M.C.Z., 9, p. 47, ’81; 18, 
DapszOy Pl 24ef.140 gas So. 


Distribution.—Yucatan Strait, 640 fms. Also off Carolina 
coast, 15-32 fms. 


(Dentisty!a) asperrimum Dall, Bull. Mus. Comp. Zool., 9, p. 40, 
78m, (as| Margarita): 18, p. 373, °89; Pilsbry, Man, 11, p. 
Misi o Om Oa tll 27) UL) Se Niy MAC May mO2 nis: 


Distribution.—Hatteras to Barbados. Also West Florida. 
100-177 fms. 


Genus SOLARIELLA S. Wood 


amabilis Jeffreys, British Conch. 3, p. 300; 5, pl. 61, f. 6, (as 
Trochus); Dall, Bull. M. C. Z., 18, p, 378, ’89; Pilsbry, Man., 
EDs Lan pl s7a he 52.) (Soy 
Distribution.—Yucatan Strait, 640 fms. Also Antillean and 


European (North Sea off Shetland Isls, 85-95 fms). 


scabriuscula Dall, Bull. M.C. Z., 9, p. 41, ’8c, (as Wargarita); 


HOMO) Pi 2u i. TO loads So.) Dilly a7 ny sn Nie Mie 5 1. 
HOMME Dl 21h Le Om TOds, | 8On 


Distribution.—Southern part of the Guli of Mexico, 539 
fms., bottom temperature 39° F. Also off Cuba. 


127 


BULLETIN 38 : I57 


aeglees Watson, Jour. Linn. Soc., 14, p. 704, ’79; Chall. Rept. 


Gastr., p. 81, pl. 5, f. 10, 785; (as Margarita); Dall, Bull. 
VE CZ Ok! AO. Ole Ss DE RYIOn OO cial Sbiayaa Via tle 1s 
Bieby Holy nGfoyw peg csiay 10O).! | SX0), 


Distribution.—Gulf of Mexico, 287-888 fms. Also off St. 
Vincent, W. I. 


aeglees var. clavata Watson, Jour. Linn. Soc. London., 14, p. 


705, Chall. Rept. Gastr., p. 82, pl. 5, f. 8, (as Margarita); 
Dali) Bulll Me CZ 85 pi280) 78omelsbry Mam. pike ps 
ZOOM DL OO} fs OSs OO oO! 


Distribution.—Yucatan Strait, 640 fms.; Culebra Isl., W. I. 
350 fms.; Pernambuco, Brazil, 675 fms. Dall and Pilsbry 
regard this as a variety of aeglees. 


lubrica Dall, Bull. M. C. Z., 9, p. 44, ’81, (as Wargarita); 18, 


D282. pl. 20h) O- 1789. bilsbrmyaiMiane atti. Qo4u ls isin ae 
250/20. 8O- Wa lly Bally, sac nlel sy INI IVs TO: 1 s@ Aly) if0 en aeeaates 
9, 9a, 89. 

Distribution.—Cedar Keys, West Florida, to St. Lucia, W. 
I. 116-805 fms. 


lubrica var. iridea Dall, Bull. M. C. Z., 18, p. 382, ’89; Pilsbry, 


Mani rn) pi22Q so. Dall mle 2 Tanase ON. Verano 
1889. 


Distribution.—Cape Florida, Gulf of Mexico, West Florida 
region. 193 fms. 


lissoconcha Dall, Bull. M. C. Z., 9, p. 41, ’81, (as Margarita); 


WOR Remo, Aeiats an tekel. “oye JEU, OL Sal INE Ao}, 641, fol 
ZINE.) 8.82, OO) 


Distribution.—Gulf of Mexico, Lat 28° N., Lon. 88° W., 
at 331 fms. 


Genus BASILISSA Watson 


alta Watson, Jour. Linn. Soc. London, 14, p. 597, 79; Rept. 


Chall) Gastrs) p.mooliplai7. i SieoMbilsb hy.) Mam ent ip: 


158 MOLLUSCA OF THE GULF OF MEXICO 128 


ATOMDInE SON ius OO sO allo Dalia Nisa Men Del TOA 
1880. 


Distribution.—-Gulf of Mexico, West of Cedar Keys; the 
Antilles and Brazil. 339-1019 fms. 


alta var.delicatulaDall, Bull. M. C. Z., 9, p. 48, 81, (as Seguen- 
Ea i8.py 384) plaz2, t./2.0 80; Pilsbrayay Mantis, \p. 421 
Desa tai 8O Dall, Ballin: Us SmiNtuMietep sn 64) ple 
Domi 24 Zann So: 
Distribution.—Gulf of Mexico to Tobago, W. I. 805 fms. 

superba Watson, Jour. Linn. Soc. London, 14, p. 598, ’79; Rept. 
ChalleiGartrp. tom pl: 7, 4h. 10,,/86s)etlsbrye Miami.) Ta |). ps 
AQ ple Bout: O77) 789; Dall, Bulls a7 Us Sunnie. pi 164s 
1889. 


Distribution.—Cape York, Australia, type locality. Also 
Gulf of Mexico west of Florida. 400-1400 fms. 
(Ancistrobasis) costulata Watson, Jour. Linn. Soc. London, 14, p. 
Goor Sou Rept Chall: Gastr. p:) 103). ple7eiia im) S011 Dalle 
Bull MC) Z., 18, p:' 384, pl. 23, f.4; 4as8o; Pilsbry,; Man, 
IDE AZO pla Bon a so oO: Dall Bale anWe i SeNi Mess 
HAL, Ole Denes vale abaya tsKos 
Distribution.—Georgia to Culebra, W. I. 


costulata var. depressa Dall,Bull. M. C. Zool. Harv. Coll., 18, p- 
3845180) bllsbiry,) Man. Tr p: 428. plu, Pian soon Wall: 
Bulle Ue SNe Mos 164.) pli 2ecit4n an oor 
Distribution.—Yucatan Strait, 640 fms. Also Antillean. 

Genus UMBONIUM Link 


bairdi Dall, Bull. Mus. Comp. Zool., Harv. Coll., p. 359, pl. 21, 
Paonia) So. eilsbrys |) Mane) “ni pela ST ple sO@d tans. (6. 
1889. 


Distribution.— Yucatan Strait, 640 fms. Also Florida reefs, 
200 fms. 


12G BULLETIN 38 I5@ 


Genus COCHLIOLEPIS Stimpson 


parasitica Stimpson, Proc. Bost. Soc. Nat. Hist., 6, p, 308, and 
he 53./ Dall, BullyiNiiG. (ZeamS)\ 91260.) (So Bulla aa). 
Sa aNG Me. p! 162) (Sorina Wem Schis 20 ans amon 


Distribution.—Charleston Harbor, S. Car. to Florida Keys 
and Tampa, West Florida. At Charleston this species lives 
under the scales of a large annelid. Pliocene, Caloosahatchie 
beds, Fla., rare. 


striata Stimpson, MS.; Dall, Bull. M. C. Z., 18, p. 360, 1889. 
Ballea7.0U. 6 S..,Ne MG) pinG2n) (8oruehr. Was ner mllmsts 
Sei 3) pL 470) ple te TON O2h acot moun. Matchelt 
SOC! 1p. 138) Keb. 192m: 

Distribution.—Beaufort, N. Car. (Jacot); Egmont Key, 
near Tampa, West Fla. (Colonel Jewett). Upper Miocene 
of North Carolina. 

Genus ADEORBIS S. V. Wood 

adamsi Fischer, Journ. de Conch., p. 173, pl. 1o, f. 11, 1875; 
Dal eer O.7S. NM VIMEO penser Sa meiisiiy alma Cotte 
Acad. Sci., 10, p. 104, 1899-1900. 

Distribution.—Cited by Dr. Dall from Cedar Keys, West 
Fla. 
Genus VITRINELLA C. B. Adams 

mooreana Vanatta, Proc. Acad. Nat. Sci. Phila., 55, p. 758, 
1903. 

Distribution.—Crooked Island, Calhoun Co., Fla. 

hemphilli Vanatta, Proc. Acad. Nat. Sci. Phila., p. 24, pl. 2, f. 
iy Zip, Meas 
Distribution. —Cedar Keys, Florida. 

interrupta C. B. Adams, Monograph of Vitrinella, p. 6, 1850; 


jot: de ,Conch:.6.\ps076,-DalluMbulla 7 MUGS: INGE Mer: 
166, 1599;) Bush) itr Couns wNcadaisciny (10), ppsjLoom Loz. 


*99-1900. 


e] 


160 MoLLUSCA OF THE GULF OF MEXICO 13 


Distribution.—Tampa, West Fla., in shallow water, to 
Jamaica. 


Genus CIRCULUS Jeffreys 


liratus Verrill, Tr. Conn. Acad. Sci., 5, p. 529, ’82, (Omalaxis); 
6, Pp. 452, °85, (Skexea); Bush, of. ctt., 6, p. 464, °85; Bull. 
VC eZ nz) Da 240. Oly MIT hal LO Oech GomnaniAcads 
SCH TOM Prats.) pl 22tui7alnT2 Tab YOO 1oOO! 

Adeorbis supranttidus var. orbignyt Dall, Bull. M. C. Z., 18, p. 
POMS Os (AH partis lr. Wel Te Siig pmai24Anog Johnson, 
WceyiPa) Bost: Soc. Ni), 7) pe rzommons) Not ov bze 722 
nischer, Journ de Conch. 6, pp.) 173) 286), 157. 


Distribution.—New England to Hatteras and West Florida. 
Lower Miocene, Chipola beds, West Fla. 


Note.—The true Czrculus orbignyi Fisher is West Indian and 
very minute, only 144 mm. in diameter,while “vatus is-2% 
mim. in diameter. 

trilix Bush, Rept. U. S. Fish Com., for 1883, p. 584, 1885, (as 
SCLC), lie ConmexAcad. | (Sei) 10% paaa7pli22 tT. 6. °1o, 
10a, 12, a-g, pl. 23, f. 10, 15, 1899-1900. 

madeonous supyanczaus Wall, Bull. Mii Cay Ze n8. p.278)0 89° 

BUS NW TSO. lanai Rema cra oan bala 
part). Not supranitidus Wood, Cat. 1842, Crag. Moll., p. 
LA 7 MPL LS tsa sa-De kos. 
Distribution.—Massachussetts to Hatteras and to Cedar 
Keys, West Fla. 15-25 fms. Pleistocene, Knapp’s No. 2 
well, Terrebonne Parish, La., at 1519-1632 feet. 


Genus LISSGSPIRA Bush 


proxima Tryon, Manual Conch., 10, p. 98, pl. 33, f. 4, 1888, 
(Gydostrema); Bush, Tr. Conn. Acad. Sei:, 10, p. 130, f. 4, 
P22.) 20 T899-1G00, 

rugulosum Verrill, Tr. Conn. Acad.,5, p. 533,’82,(Cyclostrema). 
Not of G. O. Sars, 1878. 


131 BULLETIN 38 16X 


ayine Nermill) ir: Conn Acad @nGuipy moo. pluisen 1. es. e4e (as 
Cyclostrema). Not of Jeffreys, 1883. 


trochoides Dall, Bull. M. C. Z., 18, p. 393, 1889; Dall, Bull. 37, 
U.S. N. M., p. 166, ’89, (as Cyclostrema). Not trochoides 
(Jeffreys) Sars which is a variety of pettersen7 Friele. 


Distribution.—Lat. 41° N., Lon. 65° W. to Lat. 35° N., 
Lon. 74° W. (Bush); West Florida (Dall). 


Genus CHORISTELLA Bush 


pompholyx Dall, Bull. M. C. Z., 18, p. 394, pl 28, f. 9, ’89, (as 
Gydlosizenca) Balla Wem Sui NeeVieap at On plat 2 Saute mon 
’805 Bush, ir. Conn: Acad!) Sci-. 10; pp.) 99; | 1401) 1890- 
1900. 
Distribution.—Blake Station 2, Gulf of Mexico, type locali- 
ty. Also Fernandina, Florida, to Cuba. 294-805 fms. 
Note.—This species is referred tentatively to Choristella as 
Miss Bush suggested that it might prove referable to this 
genus. ’ 


Genus EPICYN!A Moerch 


multicarinata Stimpson, MS., Dall, Bull. M. C. Z., 18, pp. 273, 
Z02,'/807) Bull27, WS) Nea pr 166)786.) (as, ze7z7ella)\. 
Bush, Tr. Conn. Acad., Io, p, 112, ’99-1900. 
Distribution.— Hatteras to West Florida. Pliocene, Caloosa- 
hatchie beds,” Fla. Upper Miocene, Duplin Co., N. Car. 
Perhaps identical with the Avrchitectonica gemma of Holmes 
listed by Hilgard from the Lake Borgne borings, La. 


Genus ETHALIA A. Adams 


The following species are of doubtful generic position. ‘They 
were excluded from /¢halia by Pilsbry (Man. Conch., 1, 
Pp. 457) when he restricted that genus and gave Ethalia 
guamensis Quoy and Gaimard, as the genotype. Dr. 
Dall has referred me to Miss Bush’s revision (Trans. Conn. 
Acad. Sci., 10, p. 116, 89-1900). Apparently she tentative- 
ly retains these three species in Hzhalia. 


162 MOLLUSCA OF THE GuLF oF MExIco 132 


suppressa Dall, Bull. Mus. Comp. Zool., 18, p. 362, ’89; Bull. 37, 
Wee NE Men pu) NOON So) 


Distribution.—Goodland Point, West Florida. Collected by 
Hemphill. 


reclusa Dall, Bull. Mus. Comp. Zool., 18, p. 361, pl. 28, f. 7, 8, 
"SOs IEEE Soe OLAS INU [Bae omnadeloy, “Skok 


Distribution.—Yucatan Strait, 640 fms. North Carolina 
coast, 12-65 fms. 


Salida Wall) Bull M.C.Z,, Harv. (Coll 18) plist. 3) 5, 780; 
MSU aoe, INE Me pr LOO) pl N 28 uikal ay GanoC) 


Distribution.—West Florida and Cuba, 310 fms. 


Genus TEINGSTOMA A. Adams 


cryptospira Verrill, Trans. Connecticut Acad. Sci. and Arts.,6, p. 
PAPE OAL an (as, Acozella)) Wall Bull. 277-9 W4y SayNE Niet. 160; 
RSOq EUS lor, Con) Acad: - Sci... 10) pyrrole EN 241 F8Oo- 
LooOs Dall Ir Wagner Insts Sei., 33) ps) 4145.92) 


Distribution.—Hatteras to Florida. Gulf coast at Crooked 
Island, Calhoun Co., Fla. 30-150 fms. Pleistocene of 
North Creek, Fla. Miss Bush regards this species as atrue 
Teinostoma. 


Genus NERITA (Linneeus) Lamarck 


pelorenta Linnzeus, Syst. Nat., 10, p. 778, 1758; Reeve, Conch. 
NCOs Oy ls Be way Sow dang Miyoyo  Wiewowe sO, jos. Bala noyll 
Ai 75-7 7cooo Mall Bull=37.) Us Sa Nt Me Koos 1£8So: 
Dall andy Simpsons Bull U2 (Si hishe) Commie. 445) 
IQOL. 
Distribution.—Florida, St. Vincent and Texas region of the 
Gulf of Mexico. 


tessellata Gmelin, Syst. Nat., p. 3685, 1792; Reeve, Conch, 
icon oy pl. ON. 7435) 494,) 6855-) dinyon wane moO py 24, ipl: 


133 BULLETIN 38 163 


Aer! Seen Dali ull eal SiN INO Goneon Dalle 
and Simpson, Bull. U. S. Fish Com., 1, p. 446, 1901, 


Distribution.—East Florida to Colon. Gulf coast, West 
Florida and Texas, shallow water. 


versicolor Lamarck, Anim. s. Vert., 6, pt. 2, p. 193, 1822; Reeve, 
Conch: Icons) o. ole 12) fase. a dakOs Somali Etiam. 
Uiisu NN. Mes) p: 166.) 780: Dall andi Simpson yBulliue S: 
ish) Com: (i) p..446.) 190r 


Distribution.—Florida Keys and West Florida to Colon, the 
Antilles and Bermuda. 


Genus NERITiNA Lamarck 


reclivata Say, Jour. Acad. Nat. Sci. Phila., 2, p. 257, 1822, (as 
Npeodoxus)\: Reeve. Conchiainon ey 34) a-b)). 55> Wallpean 
WSN SUMS. pul2sonpl iim tase nS se iyon Mane srounne 
39, pl. 12, f. 25-27, ’88; Singley, Fourth Ann. Rept. Texas 
Geol. Surv., p. 342, 1892; Mitchell, List. Texas Shells, 
Pp. 7- 
Distribution.—St. Augustine to Martinique. Gulf coast: 
Mouth of Manatee River, Tampa, Cedar Keys and St.Mark’s, 
Fla., Point au Fer and Chandeleurs, La.; Port Lavaca and 
Rockport, Tex. Pleistocene, Knapp’s No. 3 well, Terre- 
bonne Parish, La. from 1-700 feet. 


Note.—The variety palmae Dall is found at Palma Sola, 
Texas. 


virginea Linnzeus, Syst. Nat.. ed. X, p. 778, 1758, (as Nerita); 
Reeve) Conch icon o.ny plea 22a Ge ot S55 uaa 
OAS Ne Mays pil 25oM/8i5:) myonme\Vlan..) TO.) Dp: 2Qunplagia. 
Hagt-25. 37-42 Dall Maulana 7a eos ON NIG.) py OSes SO. 
Singley, Fourth Ann, Rept. Geol. Surv. Texas, p. 342, 
1892. 


Distribution.—Florida to Brazil. Gulf coast: Various local- 
ities of West Florida; Corpus Christi and Rockport, Tex. 


x64 MoLLuscA OF THE GULF OF MEXICO 134 


pupa Linnzeus, Syst. Nat., ed. X, p. 378, 1758; Tryon, Man., Io, 
Os ABS Ole WAL A ore ate ODN, JEENULS Gi OWS: SS) INi2 ME yo), 
168, ’89. 
Distribution.—Charlotte Harbor, West Florida to Jamaica. 
Shallow water. 


viridis Linnzeus, Syst. Nat., ed. X, p. 778, 1758; Tryon, Man., 
MOMS ts plu 18) £88) 1888; Dall) Bully Wu Su Ni Vi. 
Dylos. 89. 
Distribution.—Florida Keys to Barbados, W. 1. Also Texas 
region of the Gulf of Mexico. 


showalteri Lea, Pr. Acad. NatSci. Phila., p. 55, 1861; Tryon, 
(BOS TKOs FOI Get ONAN Aben Snes iacKs en IOBUN IS ybIUG) ceioie Ola Sip 
ING Me ps) K68;) .6O: 


Distribution.—Alabama and West Florida, Fluviatile. 
sparsilineata Dall, Proc, U.S. N. M., 46, p. 236, 1914. 


WNenintan spsiindet,, Aldrich, Nantilis) 24))/pi nin ps 131, 
IN@s. Jo UGUIE BO folie seme MayM toner), 


Distribution.—Upper Miocene or Pliocene. Well near Al- 
exandria, La., at 49 feet; Pine Prairie La., well at 1540 ft. 
and well % mile S. E. of Pine Praire depot, in dump. Also 
Satilla River, Ga., and Burkeville, Texas. One of the most 
widespread and characteristic species of the brackish water 
formation, extending from Georgia through Louisiana to 
Texas. 


Genus PLEUROTOMARIA Sowerby 


(Petrotrochus) quoyana Fischer and Bernardi, Journ. de Conch., 
Eee HOS. Dla Sut. 1856: Dall Balin Man Z eons 7.o) 
gon Crosse journ., de Conch), 22." pa ian s2snMall Bull: 
NICE ZEaers) De O7h pli 2ou ta Te Wu tMn Tal Me CH le nae an 
Poon mullet 27 UL SuiNG Me ps 168) samenpls wand) fosi 
1889. 


Distribution.— Yucatan, near Arrowsmith Bank, at 130 fms. 


135 BULLETIN 38 265 


Also off Barbados, 73 fms., and Marie-Galante Island, near 
Guadeloupe. 


Genus FISSURELLA Bruguiere 


(Cremides) barbadensis Gmelin, Syst. Nat., p. 3729, 1792, (as 
iRacewa)i Dall Bull 27. Ui Sy iNWuM wpe 7O.1.8o) nbd story. 
Man., 12, p. 164, pl. 37, f. 40-49, ’90; Dall and Simpson, 
Ball U2 5) Fish) Come (ie sp) Aa su oom 

Genus LUCAPINA Gray 


suffusa Reeve, Conch, Icon., in errata, 1851, (as Fissurelia); Dall, 
icrans,) Wiaener Insti Set 2) py Azdpleizan ie (22)402k 


hondurasensis Reeve, Conch. Icon., f. 70, 1851. 


cancellata Sowerby, in part, Thesaurus Conch., p. 200, pl. 8, 
(248) f187 only. 362.) Wall Mabini Wan S a4 oN MEN iMG Os 
189); bilsbiy, Manin ta. De 2OOMrDlao static pla Mstemsor 
’90; Dall and Simpson, Bull. U.S. Fish Com., 1, p. 449, 
Igol. 


Distribution.—Marco, West Florida to Vera Cruz, Mexico. 
Also Antilles and South America to the Island of Fernando 
Noronha, Brazil. 


Note.—As there has been some question whether the name 
suffusa Reeve or cancellata Sowerby should be applied to 
this species, Dr. Dall’s opinion was asked. He replied, June, 
1921, ‘‘I find nothing to change inthe synonymy of ZL. 
suffusa as given in the Wagner memoir.’’ 


adspersa Philippi, Abbild. u. Beschr., 2, p. 34, pl. 1, f. 3, 1845; 
(as) -Fissuvella) Rilsbiny, iMate fe 2h apie el OO). ls O2 eno) wie7s 
’°90; Dall and Simpson, Bull. U. S. Fish Com., 1, p. 449, 
IgOl. 


fasciata Pfeiffer. Fissurellidea fasciata (Pfeiffer) Dall, Bull. 
VRS SME p72 iSO), 


Distribution.—West Florida region of the Gulf of Mexico to 
Cubavandist + Crom Vie a 


166 MOLLUSCA OF THE GULF OF MEXICO 136 


Genus FISSURIDEA Swainson 


alternata Say, Jour. Acad. Nat. Sci. Phila., 2. p. 281, 1822, (as 
iassuneiia) Pilsbry,) lane. 12) ps 211) pleg7.\f50-53, pl: On, 
Penis) oo7) Dall and | Simpson, Bully Ues:) kish, Com,” 1; 
Pp. 450, Igor. 


Distribution.—Chesapeake Bay through the Antilles to 
Trinidad Isl. and Fernando Noronha, Brazil. Gulf coast: 
Cedar Keys, Fla.; Galveston, Corpus Christi, Texas; Vera 
Cruz and Progreso, Mexico. Also south of Nicaragua. I-50 
fms. Pliocene of South Carolina. 


eayenensis Lamarck, Anim. s. Vert., 6, pt. 2, p. 12, 1822; Reeve, 
Conch olcon? 7h) 232) Dall, Bull} a7 Uaioe NEM | p12 70) 
Ow bilsbry.) Man re) ps 212, plzz 1. above 60, 1890. 


Distribution.—Cedar Keys, Fla. to St. Lucia, W. I. Placed 
by Pilsbry in synonymy of a/fernata, but is more elongated 
and with straighter sides than that species. Perhaps to be 
classed as a variety. 


Genus PUNCTURELLA Lowe 


iriolumMe Wall Bulle MC) Z.\ 9, p.176, Si; 18ip-w4og, pls 26, f. 
8, 8b, ’89; Bull. 37, U. S. M. N., p. 168, pl. 26, f. 8, 8b, 
1889. 


Distribution. —Yucatan Strait, 640 fms. 


watsoni Dall, Bull. M. C. Z., 18, p. 403, ’89; Bull. 37, U. S. 
ING Me p68.) 780: 


Distribution.—Off Yucatan, at 200 fms. Also off Cuba and 
Barbados. 


cincularis;) Dall) Bull, Mi: (@. Z:> 9, pi75; *8n;/ 18, p. 403, pl. 26, 
Pee SO.) Bulls 7a Mey Sa Nee Mesmp ea stOSnapl 26), fai. 
7b, ’89. 


Distribution.—Florida Strait and the West Indies. 


1127) BULLETIN 38 167 


(Fissurisepta) triangulata Dall, Bull. M. C. Z., 18, p. 404, ’89; 
Bolle) S2 ING ais ipscw7onn So: 


rostrata Watson, Chall.Gastr., p. 48, pl, 4, f. 10, 7&5. Not of 
Seguenza. 


Distribution.—Yucatan coast, 200 fms. Also off Fernan- 
dina, Fla., and Culebra Isl., W. I. 


(Cranopsis) asturiana Fischer, Journ. de Conch., 30, p, 51, 1882, 
(as Rimula); Watson, Chall. Gastr., p. 45, pl. 4, f. 4, ’85; 
Dall Bulls IM Cai Zein S) (psi One Son 


Distribution.—Yucatan Strait, 640 fms. Also off Havana, 
Martinique, and Cape Florida, and in the Gulf of Gascony 
at 2018 fms. 


Genu EMARGINULA Lamarck 
(Rimula) frenulata Dall, Bull. M. C. Z., 18, p. 406, pl. 28, f. 4, 
OOM BU e 7 OES INE NED ene Ou 2 Shake Aloe 
Distribution.—West Florida, Florida Keys and Hatteras, 6- 


52 fms. 


(Emarginula) compressa Cantraine, Bull. Acad. Roy. Bruxelles, 
9, p. 2. 1835; Jeffreys, Pr. Zool. Soc., p. 679, ’83;Dall, Bull. 
Me CuOZ TS pv AOAt esos 


Distribution.—Yucatan Strait, 640 fms. Also off Havana, 
Barbados, and coast of Portugal. ‘Tertiary of Sicily. 


x68 MOLLUSCA OF THE GULF OF MEXICO 138 


GNSS) NINA ed ool NI Le RR 
ORDER POLYPLACOPHORA 


Genus LEPIDOPLEURUS Risso 


pergranatus Dall, Bull. M.C. Z., 18, p. 414, ’89, (as Leptochi- 
ton): Bull. 37, U:'S. N. M.,. p. 172, 89; Dall and Simpson, 
Balle US. hish) Come: 19.451, ply yssitavi aac. Loon: 


Distribution.—Gulf of Mexico, western Florida region, 
Porto Rico and Dominica, 114-1181 fms. 


Genus CHAETGPLEURA Shuttleworth 


apiculata Say, Amer. Conch., app., pt. 7, 1830, (as Chzton); 
Dallyaaly 27s se NEM per 72. aienih. TO. T8So% 
Pilsbryg) Mian’)! 14, p. 35, pli’ 13, tf) 75-7on 1892;) Johnson, 
Occmeanibost.. soc. Ni HH: (7, 1p. 8) Toms: 
Distribution.—Massachussetts to Haiti, Gulf coast, various 
localities of northwestern Florida and Chandeleurs, La. 0-30 
fms. 


Genus ISCHNOCHITON Gray 


limaciformis Sowerby, Proc. Zool. Soc., p. 26, ’32, (as Chiton); 
DalkeBully Mey CayZe 18. p. 45.) So svedlsbrypyMans ogy p: 
57, pl. 16, f. 9-16, ’92; Dall and Simpson, Bull. U. S. Fish 
Com p4 4.52.) LOOT 
Distribution.—Gulf of Mexico, southeastern part, Lat. 24° 
N., Lon. 83° W., at 37 fms. Also Key West and the An- 
tilles, Central America and Peru. Under stones at low 
water. 


purpurascens C. B. Adams, Proc. Bost. Soc. Nat. Hist., p. 9, 
u845. (as) Chzzon))) Dall) Bull..27. Wo Se Ne MeL) pi 172.) So: 
Edlsbiny. Manian pa SSa plait. 2oN 2annnSQ 2 ial arid 
Simpson, Bull. U.S. Fish Com., 1, p. 452, 1got. 


Distribution.—Texas region of the Gulf of Mexico, Florida 
Keys, Jamaica and Porto Rico. 


139 BULLETIN 38 ; 16g 


papillosus C. B. Adams, Proc. Bost. Soc. Nat. Hist., p. 9, 1845; 
Dall yBall 27.00 SeiNeMMelnm. 7172) 7 Sonia bia eNleian yaeAy 
Pp. 114) pl. 21-8, 40) 401892. Dall andi simpson, Bulli Weise 
Fish) Com),/1, ip: 452, 190: 


Distribution.—Tampa, West Florida, to St. Thomas, 
Wevele 


Genus CHITON Linnzeus 
fuberculatus Linnzeus, Syst. Nat., ed. X, p. 667, 1758; Pilsbry, 
Man., 14, p. 153, pl. 33, f. 58-60, 1892; Dall and Simpson, 
Bull. U.S. Fish Com., 1, p. 453, Igor. 


Distribution.—Texas and Florida to Trinidad and Ber- 
muda. 


marmoratus Gmelin, Syst. Nat., p. 3205, 1792; Dall, Bull. 37, 
WOE Se ING Mae) ips 2h So nba sbiave Mamie y 4c 0.1 15S, plana 
f. 72-76, 1892; Dall and Simpson, Bull. U. S. Fish Com., 1, 
Pp. 454, 190t. 
Distribution.—Texas, and the Antilles to Barbados. South 
to Cartagena, Colombia. 


Genus ACANTHOPLEURA Guilding 


granulata Gmelin, Syst. Nat., p, 3205, 1792, (as Chzton); Pilsbry, 
Man., 14, p. 227, pl. 50, 1892; Dall and Simpson, Bull. U. 
S. Fish Com., I, p. 454, 1901. 


pica (Gmeling Dall) YBull 27 sOm Sa ONG Ne. Pe) lat isos 


Distribution.—Charlotte Harbor, West Fla., to Trinidad, 
Bermuda and Bahamas. 


Genus ACANTHOCHITES Risso 


spiculosus Reeve, Conch. Icon., pl. 9, f. 47, 1847; Dall, Bull. 37, 
We SING ME pn 740) OO Med Sbiny NAT NS. | Dah2 nolan 
f. 60-62, 1893; Dall and Simpson, Bull. U.S. Fish Com., 


Ae ON USES Wa Keyos (3 


170 MOLLUSCA OF THE GULF OF MEXICO 140 


Distribution.—Cedar Keys, West Florida, to Barbados. 
pygmaeus Pilsbry, Tryon’s Manual Conchology, 15, p. 23, pl 13, 

PF OK 159s SOs. 

Distribution.—Cedar Keys and Key West, Fla. 


I4I BULLETIN 38 E71 


GHASS CE RHA @R@ WA 


ORDER DIBRANCHIATA 


Genus ARGONAUTA Linnceus 


argo var. americana Dall, Bull. 37, U. S. N. M., pp, £74, 200, 
plas) t. 1 a-by pleat Tazbiplv on Mit 2188.) [ounsen, 
@Occei}Pa\ Bost.’ Soc: Nat (Eista7hipai2235)LOls: 


Distribution.— Massachussetts north to 43° N. Lat., and 
south to the Antilles and doubtfully to Brazil. Gulf of 
Mexico, West Florida region. 


Note.—The typical Argonauta argo Linnzeus, Syst. Nat., 
ed. X, p. 708, 1758, is Mediterannean. It has two or three 
times as many radial folds and carinal nodules as the varie- 
ty americana. 


Genus SPIRULA Lamarck 


australis Lamarck, Anim.s. Vert. 7, p. 600, 1822; Encycl. Me’th., 
pliA6s5, tS, a,b: Walls Science) mi sins, spp. 243-2454 nego. 
Dall/and Simpson, (Bull WS; bush Come) 1p.) 359, ilies, 
LAs OOT: 


spirula Linneeus, in part. 


perone (lamarck) Dall, Bul a7 UES INY MCn loin nels 
CSA TSO: 


Distribution.—Pelagic. West Indies and the Gulf of Mex- 
ico, shells being washed ashore along West Florida coast 
and at Cameron, La., Galveston and Corpus Christi, Texas. 
A specimen with all the soft parts was also taken from the 


172 


MOLLUSCA OF THE GULF OF MExICO 142 


mouth of a fish trawled by the Albatross between the Mis- 
sissippi delta and Cedar Keys, at 324 fms. The Spzvula had 
just been seized alive by the fish and the soft parts, pre- 
viously imperfectly known, were complete. See Science, 
13896. 


ivnere are! several (tors) of |) Spezza | Siiispzrula. cot 
Linnzeus was a composite, according to Dr. Dall, and since 
the species cannot be differentiated by the shells alone no 
one can tell exactly which species Linneeus had although he 
gave America as the habitat. Huxley and Pelseneer (Chal- 
lenger Rept.) fixed the name australis Lamarck on the An- 
tillean species. 


Note.—Arvgonauta and Spirula are the only shell-bearing 
Cephalopods reported from the Gulf of Mexico. Of other 
forms, the Brazilian, Loligo brevis Blainville is cited from 
Cameron and elsewhere on the Louisiana coast, and Z. 
hemtptera Wowell from Florida and the Gulf of Mexico. 


Ma ee og Seer 
“A Bis te % 


me 
be 


ss 
pera 


BULLETINS 
OF 


AMERICAN PALEONTOLOGY 


Vol. 9 


The Miocene of Northern Costa Rica 


WitTH Novtss On Irs GENERAL STRATIGRAPHIC RELATIONS 
BY 


A. A. OLSSON 


Harris Co. 
Cornell University, Ithaca, N. Y. 
UA Savane 


OEDICAT =D 
WOSF A= 


MUSEU IM! Cm GOSsra RICA 


y 
hg 


ay 


me =n V1 OS En INE @) a) IN @) eal el ES ESN 


G@QSWA RiGA 


WITH NOTES ON ITS GENERAL STRATIGRAPHIC 


ION. 


JW 


RELATIONS 


BY 


A. A. OLSSON 


PRELIMINARY REMARKS 


GENERAL STRATIGRAPHY OF NORTHERN COSTA RICA 


THE CosTA RICAN MIOCENE 
A. THE USCARI STAGE 
B. THE GATUN STAGE 
C. THE USCARI - GATUN UNCONFORMITY 


CORRELATION 
A. SANTO DOMINGO MIOCENE 
B. BOWDEN BEDS, JAMAICA 
C. MEXICAN MIOCENE 
D. THE East Coast MIOCENE 


Tue Costa RicA GEOSYNCLINE 


is i, 
Bi 


aii 
t ¥ 


179 Costa R1cA Mr1ocENE—OLSSON 7 


PRELIMINARY REMARKS 


The fossil collections and field information, on which this 
monograph is based, resulted from over two years of field work 
by the writer in Panama and Costa Rica. The writer’s thanks 
are due to the Costa Rica Oil Corporation for permission to pub- 
lish; and he is most grateful for assistance, and encouragement 
received in the course of this work from Drs. A. C. Veatch and 
D. F. MacDonald of the Sinclair Exploration Company, to the 
Officials of the National Museum of Costa Rica and Professor 
J. Fidel Tristan of San Jose’. 


The collections were studied in the Paleontogic Laboratory 
of Cornell University, the facilities of which, Professor G. D. 
Harris, most generously placed at my disposal. 


Cornell University, 
January, 1922. A. A. OLsSson 


181 Costa Rica MrocENE—OLSSON 9 


GENERAL STRATIGRAPHY OF NORTHERN CosTA RICA 


The northern or Atlantic slope of Costa Rica, the so-called 
‘Tierra Caliente’’ is largely composed of Tertiary sedimentary 
rocks, which except for some folding, dip away from the slopes 
of the Cordillera Central towards the sea. ‘To the east, this 
sedimentary belt may be traced into the Province of Bocas del 
Toro of western Panama, where its upper and highly fossilifer- 
ous beds are well developed and exposed on the islands of the 
Chiriqui Lagoon. ‘This Tertiary sedimentary belt ends some 35 
to 40 miles east of the city of Bocas del Toro. ‘The Miocene 
sedimentary rocks of Batun and of the northern Canal Zone 
form a small independent unit, which begins about 30 miles 
west of Colon and ends close to the eastern limits of the Canal 
Zone. 


In a general way, the stratigraphic succession in northern 
Costa Rica is as follows: 


Pleistocene Slightly elevated coastalswamps, with deposits of cross-bedded 
sands, clay-marls ete. 


Pliocene Certain reef-limestones composed of coral and hydrozoa re- 
mains; also marls, blue lignitiferous clays, blue coarse sand- 
stones, and a predominating hard conglomerate, composed of 
large boulders of igneous rocks (the so-called boulder-clays of 
several writers on Costa Rican Geology). 


Miocene The Gatun Stage or Formation. Coral-reef limestones, (near 
the coast), marls, more generally a blue or blue-black sand- 
stone, dark colored shales, sandy and impure limestones, ligni- 
tes and conglomerates. Generally very fossiliferous. 


The Uscari Stage or Formation. Usually dark-colored shales 
but with some sandstones, limestones and conglomerates. The 
smaller types of foraminfera are usually quite abundant, and 
are associated with fish-scales, and small! echinoid spines, with 
mollusks generally rare. 


Oligocene Sandstones, shales and a complex of hard, often semicrystal- 
line limestones which pass through all possible gradations to 
coarse arkosic sandstones. The most abundant and characteris- 
tic fossils are the Orbitoidal foraminifera (Lepidocyclina). The 


IO BULLETIN 39 182 


age of these beds probably ranges from Lower to Upper 
Oligocene. 


THE CostTA RICAN MIOCENE 


An important part of the stratigraphic succession of north- 
ern Costa Rica is composed of rocks of Miocene age, whose 
thickness varies according to locality and completeness of sec- 
tion, from a few hundred to several thousands of feet. They are 
divisible into two stages or formations by an unconformity or 
disconformity of greater or less degree. In their natural order 
of sequence, these divisions of the Costa Rican Miocene are: 


Gatun Stage or Formation: Middle and Upper Miocene 
Uscari Stage or Formation: Lower Miocene 


THE USCARI STAGE 


The Uscari stage or formation derives its name from Uscari 
creek in the Talamanca valley of eastern Costa Rica. It con- 
sists principally of soft, dark-colored shales and because of their 
slight resistance to denundation, their outcrop frequently forms 
wide valleys and interior basins. These shales were deposited in 
quiet and moderately deep water, and their fauna is largely 
composed of the smaller types of foraminifera, such as Globzg- 
evina, Textularia, Lagena, Nodosaria and Miliola, but the 
large Orbitoidal Lepzdocycline, abundant in the underlying Oli- 
gocene, are absent. Mollusks are very uncommon in this forma- 
tion. Thesmall fauna described by Gabb from Sapote, Costa 
Rica, belongs to the lower part of the Uscari formation. 


THE GATUN STAGE 


The upper division of the Costa Rican Miocene is equivalent 
in part to the Gatun formation of the Canal Zone. The typical 
Gatun, so named from its well-known exposures at Gatun and in 
which a part of the Panama Canal and the locks at Gatun were 
excavated, lies in an independent sedimentary basin across the 


1323 Costa Rico MrocENE—OLSSON II 


northern end of the Canal Zone and in the adjacent parts of 
the Province of Colon. There the Gatun formation consists 
of blue sandstones, dark-colored shales, fuller’s earth beds etc., 
generally less than 500 feet thick. Many of its beds are very 
fossiliferous. ‘The mollusks of the Gatun stage of Panama have 
been studied by Dall, Toula, Brown, Pilsbry and Cossmann. 


In Costa Rica, the Gatun is very much thicker than in 
the Canal Zone and represents a longer depositional period. 
Paleontologically, the Gatun of the Canal Zone, seems to repre- 
sent only the lower part of the formation as developed in Costa 
Rica. It is largely of sandy character, more resistant to denu- 
dation than the Uscari shales and gives rise to a more rugged 
and hilly topography. Although typically sandy, it also contains 
locally shales, lignites, conglomerates and coralline limestones. 
In some localities the limy members are of major importance. 
The formation is characterized by marked lateral variation in 
which respect it differs. strongly from the much more uniform 
shales of the Uscari stage. ‘This feature of lateral variation 
gives rise to two important phases: that of the more typical 
Gatun, in which the beds are largely sandy in character, with 
some lignitic and coarsely conglomeritic members, and carrying 
the usual near shore Gatun fauna; and the coralline phase, in 
which the beds are more marine, with coralline limestone and 
richly fossiliferous marls perdominating, and with no lignitic 
beds. 


Along the inner margin of the Miocene belt, the Gatun be- 
longs to the first and more typical phase; while the coralline 
type is more common along the coastal zone, as at Port Limon 
and Bocas del Toro. Coralline rocks may occur at any horizon 
in the Gatun succession or it may locally be composed of mas- 
sive coral-reef limestones. Such is the case in the Limon Penin- 
sula, and such coralline limestones, by most observers, have been 
regarded as of Pliocene or even of more recent age. At Limon, 


£2 BULLETIN 39 784 


this confusion has partly arisen from the fact that these Mio- 
cene limestones are overlain by the second series, of Pliocene 
age, which partly yielded the large fauna described by Gabb, 
published shortly after his death in 1881. Dall later contribut- 
ed to the discussion of this fauna. The truly Miocene limestones 
of Limon may beseen in good exposures along the sea-front 
west of the city. ‘There they are massive in character, hard and 
recrystallized to the extent that all traces of organic remains 
have usually been obliterated or rendered uncertain of determin- 
ation. However, some sandy or marly beds are occasionally 
found intercalated in these coralline limestones, and from these 
layers we have collected a large fauna of unquestionable Mio- 
cene affinities, correlating closely in age with the Bowden beds 
of Jamaica. 


The fauna of the Costa Rican Gatun is largely molluscan. 
The following paleontological study records 334 species. Future 
collecting will doubtless greatly add to this list, as several species 
common it the Canal Zone, still remains undiscovered in Costa 
Rica. ‘The smaller types of foraminifera abound in the more 
shaly phases of the formation, as well as several species of echin- 
oids, belonging to the genera Clypeaster and Encope. Corals of 
simple and compound types occur principally in the coralline 
phase. 


THE USCARI - GATUN UNCONFORMITY 


At the close of Uscari time the Miocene sea was with- 
draw from the greater part of Costa Rica, but during the 
Gatun, it returned. These conditions are indicated by a 
stratigraphic break between the two stages, a complete change in 
their lithologic composition, the occurrence of Gatun beds in 
areas which had received no deposition during Uscari times and 
a change in their respective faunas. Although the full strati- 
graphic details of this important subject cannot be presented at 
the present time, attention is called to it, as the outstanding fea- 
ture of the Miocene stratigraphy of Panama and Costa Rica; and 
because of its bearing on correlation. 


185 Costa Rica MIocEN# —OLSSON 13 


In many parts of northern Costa Rica, the Gatun stage 
carries at its base a thick and persistent conglomerate, formed of 
coarse cobbles of igneous rocks, or of finer pebbles with alternat- 
ing layers of sandstone. This conglomerate generally rests dis- 
conformably on the Uscari, but in some cases is unconformable 
on these or older rocks. ‘This basal conglomerate is generally 
firmly cemented and because of its superior hardness, may be- 
come an important ridge former. It marks the site of high 
water-falls along mauy streams. Fossils of common Gatun 
types are abundant at many localitiesin this basal conglome- 
rate. 

Gatun rocks are frequently found in areas which contain no 
Uscari beds. In such cases, the Gatun is found resting, with 
strong unconformity, on older rocks. This condition is seen in 
the Canal Zone and along the borders of the sedimentary belt of 
the western Panama. It indicates, either a complete erosion of 
the Uscari shales during the Uscari-Gatun interval or, what is 
more likely, a greater transgression of the Gatun sea. This 
uncomformable relation of the Gatun upon older sedimentary 
beds may be seen in the Chagres Spillway in the Canal Zone, 
where the underlying rocks are sandstones, probably equivalent 
to the Mt. Lirio of Lower Oligocene age. Inthe Chiriqui La- 
goon country, of western Panama, the base of the Gatun is 
frequently a coral-reef limestone, which rests directly upon the 
older and more disturbed rocks. 


% 
> 


al sk 


The present systematic study enumerates 334 species of 
molluska of which the majority belong to the Gatun stage. A 
few of these species have not been found in Costa Rica or west- 
ern Panama, but are common in the Canal Zone, and are includ- 
ed here for the sake of comparison with Costa Rican species. 
To these we may add the 18 species described by Gabb from 
Sapote, Costa Rica, but not included in our collections and the 
following species listed below from the Canal Zone. This brings 


14 BULLETIN 39 186 


the total known Miocene fauna of Panama and Costa Rica to 
396 species. 

Volvulella micratracta Brown and Pilsbry 

Ringicula hypograpta Brown and Pilsbry 


Conus haytensis Sowerby Corbula gatunensis Toula 
Conus domingensis Sowerby Corbula sphenis Dall 
Cruns consobrinus Sowerby Corbula sericea Dall 


Conus emulator Brown and Pilsbry 

Drillia vaningent Brown and Pilsbry 

Drillia gatunensis 'Toula Solecurtus strigillatus Linné 
Drillia fusinus Brown and Pilsbry 

Drillia Zooki Brown and Pilsbry 

Drillia enneacyma Brown and Pilsbry 
Cythara heptagona Gagg 

Marginella gatunensis Brown and Pilsbry 
Anachis fugax Brown and Pilsbry 

Murex polynematicus Brown and Pilsbry 
Murex gatunsts Brown and Pilsbry 

Typhis Gabbi Brown and Pilsbry 

Malea camura Guppy 

Cyprea Henekint Sowerby, var. 

Littium nugatorium Brown and Pilsbry 
Natica bolus Brown and Pilsbry 

Natica canalizonalis Brown and Pilsbry 
Sigaretus gatunensis 'Toula 

Sigaretus (Eunaticina) Gabbé Brown and Pilsbry 
Cheilea princetonia Brown and Pilsbry 
Nucula (Acila) isthmica Brown and Pilsbry 
Pecten effosus Brown and Pilsbry 

Pecten operculariformis ‘Toula 

Pecten reliquus Brown and Pilsbry 
Amusium Toule Prown and Pilsbry 

Ostrea gatunensis Brown and Pilsbry 
Cardium durum Brown and Pilsbry 
Cardium gatunensis Dall 

Tellina equiterminata Brown and Pilsbry 


187 Costa RicA MrockNE—OLSSON 5 


Chione tegulum Brown and Pilsbry 
Pitaria cora Brown and Pilsbry 
Dosinia delicatissima Brown and Pilsbry 
Petricola millestriata Brown and Pilsbry 
Cyclinella gatunensis Dall 

Thracia gatunensis Toula 


The Uscari fauna is still poorly known, but it includes sev- 
eral very distinctive species of which the following are most 
important: 

Amusinm Lyonit Gabb 

Arca dariensis Brown and Pilsbry also Lower Gatun 
Clementia dariena Conrad also Lower Gatun 
Canccllaria (Aphera) islacolonis Maury 

Neverita neretdis Maury 

Sconsia cocleana Olsson 

Scaphella costaricana Olsson 

Ptychosalpinx ? dentalts Olsson 

Dentalium uscarianum Olsson 

The Gatun is the great fossil bearing formation of northern 
Costa Rica and Panama, and is characterlzed by a rich and high- 
ly diversified fauna. This fauna at the present numbers nearly 
400 species, but additional collecting will doubtless add consid- 
erably to this number. As we have already noted the Gatun 
beds present two important facies, the sandy and highly clastic 
phase of the typical Gatun, and the deeper water type, illustrat- 
ed by the coralline limestones and their associated marls. These 
lithological types have their own more or less peculiar and dis- 
tinctive faunas. 

The typical Gatun is the more common phase and is charac- 
terized by a molluscan fauna of heavy, thick-shelled species of 
littoral forms. In Costa Rica, the following are the most com- 
mon and characteristic: 

Arca MacDonaldi Dail Marginella MacDonaldi Dall 
Arca Pittiert Dall Voluta Alfarot Dall 
Pecten levicostatus 'Toula Strombus pugiloidis Guppy 


16 BULLETIN 39 188 


Cardium stirtatum Brown, Pilsbry Stormbus gatunensis Toula 


Chione Rowelli Olsson Natica Guppyana ‘Toula 
Chione mactropsis Conrad Turritella altiliva Courad and its 
Dosinia acetabulum Conrad varieties 


Conus molis Brown and Pilsbry 

The coralline phase is developed at several localities along 
the coast, most important of which are Port Limon and Bocas 
del Toro. Here the coralline limestones and associated marls 
form the greater part of the Gatun section, but coralline rocks 
may occur, interbedded with typical Gatun rocks, at any strati- 
graphical level. 


These coralline limestones were formed, mainly as barrier 
reefs some little distance from the old Gatun shore-line. Natur- 
ally they offered very different habitat conditions from those of 
the typical near-shore and often lacustrine Gatun. Hence we 
find a fauna of deeper-water affinities with Pleurotomids and 
thin-shelled Cones, as the most important and characteristic 
element. In localities where these coralline rocks are exclusive- 
ly developed, this fauna bears very little relation to the typical 
Gatun fauna, although the two are contemporaneous. The 
following are species of the coralline phase: 

Pecten coralliphila Olsson Mitra Alamagrensts vat. 
Antigona Flarristana Olsson coralliphila Olsson 
Gafrarium limonensis Olsson Mitra poas Olsson 

Large number of Pleurotomids J/arginella avena Valennencies 


Conus planilivatus Sowerby Olivella limonensis Olsson 
Conus limonensis Olsson Phos elegans, variety limonensts 
Conus stenostomus Sowerby Olsson 
Turritella exoleta Linné Stliquaria modesta Dall, 

var. limonensis Olsson limonensis var. Olsson 


About 13 per cent. of the Gatun fauna is identical or closely 
related to the recent species, the majority of which are found 
living along the present Caribbean coast, certain others only on 
the Pacific side in the Panama Province and a few common to 
both. ‘This large percentage of recent forms corresponds closely 


189 Costa Rica MioceNE—OLSSON 17 


with that characterizing the Chesapeake Miocene fauna of the 
eastern United States and shows their close agreement in age. 
The recent species and their closely related forms in the Gatun 
fauna are as follows: 


Arca occidentalis Phil. West Indian 

Arca umbonata Lamarck West Indian 

Arca candida Gmelin West Indian 

Arca auriculata Lamarck West Indlan 

Ostrea megodon Hanley Pacific 

Anomia simplex d’Orb. West Indian 

Phacotdes vadians Conrad var. medioantericanus 
Olsson West Indian 

Phacoides trisulcatus Conrad West Indian 

Glycymeris castaneus Lamarck West Indian 

Glycymerts decussatus Linné West Indian 

Divaricella guadrisulcata d’Orb. West Indian 

Cardium medium Linné West Indian 


Cardium spinosum var. Turtont Dall East Atlantic 
Cardium serratum Linné West Indian 
Pitaria circinata Born and its var. alternata Broderip 
West Indian and Pacific 
Antigona multicostata Sowerby Pacific 
Antigona rugosa Gmelin West Indian and Pacific 
Macrocallista maculata Linné West Indian 
Chione grus Holmes West Indian 
Tellina crystallina Chem. West Indian and Pacific 
Strigilla pistformis Linné West Indian 
Mactra exoleta Gray Pacific 
Mactra alata Spengler West Indian 
Labiosa lineata Say West Indian 
Corbula Dietziana C. B. Adams West Indian 
Tellidora cristata Recluz West Indian 
Conus proteus Hwass West Indian 
Conus floridanus Gabb var. costaricensis Olsson West Indian 
Terebra benthalis Dall var. bocasensts Olsson West Indian 


18 BULLETIN 39 190 


Turris albida Perry West Indian 
Glyphostoma dentifera Gabb West Indian 
Acteocina bullata Kiener var. costaricana Olsson West Indian 
Acteocina recta d’Orb. West Indian 
Volvula cylindrica Gabb West Indian 
Anctstrosyrinx elegans Dall variety West Indian 
Marginella avena Valen West Indian 
Murex messorius Sowerby West Indian 
Bursa crassa Dillwyn West Indian 
Natica canrena inne West Indian 
Architectonica granulata Lamarck West Indian nnd Pacific 
Astralium brevispinum Lamarck, variety dasalis 
Olsson West Indian 
Astralium caclatum Gmelin West Indian 
Sastrum nodulosum C. B. Adams West Indian 
Latirus tufundibulum Gmelin West Indian 
Northia northie variety mtocenica Olsson Pacific 
Crepidula plana Say West Indian 
Xenophora conchyliophora Born West Indian 
Oliva testicea Lamarck var. costaricensts Olsson 


It is but natural to expect that the Miocene beds along the 
northern or Caribbean coast of Central America, should contain 
a fauna largely ancestral to that of the present Caribbean, but 
along with these strictly Caribbean types, we also find several 
species, identical or closely related to forms now resticted to the 
Pacific side. ‘Their occurrence in Caribbean deposits points to a 
connection between the Atlantic and Pacific oceans during Mio- 
cend time and allows for a certain intermingling of their faunas. 
Geological observations indicate that there were at least two 
such straits crossing Central America during the Miocene, the 
main one across Costa Rica, and a second through the Darien of 
eastern Panama. 


This Pacific element of the Gatun fauna comprises many of 
the most common and characteristic species. Certain of these 
species had a wide distribution in the West Indies during Mio- 


QI Costa Rico Mi1ocENE—OLSSON 19 


cene time and have been noted by several writers. They 
comprise the following species from Panama and Costa Rica, 
listed together with the Pacific analogues: 


Panama or Costa Rica Pacific Analogue 


PAGO RAL L OVO SOM CTD paakeiscies stisesaiceire Arca grandis Sowerby 
Arca (Noetia) MacDonaldi Dali.....drca reversa Sowerby 
Ostrea megodon Hanley 

Mactira exoleta Gray 


Mactra estrellana O\SSON..........1.000+ + Mactra (Harvella) elegans Sowerby 
Semele laevis Sowerby var. 
costaricensts Olsson........semele laevis Sowerby 


Macoma panamensis Dall variety 
canalis Olsson...J7acoma panamensis Dall 


Macoma gatunensis Toula............... Macoma elongata Hanley 
Tellina crystallina Chemnitz............ also West Indian 
Antigona multicostata Sowerby 

Antigona rugosa Gmielin................. also West Indian 


Pitaria circinata Born variety 
alternata Broderip 
Cyclinella subqguadrata Hanley variety 


guitana Olfson...... Cyclinelia subguadrata Hanley 
Cyclinella beteyensis O\SSON.........2+- Cyclinella Kroyeri Philippi 
Conus vecognitus GUPPy...--.-..+2e0eeee Conus pyriformis Reeve 
Turricula lavinoides Olsson............ Turricula lavina Dall 
Cancellaria islacolonis Maury.......... Cancellaria tesselata Sowerby 
Cancellaria toroensis Olsson..........+. Concellaria tuberculosa Sowerby 
Cancellaria Pluimmert Olsson.......... Cancellaria bullata Sowerby 
Mitra Swainsont Brod. variety 
limonensis Olsson...... Mitra Swainsont Broderip 
UNIEVEMILONMETELATS, MIAULY, (21.004 .5> 23.060 Neverita glauca Humbolt 
Northia northiae Gray variety 
miocenica Olsson.....- Northia narthiae Gray 
INAIECONCOMVUTA (GUPPYs..5.22-64-de006e++-0- Malea ringens Swainson 
Oliva testacea Lam. variety 
costaricensis Olsson...... Oliva testacea Tam 


With the closing of the Miocene straits, this Pacific element 
gradually had to give way before the development of the West 
Indian fauna, but it is still to be seen in such species as, 7e/lina 
crystallina Chem. Architectonica granulata Lamarck, which 
have a distribution along both coasts. Moreover the Pliocene 


20 BULLETIN 39 192 


and Pleistocene beds of Central America contain a well-marked 


Pacific element in such species as Worthia northie Gray Oliva 
testacea Lamarck and Pecien ventricosus Sowerby. 


CORRELATION 


Miocene rocks are now known from a great many localities 
in the West Indies and the general Caribbean area of Central and 
South America. In these areas they comprise a very important 
series of sedimentary formations. Beds equivalent to the Gatun, 
appear to be the most widespread and generally are highly fossil- 
iferous. 

The following chart showing the equivalence of the Miocene 
horizons in the West Indies and elsewhere to those of Central 
America, does not differ materially from the more recent correl- 
ations proposed by Maury, Cooke or Vaughan, except in the ad- 
dition of the Uscari to the Lower Miocene of Panama and Costa 
Rica. The Gatun formation is here recognized as being of Mid- 
dle Miocene age with possibly some of its upper beds having 
Upper Miocene affinities. The Gatun, at its type locality in the 
Central Zone, is not more than 500 to 600 feet thick. In Costa 
Rica it is very much thicker, and its higher beds may range into 
the Upper Miocene. It is the exact equivalent of the Chesapeake 
Miocene of the eastern United States. In this correlation, I have 
been guided not only by the paleontologic evidence, but also by 
the field relations of the formations, studied over a wide rage of 
country in Costa Rica, and in Panama as far east as the Colombia 
frontier. . 

SANTO DOMINGO MIOCENE 

In Santo Domingo and in Haiti, Miocene rocks are exposed 
in several areas, but attain their best development along the val- 
ley of the Rio Yayue del Norte in the northern part of the is- 
land. ‘These beds are highly fossiliferous and because of the fine 
perservation of their fossils, have repeatedly attracted the atten- 
tion of paleontologists. 

The earlier writers dealing with the paleontology of the Do- 
minican Miocene, believed thatit represented but a single 


X93 Costa RIcA MIocENE—OLSSON 21 


stratigraphic unit, and Gabb insisted that no division was possi- 
ble. Dr. Maury however, as a result of paleontologic studies 
from collections carefully made in the field, was able to divide 
the Dominican Miocene into two formations, which she called 
the Gurabo and the Cercado. ‘To these Vaughan and his as- 
sociates, in more recent stratigraphic work on the island, have 
added 4 other subdivisions. The present recognized subdivisions 
of the Dominican Miocene are: 


Mao clay 
MIDDLE Mao Adentro limestone 
Gurabo formation 

(Sconsza laevigata zone) 


Cercado formation 
(Aphera islacolonis zone) 
Baitoa formation and Bulla 
conglomerate 


MIOCENE 


LOWER 


The relation of the Costa Rican and Panama Miocene to 
that of Santo Domingo is extremely close; there being about 
go species in common. Of this number about 21 are recent, 
about 26 species appear to be restricted to the Gurabo, some 27 
species are common to both the Gurabo and Cercado formations, 
and only 11 species are confined to the Cercado. The list of 
of Costa Rican and Panama species in the Dominican Miocene, 
is as follows: 

Nucula tenutsculpta Gabb Cercado and Gurabo 


*Arvca occidentalis Phil Cercado and Gurabo 
*Aryca umbonata Lamarck Cercado 

Arca Maurye Olsson Gurabo 

Arca FHlenekent Maury Cercado 


Arca goltoyaguensis Maury Cercado and Gurabo 
Arca chiriquiensits Gabb Gurabo 
*Arca auriculata Lamarck Gurabo 


% Recent 


i) 
N 


BULLETIN 39 194 


Glycymerts jamatcensts Dall Cercado 
Lime@a solida Dall Gurabo 
*Ostrea megodon Hanley Gurabo 
Spondylus gumanemocon Brown and Pilsbry, Gurabo 
*Anomia simplex d’Orb Gurabo 
Placunanomia lithobleta Dall Gurabo 
Pteria inornata Gabb Cercado and Gurabo 
*Cardium serratum Linné Cercado and Gurabo 
Cardium dominicensis Gabb 
*Cardium medium Linné Gurabo 
Protocardia gurabica Maury Cercado 
*Wactra alata Spengler Cercado 
Antigona tarquina Dall? Cercado 
Antigona Blandiana Guppy Gurabo 
*Pitarta circinata Born Cercado 
* Tellina crystallina Chemnitz Cercado 
Tellina cercadica Maury Cercado 
*Strigilla pistformis Linné Cercado 
Semele Claytont Maury Cercado 
Venericardia scabricostata Guppy Cercado and Gurabo 
Echinochama yagquensis Maury Gurabo 
Corbula vieta Guppy Cercado and Gurabo 
Corbula viminea Guppy Cercado 
** Acteocina recta d’Orb Cercado 
Acteocina triticum-tritonts Maury Cercado and Gurabo 
* Volvula cylindrica Gabb Cercado 
Bullaria paupercula Sowerby Cercado and Gurabo 
Ringicula dominicana Maury Cercado 
Terebra spirtfera Dall Cercado 
Terebra bipartita Sowerby Gurabo 
Tercbra haitensts Dall Gurabo 
Terebra Berlinere Maury Gurabo 
Conus haytensis Sowerby Cercado and Gurabo 
>? ~~ Sewalli Maury Gurabo 


“Recent 


CostA RicA MrocENE—OLSSON 


Conus stenostomus Sowerby Gurabo 

Conus proteus HAwass Cercado and Gurabo 
Conus recognitus Guppy Cercado and Gurabo 
Conus multiliratus Bose Gurabo 

Conus gracilissimus Guppy Cercado and Gurabo 
Conus marginatus Sowerby Gurado 

Conus consobrinus Sowerby Cercado and Gurabo 
Conus domingensis Sowerby Gurabo 

* Turris albida Perry Cercado and Gurabo 
Drillia venusta Sowerby Gurabo 

Drillia consors Gurabo 

Cythara heptagona Gabb Gurabo 

*Glyphostoma dentifera Gabb Gurabo 

Cancellaria Maurye Olsson Cercado and Gurabo 
Cancellaria Rowelli Dall 


Cancellaria epistomifera Guppy Cercado and Gurabo 


Cancellaria tslacolonis Maury Cercado 

Oliva brevispira Gabb Cercado and Gurabo 
Oliva cylindrica Sowerby Cercado and Gurabo 
Olivella muticoides Gabb Cercado and Gurabo 
Mitra rudis Gabb Cercado and Gurabo 

Mitra longa Gabb  Cercado and Gurabo 
*Latirus infundibulum Gmelin Gurabo 
Melongena consors Sowerby Cercado and Gurabo 
Alectrion losquemadica Maury Gurabo 
Metula cancellata Gabb Gurabo 

Murex cornurectus Guppy Cercado and Gurabo 
*Vurex messorius Sowerby Gurabo 

Typhis alatus Sowerby Gurabo 

Distortio simillima Sowerby Gurabo 

*Bursa crassa Dillwyn Gurabo 

Cassis sulcifera Sowerby Cercado and Gurabo 
Sconsia levigata Sowerby Gurabo 


*Recent 


i) 


Ww 


24 BULLETIN 39 196 


Dolium camura Guppy Gurabo 
Dolium elliptica Pilsbry and Johnson Gurabo 
Cypraca Henekent Sowerby Cercado and Gurabo 
Cypraca dominicensts Gabb Gurabo 
Strombus pugiloides Guppy Gurabo 
Serpulorbis papulosa Guppy Cercado and Gurabo 


Petaloconchus sculpturatus H.C.lea Cercado and Gurabo 
*Architectonica granulata Lamarck Cercado 


*Xenophora conchyliophora Born Gurabo 
*“Natica canrvena Linné Cercado and Gurabo 
Natica Youngi Maury Cercado and Gurabo 
Polinices subclausa Sowerby Cercado and Gurabo 
Polinices Stanislas-Meuntert Maury Gurabo 
Neritina viridemaris Maury Coreado and Gurabo 
LNeverita neretdis Maury Cercado 


Most of the above listed species belong to the Gatun forma- 
tion. Cancellaria (Aphera) islacolonis Maury, Neverita neretdis 
Maury and Corbula viminea Guppy occur in the Upper Uscari 
and in the Lower Gatun or transitional beds of eastern Costa 
Rica. This leaves but the following 8 Cercado species in the 
Gatun fauna, or a proportion of about 3.25 to 1. 

Arca Henekent Maury Winter Cay 
Glycymeris jamaicensts Dall Winter Cay 
Protocardia gurabica Maury Water Cay and Gatun 


Antigona tarquinia Dall? Saury 
Tellina cercadica Maury Saury 
Semele Claytont Maury Hotel Creek 
Corbula viminea Guppy Middle Creek 


Ringicula dominicana Maury Middle Creek, Red Cliff 
Creek and Estrella River 


*Recent. 


197 Costa RicA MioceEnm—OLSSON 25 
GENERAL CORRELATION CHART NO. I. 
Costa Rica and | Canal Santo Jama Porto 
western Panama Zone Domingo tea | Rico 
SSI SN Ese Set SC (Se Se itera SEL ee TA SRNL cI USEA EINE INT Ie Dick) TOSI ne 

| 

v . 

5 ms Limon. Monkey Las Matas 

aan Point, Bocas Stage 

m= 

Be Conglomerates 

(boulder clays) 

SRB BR MRE RB aT anes lhe tl Seah meted llinshl tome) \fecriph ea | fost. suse y Nike | % 

ed h | | 

a | Cerros de | | 

BY 2? Toro | Sal Stage hoe 

P Gatun Stage Stage Ui @ucbras 
\Bowden| dillas 

Stage act 

A Mao Clay 

4A Gatun 

jaa 5 Stage |Mao Adeotro 

o &. Gatun Stage | limestone ° 

on Gurabo Stage 

> 

s! ! 

FA € ie iS * % Ae Bees Vlies Sn bce c < < oe me | S asl) Tas v 

fa Cercado Stage | 
=  -Uscari Stage | 
e) (Sapote) | 
aN | Baiton Stage 
Reisen Reise 
Seles | ena Seed oe Ni aap fet <r Te ro KITS ach ha ‘ | 

o) | | | 

5 ‘ Emperado l’s | | 

5 Undifieren- | Culebra Stage | Cevicos l’s | 

ae tiated | Bohiocongl.|  Tabera Stage | 

© 


26 BULLETIN 39 198 


GENERAL CORRELATION CHART NO. il. 


Colombia and Mexico Eastern 
Venezuela United Staies 
* x a * * * me * * ¥ Dr x %* % * * * * * * + x% + 
Caloosahatchie 
| Waccamaw 
| 
o st x x x ¥ xX s ft * TAO TAR eM Ph Ge) Te hes oe aS 


Txuetepec, State} & Yorktown and Duplin Stage 
3 of Oaxaca a & 
is ‘<> Murfreesboro Stage 
5 Santa Rosa and | 
Cartagena and |& Barranca de v 
9 
Cumana a Santa | 
5 2, 
\2 Maria Tatetla, |s'. St. Mary’s Stage 
he] Nw 
© State of Vera aie 
° oq Cruz a) é Choptank Stage 
| 
| Calvert Stage 
eae ac Viyisen sen Wese UAC SORA R NE NESE na Cai rat igiay a ean | atl bea Niet eg |S aa) 
i a) 
Peano aay @ Shoal River marls 
ae eet om eeyan| Oak Grove marls 
Bre Hoe Bay Chipola marls 
Soa 
\<q 
x x aa w ea ileal eri wah lee CY cogil aay 2 [Py EA apy es Veo te iiMlezas Weel ea ea Se 


Tampa Stage 
Vickburgs Stage 


199 ~ Costa Rico MrocENE—OLSSON 27 


The Arca Henekent, Glycymerts jamaicensis and Protocardia 
gurabica occur with a typical Gatun fauna at Water Cay, west- 
ern Panama, associated with Sconsza, and other Gurabo shells. 
With the exception of Ringicula dominicana,the other species are 
generally rare. ‘The evidence is therfore strongly in favor of 
the equivalence of the Gatun beds with the Gurabo of Santo 
Domingo. 


BOWDEN BEDS, JAMAICA 


Next to Santo Domingo, the highly fossiliferous beds at 
Bowden, Jamaica, are the best known. From this locality, Dall 
has listed about 435 species, of which 12 per cent. appear to be 
identical with recent species. 

Recent correlation by Cooke places the Bowden beds as 
equivalent to the Gurabo formation of Santo Domingo,and there- 
fore to the Gatun formation; a correlation which is borne out by 
our own studies. About 38 Gatun species, not including recent 
forms, are here considered equivalent to Bowden shells. In gen- 
eral, the Costa Rican localities which show the closest agree- 
ment with Bowden, are the upper coralline horizons, so that the 
Bowden beds may be a little higher than the main Gurabo and 
they are so considered here. The following Bowden shells oc- 
cur in the Costa Rican Gatun. 


*Acteocina bullata Kiener 


*Acteocina recta d’Orb Petaloconchus sculpturatus Wea 
*Volvula cylindrica Gabb Alaba turrita Guppy 
Lullaria paupercula Sowerby *Crepiduia plana Say 
Terebra bipartita Sowerby * Nrtica near canrena Linne 
Conus planiliratus Sowerby Glycymeris jamaicensis Dall 
Conus stenostomus Sowerby *Arca occidentalis Philippi 
Conus granozonatus Guppy *Arca auriculata Wamarck 
Conus recognitus Guppy *Ostera megodon Hanley 
Conus interstinctus Guppy Lime@a solida Dall 

Conus gracilissimus Guppy Placunanomia lithobleta Dall 
*Turris albida Perry Echinochama antiguata Dall 


*Recent 


28 BULLETIN 39 200 
Drillia consors Sowerby Venericardia scabricostata Guppy 
Drillia venusta Sowerby Myrtea limoniana Dall 
Cancellaria Barretii Guppy Phacoides actinus Dall 

Oliva cylindrica Sowerby *Levicardium serratum Vinne 
*Latirus infundibulum Gmelin Cytherea Blandiana Guppy 
Melongena consors Sowerby Tellina lepidota Dall 

Metula cancellata Gabb *Strigilla pistformis TLinne 
Typhis alatus Sowerby Abra triangulata Dall 

Distortrix simillima Sowerby Corbula viminea Guppy 

“Bursa crassa Dillwyn Gastrochena rotunda Dali 

Cassis sulcifera Sowerby Phalium moniliferum Guppy 
Sconsia sublevigata Guppy Malea camura Guppy 

Strombus pugiloides Guppy Serpulorbis papulosa Guppy 


MEXICAN MIOCENE 


From Mexico, Boseft has described several small faunas 
from the Isthmus of Tehuantepec, and from the States of 
Oaxaca and Vera Cruz. These faunas, he considered as Upper 
Miocene and Pliocene. 


The most interesting of these localities, in the present con- 
nection, are those at Tuxtepec in the State of Oaxaca, and 
Santa Rosa and the Barranca de Santa Maria Tatetla in the State 
of Vera Cruz. From these localities, we may note together 
with their Costa Rican and other affinities, the following 
species: 


Fecten santarosanus Bose Fecien levicostatus Youla Gatun 

Amustum Mortont Ravenel Amusium luna Brown and Pilsbry 

Levicardium sublineatum Conrad Gatun Chesapeake Miocene 

Anomia simplex d’Orb Gatun-Chesapeake to recent 

Venus Ebergenvit Bose Atone ulocyma Dall Upper Chesapeake 

Solarium Villarelloi Bose Architectonica granulata Lamarck 

Lower Miocene or recent 

Vermetus pulcher Bose Serpulorbis sculpturatus H.C. Lea 
Gatun and Upper Chesapeake Miocene 

Strombus pugilis Linne Strombus pugtloidis Guppy Gatun 

Sconsia sublevigata Guppy Bowden 

Phos mexicanus Bose Gatun of Water Cay 

Melongena Mengeana Dali Solenosteira Vaughani Dall variety 


medioamericana Olsson,Gatun of 
Water Cay 


* Recent. 
+ Boletin del Instituto Geologico de Mexico, No. 22, 1919, pp. 11 - 88 


204 Costa RicA Mriockn® —OLSSON 20 


fleurotoma alesidota Dall var. 


MAGMA BOSC......000-. Drillia consors Sowerby, Gatun 
Conus Agassizi Dall variety 
multilivatus BOSe........... Conus multiliratus Bose, Gatun 
COMUSMEUYGRRATLD BOSC aN soe seces. Gatun 


The above species are from Tuxtepec and Santa Rosa. To 
these we may add; from Barranca de Santa Maria Tatetla, the 
following forms: 


Ostrea sculpturata Conrad.............. Ostrea costaricensis Olsson, Gatun 
Lucina quadrisulcata Vinné........... Divaricella quadrisulcata d’Orb 
Gatun, Upper Chesapeake Miocene to 
recent 
Levicardium serratum Vinné........ Gatun, Upper Chesapeake to recent 
Dosinia acetabulum Conrad..........-. Gatun - Chesapeake Miocene 


The above list is but a partial one of the entire fauna from 
these localities. However,’ it shows the close relationship be- 
tween these Mexican species and those of the Gatun and the 
Bowden, such as Sconsia sublevigata, Phos mexicaius, Conus 
multilivatus and others, witha few typical Chesapeake species 
such as Chione ulocyma Dall, Dosinta acetabulum Conrad and 
Cardium sublineatum Conrad. ‘This illustrates the interming- 
ling of the Gatun-Bowden-Gurabo fauna with that of the Chesa- 
peake, and proves their equivalence in age. 


THE East Coast MIOCENE 


A fine development of Miocene rocks is found along the 
Atlantic coastal plain and they extend more or less continuously 
from New Jersey south into Florida. They are continued around 
the borders of the Gulf of Mexico into Texas, but in these re- 
gions, their marine phases are deeply covered by younger forma- 
tions, and they are known only from deep wells. These Mio- 
cene beds are divisible into two series, which early were classifi- 
ed as (a) the older or sub-tropical Miocene and (b) the younger 
and cold-water Miocene. These terms are descriptive of their 
main faunal differences. The upper division consisting of sever- 


30 BULLETIN 39 202 


al formations or stages is frequently known as the Chesapeake 
Miocene or Chesapeake series, and the lower consisting solely of 
the Alum Bluff formation, may be designated for sake of discus- 
sion, the Alum Bluff series. These Miocene formations or stages 
are the following: 


| Yorktown, Duplin Stages 


Upper Murfreesboro Stage 
ml, * * * * | Chesapeake Series/* * * * * * * * 
a | |St. Mary’s Stage 
ea Middle | Choptank Stage 
Oi Calvert Stage 
oO | | 
4 | Mie a Meira aa egos.) Re | BEC RCT RA aaa a ies |e 
= | Alum Shoal River marl 
| _ Bluff 
| |Alum Bluff Series} Stage Oak Grove marl 
| Lower Chipola marl 


| 


This two-fold division is a very natural one. Each series 
represents a distinct trangression of the Miocene sea over parts of 
the Atlantic coastal plain, that of the Chesapeake being the 
more extensive. In Florida, where the Chesapeake Miocene is 
sometimes found resting upon the Alum Blufi beds, as at Alum 
and Jackson Bluffs, the contact is seen to be an erosional uncon- 
formity. At these localities, the Chesapeake Miocene is equiva- 
lent to the Murfreesboro stage or formation of Virginia and 
North Carolina, and hence the time interval represented by this 
break is equivalent to the Lower Chesapeake or the Marylandian 
Miocene. Dall,* who has paid special attention to the relations 
of the Alum Bluff and the Chesapeake series, both stratigraphi- 
cally, and paleontologically, characterizes this break as ‘‘the 
most sudden, emphatic, distinct in the whole post-Cretaceous his- 


* Trans, Wagner Free Inst., 1903, p. 1594 


203 Costa Rica Mi1ocENE—OLSSON 31 


tory of our southeastern Tertiary, and indicates physical changes 
in the surrounding region, if not in Florida itself, sufficient to 
alter the course of ocean currents and wholly change the temper- 
ature of the waters of our southern coasts.”’ 


The fauna of the Lower Miocene or Aluin Bluff series is of 
warm-water character and largely composed of Antillean types. 
In the Chesapeake series, the fauna is of cold or temperate-wat- 
er character and it finds its relations with the present Atlantic 
fauna north of Cape Hatteras. These faunal differences must be 
take into account in any correlation between the Miocene of the 
United States with that of the West Indies and Central 
America. 


In the Correlation Charts,I have placed the Gatun formation 
as equivalent to the Chesapeake series, the Uscari formation 
with the Alum Bluff. The evidence upon which this correla- 
tion is based, may be summarized as follows: 


1. The two-fold division of the Costa Rican and Panama 
Miocene, by an unconformity at the base of the Gatun forma- 
tion. 


2. Wherever the Gatun beds have been found overlain by 
more recent formations, the age of these overlying beds is Plio- 
cene or younger. 


3. The high percentage of recent species in the Gatun 
fauna, which agrees with the average as shown for the Chea- 
peake Miocene. 


4. The Gatun fauna, contains a few species which appear 
identical with strictly Chesapeake species, such as the follow- 
ing: 

 Dosonia acetabulum Conrad 

Petaloconchus sculpturatus WH. C. Lea 

Chama congregata Conrad 

Plicatula marginata Say? 


32 BULLETIN 39 404, 


The Dostnia acetabulum-Conrad is one of the most abundant 
and characteristic species of the Chesapeake Miocene and in its 
course through the Chesapeake formations, several varieties are 
developed, of great use in correlation. The typical acetabulum 
is however confined to the Upper Chesapeake formations and it 
was figured by Conradin his Medial Tertiary, from a fine large 
example from Smithfield, Virginia. It is characterized by its 
large size, broad outlines and even, concentric banding. This 
Upper Chesapeake or typical acetabulum corresponds very close- 
ly in size, form and sculpture to the Costa Rican examples. It 
is very common at certain localities in Costa Rica, especially in 
the lower part of the Gatun formation. Bose records it from 
Mexico. The VPetaloconchus sculpturatus is a widely distributed 
species in the West Indies and Caribbean Miocene, from which 
it is more generally recorded as P. dominigensts, a name given to 
the Santo Domingian shell by Sowerby. Close comparative 
study of a large collection of typical seulpturatus from Virginia 
and North Carolina, failed to show any real difference, either in 
details or sculpture or in nature of coiling. The Petaloconchus 
sculpturatus occurs in the Bowden beds of Jamaica; the Gurabo 
formation of Santo Domingo; the Quebradillas limestone of 
Porto Rico; the Springvale beds of Trinidad; and the Gatun of 
Panama and Costa Rica. It is also known from Cumana, Vene- 
zuela: Cartagena, Colombia; and Santa Rosa, Mexico. In the 
Chesapeake Miocene, it is restricted to the Murfreesboro, York- 
town and Duplin formations, that is, to the Upper Chesapeake. 
It does not occur in the overlying Pliocene so far as is known. 


5. Possibly more significant because of their greater num- 
ber are the following species, which appear for the first time in 
the Atlantic coast Miocene in the Chesapeake formations. They 
also continue into the Pliocene and generally into the recent 
fauna. 


205 Costa Rica MrockENE—OLSSON aa 


Anomia simplex d’Orb St. Mary’s to recent 
Cardium spinosum var. Tur- 
tont Dall Murfreesboro to recent 


Cardium serratum Linné © Murfreesboro to recent 
Cardium medium Linné St. Mary’s to recent 
Labiosa lineata Say Duplin to recent 
Tellina umbra Dall Duplin to recent 

Chione grus Holmes Duplin to recent 
Phacotdes radians Conrad Duplin to recent 


Phacotdes trisulcatus Conrad Chesapeake to recent 
Divaricella quadrisiulcata d’Orb Murfreesboro to recent 
Oliva sayana var.immortua 
Pilsbry and Brown typical shell, upper 
Chesapeake to recent 
Fasctolaria tulipa Linné Pliocene 


Tur Costa RICAN GEOSYNCLINE 


The wide distribution of Miocene beds around the borders 
of the Gulf of Mexico and Caribbean sea, indicates a condition 
of general deposition during Miocene time. Costa Rica was 
largely under water, at least during the Lower Miocene, and 
was the locus of a strait or geosyncline that then connected the 
Atlantic and Pacific Oceans; while Panama, parts of Nicaragua, 
and other central American areas were mainly above sea level. 
This condition is shown in the general igneous character of the 
backbone of Panama, while in Costa Rica, sedimentary rocks of 
Miocene age occur even in the high interior and at elevations of 
several thousands of feet. This geosyncline covered the greater 
part of Costa Rica, a portion of western Panama, and western 
Nicaragua. It should be noted that this area, at the present 
time, is the locus of a large number of active volcanoes. 


Certain changes of importance took place at the close of 
Uscari time, resulting in a brief withdrawal of the Miocene sea. 
The effects of these changes were widespread. These great land 
movements, which probably reached a culmination in the Plio- 


34 BULLETIN 39 206 


cene, had far reaching consequences and to them we must ascribe 

even the sudden and marked change between the fauna of the 

Alum Bluff and Chesapeake series of the eastern United States. 

It probably resulted in a partial closing of the Atlantic-Pacific 

straits and a disarrangement of the direction of oceanic currents. 

The strongly clastic character of the Gatun formation as compar- 

ed with the more shaly Uscari, is also a result of the elevation of _ 
the land. 

The Pliocene was a period of high elevation and, in Costa 
Rica, of intense volcanic activity. The coarse conglomerates, 
or the so-called boulder clays, which have perplexed most geo- 
logists who have visited Costa Rica, were formed from the 
erosion of these Pliocene lava flows and deposited as flood-plain 
material. In a few localities where these conglomerates reached 
the sea, they contain intercalated shale beds carrying marine 
fossils of late Pliocene age. 


MU 7 1909 


MONE Sax 


GWASS| GAS RROPODA 
ORDER OPISTHOBRANCHIATA 


SUBORDER TECTIBRANCHIATA 


Genus ACTAEON, Montfort 


Actzon costaricensis, n. sp. 


Shell moderately large, ovate-cylindric; body-whorl large 
and more than 3 of the total length of the shell; spire pointed, 
conic; whorls about 6, convex and with deep sutures; spiral 
sculpture of fine, nearly regular, smooth threads separated by 
deep and somewhat wider interspaces; there are about 9 spiral 
threads on the spire-whorls and about 34 on the last whorl; the 
interspaces are crossed by fine, elevated, distant threads; aper- 
ture linear-lanceolate, rounded below; columella with a single 
large fold. 

Height 19.5 mm, diameter Io mm, aperture 14. 5 mm. 

This isan unusually large and cylindrical species with a 
relatively high spire. Suggestion of the original coloration is 
indicated by a white band which encircles the middle of the 
last whorl commencing at the upper end of the aperture,— the 
general ground-color of the surface being a faint purple or red- 
dish brown. ‘The interspaces between the spirals are crossed by 
fine, elevated threads, but they appear merely punctate on the 
early whorls. 

Gatun Stage: Hill No. 3, Banana River. 

Middle Creek. 


Genus ACTEOCINA, Gray 


Acteocina bullata Kiener, var. costaricana, n. var. Plate 15 figures 20, 21 
cf. Tornatina bullata Kiener, Icon. Coq. Viv., p. 5, pl. 1, fig. 4. 


206a a 34a 


ct 


34b BULLETIN 39 206) 


Shell rather large, moderately solid; outlines subcylindrical 
but a little wider about the anterior half of the shell; the spire 
is short, composed of 4 or more whorls, and a small nucleus of 
the usual type, asin canaliculata,; sutures channelled; the sur- 
face appears smooth, until slightly magnified when it is seen to 
be covered with very fine spiral lines; columella carries a single 
strong fold. ; 

Height 10, diameter 3, aperture 9 mm. 

The Costa Rican fossils are related to the recent A. bullata 
Kiener, of the West Indies but differ in their shorter spire and 
are somewhat more expanded about their anterior half. It is the 
largest species in the Costa Rican Miocene, often reaching a 
length of 1o mm. Its surface is covered with fine, spiral lines 
and the columella carries a single, strong plication. 

Gatun Stage: Middle Creek. 

Coll. 4, East Grape Point Creek. 


Acteocina recta d’Orbigny 


' Bulla vecta d’Orbigny, 1845, Delas Sagra, Hist. Pol. y Nat. Isla de 
Cuba, vol. 5, p. 67, no. 55; Atlas 8, pl. 4, dzs, figs. 16-20. 
Tornatina recta Gabb, Trans. Phil. Soc., vol. 15, p. 246. 
Tornatina coix-lacryma Guppy, 1876, Quart. Journ. Geol. Soc. Lon- 
don, vol. 22, p. 518. Probably not of Guppy, 1867. 
Acteocina recta Maury, 1917, Bull. Amer. Pal., vol. 5, p. 178, pl. 3, 


sete ey 

A small species, typically with a short exserted spire and a 
broadly cylindrical form. The Costa Rican examples are all 
less than 3mm in height. There is considerable variation in 
the height of the spire, which may be sunken in some cases as 
illustrated by the larger cozx-lacryma Guppy from the Miocene 
of Venezuela and Martinique, to others with high spires and 
strongly descending whorls as in the recent canaliculata Say. 
However they all appear to belong to the same species. From 
the young of bullata var. costaricana, they are at once distinguish- 
ed by their smooth, plain surface. 


Gatun Stage: Coll. 7, Estrella River. 
Coll. 5, Red Cliff Creek. 


206c Costa Rica M1ockENE—OLSSON 34c 


Subgenus CYLICHNELLA, Gabb 
Acteocina triticum-tritonis Maury 
Cylichnella bidentata Gabb, 1873, (in part) Trans, Amer. Phil. Soc., 
vol. 15, p. 273. Not of d’Orbigny, 1845. 
Cylichna bidentata Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 
32, p. 518. 
Acteocina (Cylichnella) triticum-tritonis Maury, 1917, Bull. Amer. 
ale VOUS Py L7O. Pla siutie., Ae 
This shell was separated by Dr. Maury from the recent 67- 
dentata d’Orbigny, with which Gabb, Guppy and others had 
identified it. In ¢vzticum-tritonts the anterior of its plications is 
very small, the posterior one large, which is the reverse of the 
conditions as seen in the living d¢dentata. ‘The base of the shell 
is finely grooved with incised, spiral lines. ‘The specimens from 
Costa Rica agree very closely with typical specimens from the 
Miocene of Santo Domingo. 
Gatun Stage: Middle Creek. 


Genus VOLVULA, A. Adams 


Volvula cylindrica Gabb 

Volvula cylindrica Gabb, 1873, Trans. Amer. Phil. Soc.. vol. 15, p. 
246. 

ie cylindrica Guppy, 1876, Quart. Journ. Geol. Soc. London, 
vol. 32, p. 518. 

cf. Bulla (Volvula) cf. oxytata Toula, 1908, Jahrb. der K-K. Geol. 
Reichsanstalt, Wien, vol. 58. p. 709, pl, 28, fig. 4. 

Volvula cylindrica Maury, 1917, Bull. Amer. Pal., vol. 5, p. 180, pl. 3, 
fig. 5. 

This, is a small species of no unusual characters. It is nar- 
rowly cylindrical in form, shortly produced and pointed at its 
posterior extremity. It was described by Gabb from the Mio- 
cene of Santo Domingo, but Dall united with it, the oxytata 
Bush, in which case, it is also found in the recent fauna. Gabb 
records acuta d’Orb. from the Pliocene of the Limon Peninsula. 

Uscart Stage: Middle Creek. 


Genus BULLARIA, Rafinesque 


Bullaria paupercula Sowerby Plate 4, figure 14 


34d BULLETIN 39 206d 


Bulla paupercula Sowerby, 1849, Quart. Journ. Geol. Soc. London, 
vol. 6, p. 52. 

Bulla paupercula Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 
246. 

Bulla paupercula Guppy, 1874, Geol. Mag. London, p. 437. 

Bulla paupercula Guppy, 1876, Journ. Geol. Soc. London, vol. 32, p. 
518. 

Bulla paupercula Dall, 1890, Trans. Wagner Free Inst. Sci., vol. 3, 


pt. 1, p. 18. 
Bulla striata Dall, 1903, Trans. Wagner Free Inst., Sci., vol. 3, pt. 6, 


. 1583. 
ie ee Maury, 1917, Bull. Amer. Pal., vol. 5, p. 182, pl. 
Balen Se 
Bullaria paupercula Maury, 1920, N. Y. Academy of Science, Tertiary 
Mollusca of Porto Rico., vol. 3, pt. I, p. 75. 

This species is rare in Costa Rica, and all our specimens are 
from the Lower Gatun rocks of Middle Creek. It is possibly 
identical with the recent Aullaria amygdala Dillwyn, abundant 
on the beaches along the northern coast of Panama and Costa 
Rica. 

Gatun Stage: Middle Creek. 


Genus RINGICULA, Deshayes 
Ringicula dominicana Maury 
Ringicula semistriata ? Gabb, 1875, Trans. Amer. Phil. Soc., vol. 15, 
p. 225. Not #. semistriata d’Orbigny. 
Ringicula dominicana Maury, 1917, Bull. Amer. Pal., vol. 5, p. 185, 
Oi ain semegy Nae, 

This is the most abundant of the Opisthobranch mollusks in 
the Miocene beds of Costa Rica. The whorls are in general more 
strongly sculptured with incised lines than typical dominicana 
from the Miocene of Santo Domingo, and the spiral lines extent 
over the greater part of the shell,except for a wide, smooth band 
about the suture. 

The R. hypograpta Brown and Pilsbry,from Gatun is a larg- 
er shell with a heavier outer lip and more crowded spiral lines 
about the base. ‘The Costa Rican shells measure about 1.5 mm 
or less in height. 

Gatun Stage: Middle Creek. Coll. 5, Red Cliff Creek. 

Coll. 7, Estrella River. 


Ww 
qn 


207 Costa Rico M1ocENE—OLSSON 


ORDER CTENOBRANCHIATA 
(A) SUPER-FAMILY TOXOGLOSSA 


Genus TEREBRA ApDANSON 


Terekra subsulcifera Brown and Pilsbry 
Terebra subsulcifera Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. 
Phila., vol. 63, p. 339, pl. 22, fig. 7. 


Terebra (Myurella) subsulcifera Cossmann, 1913, Jour. de Conchylio- 
logie, vols 61, ps 14) ple tr, e.)25: 


The single, fragmentary specimen comes from the Gatun 
beds of Mt. Hope in the Canal Zone. From ézpartita, which it 
resembles in its sculpture, this species and haztenszs Dall, differ 
in having only a single columellar fold. 


Gatun Stage: Mt. Hope, Canal Zone. 


Terebracf. haitensis Dall Plate 1, figure 3 


Terebra haitensis Dall, 1895, Proc. U. S. Nat. Mus., vol. 18, p. 35. 
Terebra haitensis Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, pt. 


- 6, p. 1632, pl. 59, fig. 30. 
Terebra haitensis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 194. pl. 4, 


185) Bie 
The collection contains a single, imperfect shell from Sousci 
Creek, which should probably be referred to this species. From 
the swbsulcifera, it differs in its more numerous and oblique ribs, 
and wider, less pronounced spiral bands. Like subsulcifera, the 
columella carries a single, sharp plication. Our specimen con- 
sisting of only 5 whorls measures 37 by 13 mm. 


Gatun Stage: Sousct Creek, a branch of Fone Creek, C. R. 
Terebra bipartita Sowerby Platemmmehourent, 2. 


Terebra bipartita Sowerby, 1849, Quart. Jour. Geol. Soc. London, vol. 
5; P- 47. 


36 BULLETIN 39 208 


Terebra bipartita Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 
225. 

Terebra (Acus) bipartita Dall, 1895, Proc. U.S. Nat. Mus., vol. 18, 
p-. 38. 

Terebra bipartita Maury, 1917, Bull. Amer. Pal., vol. 5, p. 187, pl. 3 
fig. 14. 


This species, possessing much the same type of sculpture as 
sulcifera, subsulcifera, haitensis etc., is recognized by having two, 
instead of one columellar plication. Full-grown specimens 
have a bipartite sculpture while in sz/cifera and the others it is 
tripartite. 


The Costa Rican collection contains three specimens from 
Saury creek, near Cahuita, the largest of Io incomplete whorls 
mearures 77 by I9 mm. 


Gatun Stage: Zone E. Saury Creek 


Terebra gatunensis Toula Plate 1, figures 4-6 


Terebra gatunensis Toula, 1909, Jahrb. der K-K Geol. Reichsanstalt, 
vol. 58, p. 705, pl. 25, fig. 14. 

Terebra gatunensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. 
Phils.) volii63)) p.339pls.22sihige 2s 

Terebra wolfgangi Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. 
Phil., vol. 63, p. 340, pl. 22, figs. 1, 3-6, but not of Toula, 1908. 


Terebra (Myurella) gatunensis Cossman,1913,Jour. de Conchyliologie, 
vol, 61, p. 13, pl. 1, figs. 26-29. 


In the Canal Zone, the Gatun beds contain two common 
species of Terebra, which were described by Toula as gatun- 
ensis and wolfgangi. ‘These species have been more or less con- 
fused by Brown and Pilsbry, so that their figures and description 
of wolfgang?, really represent gatunensis. Their figure 2 isa 
large and less common variety of gatunensis. ‘Toula’s figure, al- 
though of a young imperfect shell, is clear and shows the main 


209 Costa RicA MiockENE—OLSSON 


Ww 
“I 


characters of the species, the spirally striated, faint sutural sul- 
cus and fine spiral scu)lpturing of the whorls. He however in 
his notes, allies this species with the recent 7. dislocada of Say, 
which is misleading. Toula’s figure can be duplicated by scores 
of specimens in our collection. 


Terebra gatunensts is characterized by its large size, (length 
50 mm or more), its faint sutural sulcus which may be smooth 
or with 2 or 3 faint spirals, showing best on young shells. The 
spiral cords of the main body of the whorl are variable in num- 
ber, spacing and strength, as well as the longitudinal ribs. A 
large coarse variety, with persistent sulcus, and heavier sculp- 
ture occurs rarely and is shown by Brown and Pilsbry as their 
figure 2. The pillar is nearly smooth, carrying a very broad 
fold above, and a small one below. 


Gatun Stage: Gatun and Mt. Hope, C. Z. 
Terebra wolfgangi Toula Plate 1, figure 11, 12 


Terebra wolfgangi Toula t909, Jahrb. der K-K Geol. Reichsandstalt, 
vol. 58, p. 705, pl. 28, fig. 7. 


Not Zerebra wolfgangi Brown and Pils. which is 7. gatwnensis Toula. 


Although 7. gatunensis and wol/gangz have not been collect- 
ed by us except in the Canal Zone, where they are abundant, 
they are included here for the sake of completeness. It is also 
not unlikely that future collecting may still reveal their presence 
in the synchronous beds in Costa Rica and adjacent parts of 
Panama. 


T. wolfgangi differs from the common 7. gatunensis by its 
long, tapering form, deep, persistent sulcus anda sutural band 
generally free from spirals. It is also a much smaller species, 
the largest specitnen in our collection measuring about 38 mm in 
length. 

Gatun Stage; Mt. Hlope, C. Z. 


38 ; BULLETIN 39 220 


Terebra gausapata Brown and Pilsbry Plate 1, figures 8, 10 


Lerebra gausapata Brown and Pilsbry, April 1911, Proc. Acad. Nat. 
Sci) Phila.) iveli163,)p./ 340) (pls 22) fies. (8. 9: 

Tevebra acuaria Toula, Dec. 1911, Jahnb. der K-K Geol. Reichanstalt, 
vol. 61, p. 19, pl. 2, fig. ro. 


Tevebra (Myurella) acuaria Cossm., 1913, Journ. de Conchyliologie, 
vol. 61, p. 12, pl. 1, figs. 19-24. 


The 7. gausapata, was described by Brown and Pilsbry, 
from young specimens, measuring only 19 mm in length. Later 
in the same year appeared Toula’s 7. acuaria, also based on 
young shells. 


The important characters of this species, are its nearly 
straight ribs, deep sulcus, a sutural band with faint spirals 
and even sculpture. Figures 23 and 24 of Cossmann I believe 
represents wolfgangt. T. wolfgang? as will be noted from the 
accompanying figures is a much more slender species and usually 
with a smooth sutural band. The pillar carries two broad folds, 
which are better developed than in wol/gangi or gatunensts. 


Length 33, breadth 8.5 mm. 


Gaiun Stage; Mb. Tope (Cay 
Water Cay. 


Terebra spirifera Dall Plate 1, figure 13 
Terebra dislocata Gabb, 1873, Trans. Amer. Phil. Sci., vol. 15, p. 225, 
Not of Say. 


Terebra (Acus) bipartita Sowerby, variety spirifera Dall, 1895, Proc. 
U. S. Nat. Mus., vol. 18, p. 38. 


Terebra (Oxymeris) bipartita Sowerby, variety spirifera Dall, 1903, 
Trans. Wagner Free Inst. Sci., vol. 3, pt. 6, p. 1632, pl. 59, fig. 13. 


Terebra spirifera Maury, 1917, Bull, Amer. Pal., vol. 5, p. 188, pl. 3 
figs. 15, 16. 


In 7. Gatunensis and wolfgang? the pillar is nearly smooth 


Axi Costa R1icA M1rockNE—OLSSON 39 


or with two board, sub-obsolete folds. In spzrifera, the two folds 
are strong and recall those seen in the recent 7: dislocata. The 
sutural band is strongly sculptured by the end of the ribs which 
are entirely dislocated by the deep sulcus. Typical specimens 
have three or four, even spirals, which are stronger than the 
longitudinal ribs. Variation occurs in the number, and regu- 
larity or these spirals, 


Length 43, breadth 9 mm. 


Gatun Stage: Banana River, Hill No. 1. 
Water Cay. 


Terebra spirifera Dall, variety midiensis, n. var. Plate 1 figure 7 


This variety, represented only by a single specimen from 
Middle Creek, C. R., differs from typical spzrifera, in having on- 
ly three spirals, which are broad and strap-like in form. 


Gatun Stage: Middle Creek. C. R. 


Terebra Berlinerae Maury Plate 1, figures 17, 18 


Terebra Berlinere Maury, 1917, Bull. Amer. Pal., vol. 5, p. 198, pl. 
Als SONA, 9/3) le 


This beautiful species was described by Dr. Maury, from the 
Cercado (Miocene) formation of Santo Domingo. ‘The whorls 
are slightly convex, and its sculpture-is closer, finer and more 
delicate than is seen in any of the associated Costa Rican Tere- 
bras. Our specimens, all imperfect, were collected from the 
hard Gatun sandstones of Boucary Creek. The most complete 
specimen measures 21 by 6.5 mm. 


Gatun Stage: Boucary Creek. 


40 BULLETIN 39 | 21zZ 


Terebra estreliana, n. sp. Plate 1, figures 9, 16 


Shell small; nucleus of three smooth tapering whorls; post- 
nuclear whorls about 9, sculptured with a faint sutural sulcus, 
which is lacking or nearly so from the early whoris; axial sculp- 
ture of about eleven, straight or slightly oblique ribs which pass 
from suture to suture; but are nearly lacking from the base; 
spirals lacking or very faint on the spire-whorls, so that they ap- 
pear smooth and solid; the base of the last whorl is strongly con- 
tracted and shows four or five, very faint spirals; pillar straight 
and smooth; beak twisted. 


Length 7.5, diameter 2.5 mm. 


This is a small species of rather solid appearance, strong 
ribs, but otherwise nearly smooth whorls. The spirals are very 
faint and are seen best only when viewed at right angles to the 
source of light. Our specimen came from the Estrella River. 


Gatun Stage: Zone 7, Estrella River, C. R. 


Terebra costaricensis, n. sp. Plate 1, figure 19 


Shell small and slender, with about three, long, tapering 
smooth, nuclear whorls; post-nuclear whorls eleven; sutural 
band rather large and prominent, strongly tuberculated; the 
spire-whorls are slightly concave and have at their base or just 
above the suture, a revolving series of tubercles, which are the 
enlarged bases of the longitudinal ribs; longitudinal ribs on the 
last whorl number about twelve, are nearly straight and large 
and strong on each side of the suture, low and weak in the mid- 
dle of the whorls; the surface of the whorls are otherwise smooth 
or in some shells feebly spiralled; base slightly rounded with 
eight or more spirals which become finer as they approach the 
canal; the longitudinal ribs are continued across these spirals. 

Length 13, diameter 3 mm. 


The above description is based mainly on two specimens 


214 Costa Rica M1ockNE—OLSSON Al 


from Hill No. 3, of the Banana River. Our collection contains 
several smaller shells from Zone 7, of the Hstrella River in 
which the base is somewhat more heavily sculptured and with 
very fine spirals over the main surface of the whorls. 


Terebra benthalis Dall var. bocasensis, n. var. Plate 1, figures 25, 26 


Cf. Terebra (Acus) benthalis Dall, 1889, Bulli. Mus. Comp. Zool., vol. 
18, p. 65, pl. 29, fig. 6. 

This species, fairly abundant in the shale beds, just north 
of the city of Bocas del Toro, Panama, agrees closely with Dall’s 
figure in the Blake Report, except that the revolving series of 
tuberclesin the fossil shells seem to be lower and broader. 
The whole surface on slight magnification is seen to be finely 
spirally striated. 

Terebra benthalis was described by Dall from specimens 
dredged from Ioo to 400 fathoms of water off the Morro Light, 
Havana, Cuba. The species belongs to the section Fusoferebra 
Sacco. 

Length 27, diameter 6.5 mm. 


Gatun Stage: Bocas del Toro, Panama. 
Terebra pumbriensis, n, sp. Plate 1, figure 27 


Shell of medium size, composed of 9 whorls (type with a 
small portion of the apex lost); the early post-nuclear whorls 
(the 1st 3 or 4) with two revolving rows of tubercles, formed by 
the intersection of fine, oblique riblets and two spiral bands; on 
the 4th whorl of our specimen, a median spiral makes its ap- 
pearance and rapidly increases in strength so that the later 
whorls have three rows of tubercles, the upper or most posterior 
one being somewhat the strongest; the longitudinal ribs are nar- 
row and oblique, and on the last whorl number about 20; the 
space between the ribs have three or four fine, incised growth- 
lines; periphery of the base has an additional tuberculated spiral, 


42 BULLETIN 39 214 


below which is a band composed of four smooth, close spirals ; be- 
tween these spirals and the keel is a deep, concave zone; pillar 
nearly straight and smooth, with a single sharp keel on the 
back. 


Length 20, diameter 4.75 mm. 


Of this interesting and distinctive species, we have only a 
single, imperfect specimen from the Gatun beds of Pumbri 
Creek. The three rows of tubercles are evenly spaced and there 
is no sutural band. 


Gatun Stage: Pumbri Creek, C. R. 
Genus CONUS Linné 
Conus molis Brown and Pilsbry Plate 2, figures 1, 2 


Conus molis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., 


VOl6gh pi 242 wplyi2guntomis 
Conus molis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 200. 


This is the common Cone in the Gatun beds of Panama and 
Costa Rica and large specimens may reach a length of nearly 150 
mim. ‘The upper surface of the whorls carry 5 or more fine 
spirals and perfect examples of young shells have a few of the 
early post-nuclear whorls finely coronated. Described by Brown 
and Pilsbry from the Canal Zone, the species has also been col- 
lected by us at several stations along the Panama coast between 
Colon and Bocas, at Water Cay, and along the Banana River in 
Costa Rica. A large specimen from Water Cay measures: 
length 144 mm, width 81 mm. 


Gatun Stage: Gatun, Water Cay, Banana River, ete. 


Conus stenostomus Sowerby Plate 3, figure 2 


Conus stenostoma Sowerby, 1849, Quart. Jour. Geol. Soc. London, 
vol. 5 p. 44. 

Conus stenostoma Guppy, 1866, Zd., vol. 22, p. 287, pl. 16, fig. 2. 

Conus stenostoma Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 
230. 


i 
ae 
CH 


Costa Rico MioceNE—OLSSON 43 


Conus catenatus Guppy, 1876, Quart. Jour. Geol. Soc. London, vol. 32, 
Pp. 527, in part b 

Conus stenostomus Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, 
pt. 6, p. 1583. 


Canus stenostomus Maury, 1917, Bull. Amer. Pal., vol. 5. p. 203, pl. 6, 
fig. 4. 


A species characterized by its sharp angulated shoulder, 
concave upper surface of its whorls and attenuated anterior can- 
al. Well-preserved examples showing a highly polished sur- 
face and traces of yellow blotches of color markings were col- 
lected quite abundantly in the coralline of Port Limon. The 
species also occurs in Jamaica and Santo Domingo. 


Gatun Stage: Port Limon. 


Conus concavitectum Brown and Pilsbry 


Conus concavitectum Brown and Pilsbry, 1911. Proc. Acad. Nat. Sci. 
Philal vol. 62. p. 341, pl. 23. figs 54 6. 

Conus (Lichoconus) concavitectum Cossmann, 1813, Jour. de Conchy- 
liologie, vol. 61, p. 43, pl. 4, figs. 3, 4. 


In adult shells, the spire whorls become concave. The 
species is also recognized by its sharp shoulder and low spire. 
It may be confused with varieties of proteus but the sharp should- 
er angle and spiralled spire whorls will separate it. Our speci- 
mens come from Water Cay, and measure 48 by 27 mm. 


Gatun Stage: Canal Zone and Water Cay. 


Conus proteus Hwass Plate 2, figures 3, 4 


Conus proteus Wwass, 1789, Enc. Meth. vers, 1 pt. 2, p. 682. 

Conus leoninus WHwass 1789, Enc. Meth. vers, 1 pt. 2, p. 683. 

Conus proteus ? Gabb, 1873, Trans, Amer. Phil. Soc., vol. 15, p. 232. 

? Conus Berghausit ? Gabb, dem, p. 232. Not of Hoernes, Foss. Wien. 
Becki ly ii hige 38 

Conus leoninus Gabb, 1881, Jour. Acad. Nat. Sci. Phila., vol. 8, 2nd 
series, p. 359. 


44 BULLETIN 39 216 


Conus proteus Dall, 1890, Trans. Wagner Free Inst. Sci., vol. 3, pt. 1, 
p. 26. 


Conus proteus Maury, 1917, Bull. Amer. Pal., vol. 5, p. 206, pl. 6, 
sifegy 10h 


This isa common species in the Miocene and Pliocene beds of 
Costa Rica and Panama. It is generally possible to distinguish 
two forms, a smaller one which agrees with typical examples of 
proteus in form and coloration (which is frequently preserved, ) 
and a larger and heavier type, like the so-called /eontnzus. ‘This 
second variety may show several dark spiral bands in addition to 
the square yellow blotches. Such markings we have not observ- 
ed on recent examples of /eonznus. 


Gatun Stage: Banana River, Port Limon, Water Cay. Plio- 
cene: Monkey Point; Bocas. 


Conus Veatchi, n. sp. Plate 2, figures 5, § 


Shell rather large, heavy and broad, with a low, nearly flat 
spire, but pointed, slightly projecting apex; spire-whorls about 
Io in number, witha slightly raised border near the suture and 
marked with fine spirals which on the last whorl number ro or 
11; shoulder of the last whorl rounded or beveled; last whorl, 
large and broad, nearly smooth above, except for the very faint 
spirals, but with numerous, fine, irregular spirals on the canal; 
pillar straight; aperture straight and narrow, slightly wider 
below. 


Length 43, breadth 28 mm. 


The present collection contains two specimens, the larger of 
which from Water Cay, serves as the holotype. The species is 
very unlike any of the associated Gatun species in its low, near- 
ly flat spire. The Conus demiurgus Dall, of the Chipola Mio- 
cene of Florida, is a related species. 


It is named for Dr. A. C. Veatch, Director of the Hxplora- 
tion Work of the Sinclair Consolidated Oil Corporation, for his 


217 Costa RicA MIocENE —OLSSON 45 


interest and co-operation in obtaining the extensive collections 
which form the basis of the present work. 
Gatun Stage: Water Cay, Panama. 


GOO On VE 
Conus floridanus, var. costaricensis, n. var. Plate 3, figures 3, 9 


Shell conic, witha moderately high spire of about eleven 
whorls; profile of the spire flat or slightly concave to the long 
projecting apex seen in well-preserved specimens; shoulder angle 
rather sharp with the whorls slightly concave above between the 
sutures; spire-whorls generally show 3 or more faint, raised 
spirals and strongly arcuated growth lines; the last whorl below 
the shoulder angle is usually nearly ‘smooth above (sometimes 
with faint spirals) but with heavy regular spiral groovings to 
the number of about fifteen on the canal; the canal is long, rap- 
idly attenuated and usually somewhat flexed below. 


Height 44, breadth 21 im. 
Type from the Gatun beds of the Banana River. 


This fossil, quite comimon in Costa Rica, should probably be 
classed as a variety of the recent Conus floridanus Gabb, differing 
only in always being somewhat more slender. They also agree 
closely with Dall’s figure of Conus chifolanus Dall from the 
Chipola Miocene of Florida. 

Care should be taken in distinguishing this from C. 
zmitator of the Canal Zone. In that species, the full-grown shell 
is larger, with nearly straight sides and finer irregular spirals on 
the canal. 


Gatun Stage: Banana River; East Grape Point Creek. 
Conus imitator Brown and Pilsbry Plate 2, figure 6 


Conus tmitator Brown and Pilsbry, April, Ig11, Proc. Acad. Nat. Sci. 
Phila., vol. 63, p. 342, pl. 23, fig. 4. 

Conus Dallt Toula, Dec., 1911, Jahrb. der K-K Geol. Reichsanstalt, 
Wien, vol. 61, p. 508, pl. 31, figs. 23 a-d. 


46 BULLETIN 39 218 


Conus (Lithoconus) Dalli Cossmann, 1913, Jour. de Conchyliologie, 
VOM GI pA pls Baines 2 OniaT ama MAN TO SEIZE Sy 

? Conus, Pall Maury, 1917, Bull! Amer Pale vol. 5) p.) 212), pl. gy 
fig. 15. 

The Conus Dalli Toula is doubtless the young of zmztator 
described earlier in the same year by Brown and Pilsbry. Fig- 
ure 6 is of a specimen from Mt. Hope which measures 51 by 2: 
mim. From Costa Rica, the collection contains a series of per- 
plexing Cones, which probably should be referred to this speces. 
From costaricensis they differ in their nearly straight sides, more 
irregular spiral groovings on the anterior canal, and they usually 
show faint spirals over most of the body-whorl, which I have 
not observed on costaricensis. A large shell from Zone C of 
Comadre Creek near Cahuita, measures 65 by 29 min. 


Gatun Stage: Mt. Hope and Gatun C. Z. 
Water Cay, Panama. 
Banana River, East Grape Point Creek, 
Comadre, Middle Creek, C. R. 


Conus recognitus Guppy Plate 2, figure 9 


Conus solidus Sowerby, 1849, Quart. Jour. Geol. Soc. London, vol. 6, 
p- 45, Not C. solidus Sowerby, 1841. 

Conus solidus Guppy. 1866, Quart. Jour. Geol. Soc. London, vol. 22, 
P. 287.0 pl. (26) fig.) 1. 

Conus recognitus Guppy, 1867, Proc. Sci. Assoc. Trinided, p. 171, Har- 
ris Reprint, Bull. Amer. Pal., vol. 8, p. 198. 

Conus pyriformis Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 229, 
Not of Reeve, 1843. 


Conus vecognitus Guppy, 1876, Quart. Jour. Geol. Soc, London, vol. 


32, Pp. 527. 

Conus vecognitus Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, pt. 
(CN OBInansgo rey 

Conus recognitus Maury, 1917, Bull. Amer. Pal., vol. 5, p. 209, pl. 7, 
fig. 9. 


This species is easily recognized by its well-rounded should- 
er and appressed sutures. It is closely related to the recent C. 


BLO Cosra Rica M1ockENE—OLSSON 47 


pyriformis Reeve of the west coast of Central America but the 
fossil shells have generally a lower and flatter spire. It is also 
known from the Miocene of Santo Domingo, Jamaica and 


Trinidad. 
Gatun Stage: Banana River, Hill No. 3. 


Conus limonensis, n. sp. Plate 3, figures 7, 8 


Shell rather large, but delicate and rather slender; spire low, 
concave to the slightly projecting apex; spire whorls Io or more, 
nearly flat between the sutures or slightly concave due toa 
slightly raised border; spire-whorls with 3 or 4 low spirals which 
become faint on the later whorls so that they are nearly smooth; 
sutures deep; shoulders of the last whorl angled or somewhat 
raised; body-whorl long and slender, polished and smooth, ex- 
cept fora few faint spirals near the tip of the canal; anterior 
canal long and straight, evenly tapering. 

Length 41, breadth 24 mm. 


This species is based on 2 specimens from the coral lime- 
stones of Port Limon, where it is associated with Conus stenost- 
omus and planiliratus. Froim stenostomus, its straight canal, low 
spire and nearly smooth, polished surface is most noticeable. 


Gatun Stage.) Port Lemon, \C. R: 
Conus musaensis, n. sp. Plate 1, figures 22, 24 


Shell small and solid; spire short, conic, composed of about 
2 pellucid, globular and projecting whorls and 6 post-nuclear 
whorls; the spire-whorls are flat or slightly cancave and project 
slightly over the anterior sutures; body-whorl with the upper 
half smooth, but with 12 deep grooves on the anterior canal 
which produce wide, spiral bands: these grooves are delicately 
incised by raised longitudinal lines; color markings are some- 
times preserved, which appear as 8 narrow, gray bands on the 
“upper half of the whorl. 


Ny 


Length 19, breadth 9 mm 


This small species, the Gatun analogue of the recent West 


43 BULLETIN 39 220 


Indian Conus Agassizt Dall, is abundant along the Banana 
River and elsewhere in Costa Rica. 


Gatun Stage: Banana River, Zone 6 Red Cliff Creek, C. R. 
Conus cf. interstinctus Guppy Plate 3, figure 12 


Conus interstinctus Guppy, 1866, Quart. Jour. Geol. Soc. London, vol. 
22) DA 2oS ime MO. sh oa ah 

A single specimen from Hill 1, of the Banana River, agrees 
closely with Guppy’s figure of this Jamaican species. The shell 
measures 39 by zo mm. ‘The spire is moderate in height, with 
a slightly concave profile. The spire-whorls are smooth between 
the sutures but also show a slight coronation. ‘The last whorl 
carries about 15 narrow, widely spaced, spiral cords which an- 
teriorly are slightly beaded. 

Dr. Maury unites zzdersttuctus, with Sowerby’s catenatus, 
but that species is based on a very young and scarcely distin- 
guishable specimen. We therefore prefer to compare the Costa 
Rican sheil with Guppy’s larger and well-figured zzterstinctus. 


Gaiun Stage: Banana River, Hill No. 1. 
Conus cf. Sewalli Maury 


Conus Sewalli Maury, 1917, Bull. Amer. Pal., vol. 5, . 201, pl. 5, fig. 3, 
pl. 6, fig 3. 

The collection contains 2 young shells which may belong to 
this Dominican species. They differ from the preceding zter- 
stinctus in being shorter and broader, and the spire-whorls in 
place of being smooth are marked with 5 or 6 strong spirals. The 
last whorl is sculptured with 15 or more widely spaced beaded 
spirals. The dimensions of the largest shell are: Length or 
height 30, breadth 19 mm: 


Gatun Stage: Banana River. 
Conus toroensis n. sp. Plate 2, figure 7 


Shell of medium size, with a conic spire a little more than 
¥ the length of the aperture; the whorls of the spire numbering 


2ar Costa RicA MI1ocRENE—OLSSON 49 


12 plus are flat or slightly concave and bordered below by a low 
carina projectly slightly above the suture; this carina on all ex- 
cept the last whorl bears low nodes, about 20 to the later whorls; 
the spire-whorls are otherwise smooth except for the arcuate 
growth lines; the last whorl is a tapering cone, nearly smooth, 
except for the faint spirals about its lower one-third; the aper- 
ture is narrow, straight, with a thin arcuate outer lip. 


Height 45, diameter 21, aperture 33 mm. 


This shell belong to the Conus consobrinus group, agreeing 
in its form and strongly nodulated spire whorls. The main 
difference is that éovoensis is nearly smooth, the spirals showing 
only on the lower one-third, while cozsobyinus has most of the 
last whorl covered with beaded spirls. 


Gatun Stage: Toro Cays. 


Conus (Chelyconus) tortuosostriatus Toula Plate 1, figure 15 


Conus (Chelyconus) tortuosostriatus Toula, 1911, Jahrb. der K-K. 
Geol. Riechsandstalt, Wien, vol. 61, p. 508, pl. 31, fig. 22. 

Conus (Hemiconus) tortuosostriatus Cossmann, 1913, Jour. de Conchy- 
liologie, vol, 61, p. 40, pl. 3, figs. 28, 29. 

Conus tortuosostriatus Maury, 1917, Bull. Amer. Pal., vol. 5, p. 205, pi. 
6, fig. 9 


This shell should probably be considered as a short variety 
of gracilissimus Guppy, differing chiefly in its broader form and 
shorter spire. 


The body-whorl is sculptured with about 24 raised,even, spiral 
cords, separated by deep, even interspaces. These interspaces 
are crossed by neat, incised lines, corresponding to the lines of 
growth. ‘The nucleus consists of about 3, long, tapering smooth 
whorls, after which follow several turns of carinated and finely 
coronated post-nuclear whorls. 


Length 22, breadth 9 mm. 
Gatun Stage: Bocas del Toro. 


50 BULLETIN 39 222 


Conus granozonatus Guppy Plate 3, fig2 15 


Conus granozonatus Guppy, 1866, Jour. Geol. Soc. London, vol. 22, p. 
237) MplyTOw ioe 
This is a rare species in Costa Rica, our collection containing 
only a single specimen from the Gatun beds of the Rio Blanco. 
It is characterized by a rather high, conic spire, with smooth or 
only slightly coronated whorls. ‘The last whorl has about 18, 
wide, spiral bands, which areheavily granulated above. The 
specimen agrees well with Guppy’s figure of the Jamaican 
shell. 
Length 25, breadth rz mm. 
Gatun Stage: Rio Blanco, C. R. 


Conus planitiratus Sowerby Plate 3, figures 10, 13 


Conus planiliratus Sowerby, 1849, Quart. Jour. Geol. Soc. London, vol. 
6, p. 44. 

Conus planiliratus Guppy, 1866, Sdem. vol. 22, p. 287, pl. 16, fig- 
Tbe) P77 


Conus planiliratus Gabb, 1873, Trans. Amer. Phil. Soc., p. 230, in 


part. 

Conus planiliratus Guppy, 1876, Quart. Jour. Geol. Soc. London, vol. 
Aiea! ON) ENaLeNe 

Conus planiliratus Dall, Trans. Wagner Free Inst. Sci., vol. 3, pt. 6, 
Pp. 1583. 


Couus planiliratus Cossmann, Jour. de Conchyliologie, vol. 61, p. 48, 
PIs wmSs25h2onbore 

Conus regularis Gabb, Jour. Acad. Nat. Sci. Phila., vol. 8, 2nd series, 
p- 359, pl. 46, figs. 45-48. Not of Sowerby. 

Conus planiliratus Maury, Bull. Amer. Pal., vol. 5, p. 209, pl. 7, 
fig. 10. 

The occurrence of this species in Costa Rica is limited to 
the coral limestones of Limon, Gabb had specimens from here 
which he identified with the recent West Coast Conus regularis 
of Sowerby. The Limon shells agree very well with Guppy’s 
figure in the Quarterly Journal and with Cossmann’s figures of 
Bowden and Mindi specimens. The species appears to be closely 


223 Costa Rico MirocENE—OLSSON 51 


related to the recent Conus Stimpsont Dall from the Florida 
Seats CbTOe NUE, Sah Nabi Mids.) voli 24m pa503,pluizon tion!) 7! 
1903). 

Surface of whorls are heavily sculptured with 25, 30 or 
more subequal spiral cords. Upper surface of whorls, flat or 
slightly concave and with 3 or more spirals. ‘The shoulder is 
sharp and bounds a rather high, conic spire. 

Length 31, breadth 16 mm. 


Gatun Stage: Port Limon. 
Conus planiliratus, var. hocasensis, n. var. Plate 3, fig. 14, 16 


Shell of medium size, solid witha broad, stubby outline; 
spire low, conic with nearly straight sides and composed of seven 
or more whorls; the rst 4 or 5 post-nuclear whorls are coronated; 
spire-whorls with 3 faint spirals and curved growth-lines; body- 
whorl large and wide, with 25 strong but somewhat irregular 
simple spiral cords; their interspaces are wide and with crowded, 
raised lines of growth and sometimes intercalated spirals; anter- 
ior canal nearly oe 


Length 32, breadth 17 mm. 


This Cones which should probably be considered as distinct, 
is fairly common in the shale beds, belonging to the Gatun 
Stage on Bocas Island. From typical planiliratus it GTS in its 
broad stubby form and coarse sculpturing. 


Gatun Stage: Bocas del Toro, Panama. 
Conus marginatus Sowerby Plate 1, fig. 20 


Conus marginatus Sowerby, 1849, Quart. Jour. Geol. Soc. London, 
vol. 6, p. 44. 

Canus marginatus Gabb, 1873, Trans. Amer. Phil. Soc., vol. 16, p. 
230. 

Conus marginatus Guppy, 1876, Quart. Jour. Geol. Soc. London, vol. 
32, p. 528, pl. 20, fig. 5. 

Conus marginatus Maury, 1917, Bull. Amer. Pal., vol. 5, p. 210, pl. 7, 
shoes 


52 BULLETIN 39 224, 


On the basis of Guppy’s figure in the Quarterly Journal, 
this cone is seen to belong to the type exemplified by the recent 
C. verrucosus, with its sulcated sides and appressed or beveled 
shoulder angle. Our collection contains a single specimen from 
Zone 5 of Red Cliff Creek which is very close to Guppy’s figure. 
Cossmann’s specimen of marginatus from Martinique should 
probably be referred to some other species. 


Gatun (stages: Zone Gi ean Glas Graces GLiree 
Conus gracilissimus Guppy Plate 1, figure 14 


Conus gracilissimus Guppy, 1866, Quart, Jour. Geol. Soc. London, 
vol. 22, p. 288, pl. 16, fig. 4. 

Conus Orbignyi Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 230, 
Not C. Ordtenyt Audonin, 1830. 

Conus gracilissimus Gappy, 1876, Quart. Jour. Geol. Soc. London, vol. 
32, P- 527+ 

Conus gracilissimus Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, 
DEVON spe SOs: 

Conus gracilissimus Cossmann, 1913, Jour. de Conchyliologie, vol. 61, 
Plea mess 

Conus gracilissimus Maury, 1917, Bull. Amer. Pal., vol. 5, p. 204, pl. 
6, figs. 8. 

A small, but long and slender cone, sulcated with 25 or 
more, strong, regular spiral cords. The spire is long and slend- 
er with slightly coronated whorls and the area between the su- 
tures is delicately decussated by 3 or more spirals crossed by the 
raised and arcuated lines of growth. 


Our collection contains specimens from Bocas and Limon, 
the largest measuring 26 by 9 mm. 


Gatun Stage: Bocas del Toro, Pan., Port Limon, C. R. 


Conus Burckhardti Bose Plate 3, figures 4, 5 
Conus Burckhardti Bose, 1¥906, Bull. de Inst. Geol. de Mexico, 
numero 22, p. 50, pl. 5, figs. 39, 40. 


Shell of moderate size, witha rather high slightly concave 
spire and long, acute apex; sutural whorls smooth, except for the 


YI 
W 


225 Costa Rica MIocENE —OLSSON 


faint arcuate lines of growth and very indistinct spirals; the 
earlier whorls have a carinated suture but are only very faintly 
coronated; last whorl heavily sculptured with about 20, wide, 
even spiral bands, separated by grooves of about 1-3 of their 
width; these grooves are faintly incised by the longitudinal 
lines of growth; the spiral bands are slightly wider above 
and smooth, becoming granulated on the canal; this granulation 
appears on the posterior side of the bands only; aperture long and 
narrow. 


Length 36, diameter 14.5 mm. 
33 (apex broken) diameter 15 mm. 


The original specimen of Conus Burckhardti was described 
from the Miocene of Mexico, State of Oaxaca. Our specimens 
from Panama and Costa Rica area little larger, but show the 
characteristic high, smooth, spire and strong, flat, spiral bands of 
the body-whorl. 


Conus tortuosopunctatus "Toula is a similiar but smaller 
species. The difference in sculpture of the two species may be 
seen in the accompanying figures, the spiral bands of fortuoso- 
punctatus being double or incised, while in Burckhardt, they are 
broad and simple. 


Gatun Stage: Water Cay, Panama. 
Zone 2, East Grape Point Creek. 


Conus Harrisi, n. sp. Plate 3, fig. I 


Shell of moderate size, subcylindric with a high, even, conic 
spire; spire-whorls about 6 plus, flat, with close sutures, except 
for the upper whorls which are slightly carinate; the surface of 
the spire-whorls is nearly smooth, except for a few faint spirals; 
the last or body whorl is strongly sculptured with widely spaced 
and sub-regular grooves; these grooves form about 14 wide, 
spiral bands about the lower 3-4th of the last whorl; the upper 
ath below the shoulder angle forms a wide, smooth band; below, 
the spiral bands are generally smooth except those about the lower 


54 BULLETIN 39 226 


half of the shell, which may become, mesially divided and bear 
afew small granulations or beads about their upper half; aper- 
ture sublinear. 

Height 51, diameter 23 mm. 


A single specimen of this elegant species was collected from 
the Gatun of the Banana River. It is a species of the Conus 
Burckhardt type but much larger and when perfect reaching a 
length of nearly 60 mm. The sculpture is of wide bands formed 
by deep, and regularly spaced grooves about the lower 3-4ths of 
the shell. 


Gatun Stage: Hill No. 3, Banana River. 


Conus multistriatus Bose Plate 1, figures 21, 23 


Conus Agasstzt Dall, variey multistriatus Bose, 1906, Bull. de Inst. 
Geol. de Mexico, numero 22, p. 49, pl. 5, figs. 34-38. 


Conus gaza Johnson and Pilsbry, 1g91t, Proc. Acad. Nat. Sci. Phila., 
vol. 63, pl. 23, figs. 2, 13. 


Conus gaza | Maury,.)1917, Bulle) Aimer) kale, vol) '5;)) sp. ) 210.) play 
Aigehen2) 

This is the Conzws gaza of Johnson and Pilsbry described 
from the Miocene of Santo Domingo and the Isthmus. It isa 
small, pretty species, recognized by its nearly biconic outlines 
and neat sculpture. The spire is high; with nearly smooth, spire- 
whorls and carinated sutures. The last whorl is rather short and 
heavily sulcated with about 18, regular, spiral cords, separated 
by intervals of their own width. 


All our specimens come from Water Cay, where the species 
is fairly abundant. The largest specimen measures 20 by II 
mm. Bose’s original specimens are from the Miocene of the 
State of Oaxaca, Mexico. They are somewhat larger, measuring 
Zieh) OMA), a's) panbaol yi 

Gatun Stage: Water Cay. 
Conus tortuosopunctatus Toula Plate 3, figures 6, 11 


Conus tortuosopunctatus Toula, 1911, Jahrb. der K-K. Geol. Reichsan- 
stalt, Wien, vol. 61, p. 507, pl. 31, fig. 21. 


On 
Ui 


227 Costa Rica MrocknE—OLSSON 


Shell small, with a high smooth, conic spire of about 12 
whorls the earlier post-nuclear whorls are finely nodulated, the 
succeeding have the sutures carinated and then simple; body- 
whorl with 20 or more strong, beaded, spiral cords; three spiral 
cords are usually double or incised with a median line, especially 
on the anterior canal; the beading of the spiral cords is develop- 
ed best on their posterior portions; interspaces of the cords near- 
ly as wide as the cords themselves, smooth or with faint, longi- 
tudinal lines and frequently a small median spiral. 


Length 30, breadth 13 mm. 
Gatun Stage: Mt. Flope. 


FAMILY TURRITIDA 
Genus TURRICULA Schumacher 
SECTION SuRCcULA H. anp A. ADAMS 
Turricula lavinoides, n. sp. Plate 4, figure 6 


Shell rather large and moderately solid; whorls about 12 in 
number, heavily sculptured with ribs and spiral cords; nucleus 
of two smooth, slightly bulbous whorls; post-nuclear whorls in- 
creasing uniformily in size; concave above, widest below the per- 
iphery; suture strongly appressed with a spiral cord in front of 
it; anal fasciole flat or concave, appearing nearly smooth except 
for about three, small, widely spaced spirals; remainder of 
whorls strongly sculptured with 8 or 12 axial ribs which are 
lacking from the anal fasciole and from the base of the last 
whorl; spiral sculpture of 3 or 4 strong cords on the spire-whorls, 
becoming 10 or more on the last whorl in addition to those of the 
anterior canal; finer, intercalated threads are present between 
the main spiral cords on the later whorls; the interval between 
the spirals is finely, longitudinally striated by long growth lines; 
anterior canal long and nearly straight, slightly calloused along 


56 BULLETIN 39 228 


the inner lip; outer lip thin, with a shallow anal sinus, lying in 
the sutural fasciole. 


Length 54, diameter 20, last whorl 38 mm. 


This elegant species may be compared with the 7. jaguensis 
Sowerby from the Santo Dominican Miocene and the recent 7. 
lavinia Dall from the West Coast of Mexico. The Dominican 
species lacks the sutural cord, as well as differing in details of its 
sculpture. 7. /avinia, appears to be more closely related, differ- 
ing in its finer sculpture and proportions. 


Gatun Stage: Hill ra, Banana River. 
Turricula favinoides var. limonensis, n. var. Plate 4, figure 12 


Shell of medium size and with about 10 whorls; nuclear 
whorls about 2 in number, at first smooth, becoming finely rib- 
bed on the Jast half; post-nuclear whorls spirally and axially 
sculptured; anal fasciole concave, smooth, with a strong spiral 
cord, just in front of the appressed suture; axial sculpture of 
about 8 sharp or pointed ribs on the periphery of each whorl, 
lacking entirely from the anal fasciole and from the base of the 
last whorl; spiral sculpture of 3 cords on the early spire-whorls 
becoming more numerous on the later, due to the intercalation 
of additional spirals; on the last whorl, the spirals number about 
22,including those on the canal; anterior canal long and straight; 
anal sinus shallow; outer lip thin. 

Length 47, diameter 18, last whorl, 32 mm. 
67 27 47 

This shell when full-grown appears distinct from the /avz- 
noides but young specimens cannot be easily separated. This 
variety differs from the typical form, by its sharp, narrow, and 
widely spaced axial ribs, which produce a sharp, angled and 
not rounded shoulder to the periphery of each whorl. The anai 


fasciole is also wider and more smooth. 


Gatun Stage: Toro Cays. Port Liman, coralline limestones. 


229 Costa Rica M1ockNE—OLSSON 57 


Turricula taurina, n. sp. Plate 5, figure 3 


Shell small, slender and fusiform; mucleus of nearly two 
smooth, convex whorls; post-nuclear whorls about seven; su- 
tures appressed and bordered by a spiral cord; fasciole about 
one-half the width of the spire-whorls, smooth; the sculpture of 
the whorls is spiral, axial ribs lacking; on the spire-whorls there 
are four raised spiral cords with the flattened interspaces about 
three times the width of the spirals themselves; on the last whorl 
and anterior canal, there are 18 or 19 spirals; anterior canal, 
long, slender and strongly twisted or curved; outer lip simple; 
anal sinus as indicated by the growth lines is wide, moderate- 
ly deep and occupies the middle of the fasciole. 


Length 25, diameter 8, last whorl 16 mm. 


This isa small and delicate species, characterized by its 
slender, fusiform outlines and lack of an axial sculpture. The 
spirals are narrow, raised cords, rather widely spaced. The 
single specimen was collected at Bocas del Toro. 


Gatun Stage: Bocas del Toro, Panama. 


Turricula Terryi, n. sp. Plate 5, figure 4 


Shell small, slender and fusiform, with a long, evenly taper- 
ing spire and slender, twisted anterior canal; nucleus of two, 
smooth, small whorls, followed by 8 sculptured post-nuclear; su- 
ture appressed, bordered by a low, but wide, spiral cord; upper 
half or more of each spire-whorl, flat or slightly concave and 
forms the fasciole; fasciole nearly smooth or sculptured with 
few, low, faint spirals; early spire-whorls strongly sculptured 
with 12 or 13 axial ribs which are most heavy on the periphery 
of each whorl; these ribs become progressively weaker and on the 
last whorl are only feebly developed on the shoulder or periph- 
ery; spiral sculpture persistant but weak, consisting on the spire- 
whorls of 4 cords on the lower half, alternating with much weak- 
er spirals: the fasciole as already noted is nearly smooth, or very 
weakly spiralled; on the last whorl, there are 20 spirals, includ- 


58 BULLETIN 39 230 


ing those on the anterior canal; aperture elliptical with a thin 
outer lip and long twisted anterior canal. 


Length 23.5, diameter 7, last whorl 15 mm. 


The small exposure of Gatun shales, on the east shore of 
Columbus Island, about one mile north of the city of Bocas del 
Toro, has yielded a large number of small interesting gastropods. 
More than a dozen species of Pleurotomids were collected at this 
place by Mr. R. A. Terry and the writer during a short period 
of collecting. 

The 7. faurina, also from this locality, is somewhat similiar 
but differs by its more convex whorls, deeper fasciole and prati- 
cally lacks all axial sculpturing. 

Gatun Stage: Bocas del Toro, Panama. 


Genus TURRIS Bolton 
Turris albida Perry Plate 4, figres I, 2 


FPleurotoma albida Perry, 1811, Conch. Expl., pl. 32, fig. 4. 
Fleurotoma virgo Wamarck, 1822, An. s. Vert., vol. 7, p. 94. 
Pleurotoma haitensis Sowerby, 1849, Quart. Jour. Geol. Soc. London, 


vol. 6, p. 50. 
Fleurotoma virgo Moore, 1853, Quart. Jour. Geol. Soc. Lond., vol. 9, 
Paso: 


Fleurotoma barretti Guppy, 1866, Quart. Jour. Geol. Soc. London, vol. 
22M ZOOMIPLe) V7 Aad Sa Os 

Pleurotoma antillarum Crosse, 1865, Journal de Conchyliologie, vol. 
13, p. 34, pl. I, fig. 8. Not of d’Orbigny. 

Turris (Surcula) virgo Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, 
Pp. 206, 207. 

Pleurotoma albida Dall, 1890, Trans. Wagner Free Inst. Sci., vol. 3, 
pt. 1, p. 28, pl. 4, figs.da. 

Pleurotoma albida Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. 
Phila., vol. 63, p. 343. 

Pleurotoma haitensis Cossmann, 1913, Jour. de Conchyliologie, vol. 
61, p. 16, pl. 2, figs. 1-4. 

Pleurotoma cf. antillarum Cossmann, 1913, Jour. de Conchyliologie, 
VOL OLN panei ple 2 wes s5 ho! 

Turris albida Dall, 1915, Bull. go, U. S. Nat. Museum, p. 38, pl. 5, fig. 
Tu DPV CA NATO NT) 


223i Costa Rico M1iocENE—OLSSON 59 


Turris albida Maury, 1917, Bull. Amer. Pal., vol. 5, p. 214, pl. 8, figs. 
4-8. 

This is a common species in the Gatun beds of Central Amer- 
ica. ‘The usual form is a large, strongly sculptured and should- 
er shell, which corresponds best to the variety described by 
Guppy from Jamaica as darreééz. Our largest specimen from 
Toro Cays, Panama measures as follows: Length 86, diameter 
27, last whorl 56 mm. 7. a/dida, is alsoa common fossil in the 
Miocene of Santo Domingo, Jamaica, Trinidad and Venezuela. 
Its geologlcal range is from the Oligocene to the recent. 

Gatun Stage: Gatun,C. Z. 

Toro Cay, Water Cay, Panama. 


Genus LEUCOSYRINX Dall 


Leucosyrinx chloris, n. sp. Plate 5, fignre 19 


Shell subfusiform, thin, with an acute spire and a long, 
straight, anterior canal; nucleus small, globular of about 2, 
smooth, convex whorls; post-nuclear whorls about 7; the spire- 
whorls are strongly angled in the middle, leaving a wide, con- 
cave or sloping zone about the upper suture; the sculpture con- 
sists of small, more or less confluent tubercles on the periphery 
of the whorls and number on the last whorl about 14; in addi- 
tion, the whole surface is strongly sculptured with raised, sub- 
equal, spiral threads; base strongly contracted tothe long, 
straight, anterior canal; the outer lip is broken; anal sinus, as 
indicated by the growth lines, lies in the concave zone, close to 
the upper suture. 

Length 14.50, diameter 5.00, aperture 7.25 mm. 

This appears to be a true Leucosyrinx, characterized by its 
thin shell, long, anterior canal and acute spire. It occurs in the 
Bocas shales with Dyrillia lithocolletotdes, cocostna, Tevrebra benth- 
alts var. bocasensts and others, shells which are all closely related 
to recent deep-water species. 


Gatun Stage: Bocas del Toro. 


60 BULLETIN 39 232 


Genus ANCISTROSYRINX Dall 
Ancistrosyrinx elegans Dall, variety Plate 4, figure 17 


Ancistrosyrinx elegans Dall, 1881, Bull. Museum Comp. Zool., vol. 
9, P. 54. 
Axcistrosyrinx elegans Dall, 1889, Bull. Museum Comp. Zool., vol- 
LSM PU 2hp A, Ous ple soul See 
The discovery of this elegant, recent species in the Gatun 
beds of Costa Rica is of more than usual interest, the record 
being based on a single, small but otherwise quite typical speci- 
men from Hill 1a of the Banana River. Comparison with Dall’s 
figure in the Blake Report, shows no important difference, ex- 
cept that the sculpture of the fossil shell is somewhat finer. 
A. elegans, is recorded by Dall from the Florida Reefs and 
from 805 fathoms of water off Havana, Cuba. 


The dimensions of our shell is as follows: Length 15, diam- 
eter 5.75, last whorl 11 mm. 


Gatun Stage: Hill ra, Banana River. 
Ancistrosyrinx Dalli, n. sp. Plate 4, figure 16 


Shell larger than the preceding and very nearly smooth; nu- 
cleus very small and smooth; post-nuclear whorls seven or more 
with a large and prominent, dentate or spinous keel; between 
the keel and the suture, there is a wide, nearly flat area, which 
carries a strong, smooth, median, spiral cord, behind which lies 
the small concave anal fasciole; the rest of the shell is smooth 
and without spirais;on the last whorl, the peripheral keel carries 
about 18 short, broad, spiny teeth; the canal is long, slender and 
straight; outer lip thin and fragile. 

Length 14, diameter 7, last whorl 9.5 mm. 
20 10, 15 mm. 


Two specimens are represented in our collection, and the 
smaller but more perfect specimen is selected as the Holotype. 


It differs from the described species of Axncistrosyrinx by its 


233 Costa Rica MioceENE—OLSSON 61 


nearly total lack of spiral sculpturing. The species is named 
for Dr. W. H. Dall of the National Museum, the author of the 
genus Azncistrosyrinx. 


Gatun Stage: Toro Cay, Water Cay, Panama. 


Genus DRILLIA Gray 
Drillia venusta, Sowerby Plate 4, figure 9 


Fleurotoma venusta Sowerby, 1849, Quart. Jour. Geol. Soc. London, 
VOl--65)p. 50; pl: Lo, fg. 7. 

LPleurotoma venustum, Guppy, 1866, Quart. Jour. Geol. Soc. London, 
VOls 22)" ps 280) 

Turris (Drillia) venusta, Gabb, 1873, Trans. Amer. Phil. Soc., vol. 
15, p. 208. 

fleurotoma venusta, Guppy, 1876, Quart. Jour. Geol. Soe. London, vol. 
Behe D 527): 

Fleurotoma venusta, Guppy and Dall, 1896, Proc. U. S. Nat. Museum, 
vol. 19, p. 305. 

Drillia venusta Maury, 1917. Bull. Amer. Pal., vol. 5, p. 216, pl. 8, 
BUSRS) AS TEA, 


The Drillia venusta and consors of Sowerby, are the two 
commonest Pleurotomids in the Gatun beds of Costa Rica. 
Both are characterized by a neat, regular and reticulate sculp- 
ture of spirals and axial ribs. In vwenusta, the anal fasciole is 
narrow, and appears asa deep constricted zone, encircling the 
upper part of each whorl. The spirals are narrow bands, pro- 
duced between deep, incised lines and in typical specimens num- 
ber on the last whorl about 18. Thecanal is rather long and 
usually more or less twisted. 


Drillia venusta, occurs also in the Miocene of Jamaica, 
Trinidad and in Santo Domingo. 


Gatun Stage: Gatun, C. Z. 
Banana River, Port Limon. 


Drillia consors, Sowerby Plate 4, figures 8, Io, 13 


Pleurotoma consors, Sowerby, 1849, Quart. Jour. Geol. Soc. Londen, 
WO, ©, {Os GOs 


62 BULLETIN 39 434 


FPleurotoma consora, Guppy, 1866, Quart. Jour. Geol. Soc. London, 
vol. 22, p. 280. 


Turris (Drillia) militaris, Gabb, 1873, Trans. Amer. Phil. Soc., vol. 
15, p. 207. Not of Hinds, 1843. 

fleurotoma consors, Guppy, 1876, Quart. Jour. Geol. Soe. London, vol. 
32, Pp. 527- 

Drithia militaris, Gabb, 1881, Jour. Acad. Nat. Sci. Phila., vol. 8, and 
series, p. 337. 

FPleurotoma alesidota, Dall, var. magina Bose, 1908, Bol. Inst. Geol de 
Mexico, No. 22, p. 47, pl. 5, figs. 30, 31, 33, 45, 

Pleurotama sp. aff. Pl. alesidota (Dall) var. mactlenta Toula, tort, 
Jahrb. der K-K Geol. Reichsanstalt Wiea,vol. 61, p. 505, pl, 30, fig. 
11. Not alesidota or macilenta Dall, 1889. 

Drillia consors Brown and Pilsbry, 1911, Proc. Acad Nat. Sci. Phila., 
vol. 63, p. 345. 

Drillia consors, Cossmann, 1913, Jour. de Conchyliologie, vol. 61, p. 
20, pl. 2, figs. 8-14. 


Drillia consors, Maury, 1917, Bull. Amer. Pal., vol. 5, p. 216, pl. 8, 
figs. 15, 16. 


This species is easily distinguished from the vezusta, by its 
more slender form, wide but shallow anal fasciole, sharper sculp- 
ture and straight, longer, anterior canal. ‘The most common 
variety, illustrated by figure 10, averages in length about 35 
mm. The axial ribs are numerous (about 28), straight or 
slightly oblique and only slightly heavier than the raised spiral 
cords. In addition the surface is overrun by fine and almost mi- 
croscopic spirals. Figure 8 of a large variety from Grape Point, 
has about 9 whorls and measures 60 mm in length. The early 
spire-whorls have the usual sculpture as seen on typical cozsors. 
On about the seventh whorl, the ribe increase suddenly to near- 
ly twice their normal number. They gradually become obsolete, 
so that on the last whorl, pratically only the spirals are left. 
The fine, microscopic spirals seen on typical consors are lack- 


ing. 
The Drillia alesidota and its variety macilenta Dall of the 
recent West Indian fauna, are closely related to consors. 


235 ~ Costa RrcA MrockenE—OLSSON 63 


Gatun Stage: Mt. Hope, C. Z. 
Banana River, Port Limon. 
Boras. 


Brillia cf. moensis, Gabb 


Drillia moensis, Gabb, 1881, Jour. Acad. Nat. Sci. Phila., vol. 8, 2nd 
SCHIES, py 3514 Pl. 46.) fe. 33. 


A single small speimen of a Drzllia was collected from the 
coral limestones near Limon, which agrees partly with Gabb’s 
figure and description. Our specimen has 7 instead of 8 ribs, 
and lacks the sutura! spiral mentioned by Gabb. The dimen- 
sions of this shell are: Length 12.5, diameter 4.5, last whorl 
7 mm. 


Gatun Stage: Port Limon. 
Drillia papaya, n. sp. Plate 4, figure 5 


Shell of medium size, slender and subfusiform; whorls 10 
(mucleus eroded); anal fasciole, a deep concave zone, occupying 
about 1-3 of a whorl interval and bordered above by a small su- 
taral cord; sculpture of heavy, knob-like ribs, crossed by spiral 
threads of different strengths; axial ribs about 6 on the last 
whorl, lacking from the fasciole and from the base and anterior 
canal; spiral sculpture of primary secondary and tertiary threads, 
there being about four primary threads on each spire-whorl, 
on the last whorl there are 8 or more primary threads, with an 
intermediate secondary and finer tertiary threads on each side; 
the anterior canal has numerous threads of varying strength; 
aperture long and subelliptical; anterior canal long and slightly 
bent backwards; anal sinus asindicated by the growth lines is 
deep and wide. 


Length 32, diameter ro, last whorl 19 mm. 
A single specinien was collected from certain sandstones of 
Lower Gatun age from a small tributary stream of the Rio 


Cocles near Old Harbor, C. R. The recent Drillia haliostrephis 
Dall from the Gulf of Mexico appears closely related, but the fos- 


64 BULLETIN 39 236 


sil shell has a more slender spire and slightly longer anterior 
canal. 


Gatun Stage: Rio Cocles, C. R. 
Drillia cocees, n. sp. Plate 5, figure 7 


Shell small, slender and fusiform; whorls Io or more, nu- 
cleus eroded; fasciole deep and concave, about % of the width of 
a spire-whorl, bordered above by a strong sutural cord, other- 
wise smooth; sculpture of strong axial ribs, which are lacking 
from the fasciole and from the base of the last whorl; these ribs 
number on the last whorl about 10; the ribs are crossed by 
strong, raised cord-like spirals, there being at first 2 on the earl- 
iest spire-whorls and on the later 3; the last whorl shows 8 
spirals and the wide interspaces are occupied by 3 or more small- 
er ones; additional and more crowded spirals occur on the long 
slender canal; anterior canal long and slender, slightly bent. 


Length 21, diameter 6, last whorl 12 mm. 


A small, very slender and fusoid species with strong, persis- 
tent spiral and axial sculpture. A single specimen was collect- 
ed at Coco Plum, a small cocoanut plantation about 40 miles east 
of Bocas del Toro. 


Gatun Stuge: Coco Plum. 


Drillia chiriquiensis, n. sp. Plate 5, figure 2 


Shell solid of medium size; whorls 7 (apex lost), of slightly 
convex profile; anal fasciole small but quite deep and nearly 
smooth; suture strongly appressed and bordered just below by a 
strong, elevated spiral cord; sculpture of numerous, somewhat 
oblique riblets and numerous weak spirals; the ribs commence 
at the lower edge of the fasciole, pass slightly obliquely across 
the whorls to the suture or on the last whorl across the base to 
the canal; the ribs on the last whorl number 15 or 16; spirals of 
numerous weak threads which are generally alernating in charac- 
ter; the spirals are very small on the fasciole, quite large and 


237 Costa RicA M1IocENE—OLSSON 65 


coarse on the anterior canal; the outer lip is heavily thickened 
by a large rib; anterior canal short and bent. 


Length 25, diameter 8, last whorl 15 mm. 


A single shell from Bocas, serves as type for this species. It 
recalls the D. consors Sowerby, but differs in its much finer 
sculpture. 


Gatun Sage: Bocas del Toro, Panama. 
Drillia citria, n. sp. Plate 5, fiyures-12, 13 


Shell small; whoris about 8, of which the first 2 belong to 
the small, obtuse, smooth nucleus; the post-nuclear whorls are 
strongly sculptured with ribe and spirals; the upper 1-3 of the 
spire-whorls carries a concave fasciole, bordered posteriorly by a 
strong sutural cord to the rst post-nuclear whorl; the fasciole is 
smooth or only faintly sculptured with fine spirals; the sculpture 
consists on the last whorl of about 9 ribs which fade out rapidly 
on the base and on the fasciole; the ribs are crossed on the early 
spire-whorls by 2 spiral cords, becoming 3 on the later; on the 
last whorl there are 12 more of which 6 are on the anterior ca- 
nal; the intervals between the spirals, are 3o0r more times as 
wide as the spirals themselves and are sculptured with fine 
spirals threads; aperture sub-elliptical, with a short and slightly 
twisted anterior canal; anal sinus moderate. 

Height t1, diameter 4 mm. 


A small species, common in the coralline phase of the Gatun 
at Port Limon. 


Gatun Stage: Port Limon. 


Drillia aquanica, n. sp. Plate 5, figures 16, 17 


Shell small, slender with a long spire and a shorter last 
whorl; whorls about 11, the nucleus composed of 2 small, smooth 
convex whorls and about 9 post-nuclear; whorls strongly sculp- 
tured with ribs and spiral cords; fasciole nearly 1-3 the width of 
the spire-whorls, marked with fine spirals and bordered postert- 


66 BULLETIN 39 238 


orly by a strong sutural cord; the spiral sculpture consists on 
the early whorls of 2, later 3 and finely 4 cords on the penulti- 
mate whorls; the last whorl has 8 spiral cords and 10 others on 
the canal; the spiral cords are separated by interspaces of slight- 
ly greater width; the interspaces are finely striated with small 
spiral threads; the axial sculpture consists of about 9 knot-like 
ribs over which pass the enlarged spiral cords; the ribs pass over 
the base of the last whorl to the canal and slightly over the fac- 
ciole; aperture elliptical with a short, straight canal. 


Height 18, diameter 6, last whorl 9 mm. 


This is a diminutive of Drillia fusiformis of the Miocene of 
Santo Domingo, the two species having practically the same 
sculpture. Drillia fusiformis of 11 whorls measures 35 mm in 
length, while agzanica of the same number of whorls hasa 
length of only 19 mm. The Maury collection contains a small 


but quite typical specimen of aguanica from the Rio Gurabo, 
Santo Domingo. 


Gatun Stage: Water Cay. 


Drillia limonica, n. sp. Plate 4, figure 15 


In general form like Drillia consors Sowerby, with a long 
spire and but slightly shorter anterior canal; whorls about 8% of 
which 1% belongs to the small, smooth, blunt nucleus; the 
post-nuclear whorls are sculptured with ribs and weaker spirals ; 
on the 1st three whorls, the anal fasciole is but slightly differ- 
entiated; it increases slowly in strength until on the later whorls 
it has become a concave zone asin comsors; a sutural cord is in- 
troduced on the 2nd post-nuclear whorl and is present on all the 
succeeding whorls; on the spire-whorls there are about 11, wide- 
ly spaced ribs; these ribs are lacking from the anal fasciole, ex- 
cept on the very earlest whorls, where they are faintly continued 
towards the upper sutures; on the last whorls, the ribs are small- 
er and number about 13; between these main ribe on the last 
whorl there appear occasionally smaller and shorter ribs which 
do not reach to the edge of the fasciole; there are 5 faint spirals 


239 Costa Rico MrockNE—OLSSON 67 


crossing the ribs on the spire-whorls and very much finer ones in 
the anal fasciole; the last whorl shows about ro spirals together 
with others on the canal; aperture elongate with the anterior 
canal rather long and straight. 


Length 19, diameter 6.5 mm. 


Like the Drillia consors in form, this species is distinguish- 
ed by its fewer ribs and shallower anal faciole. The ribs on the 
earlier whorls are oblique, but become very nearly straight and 
vertical on the later. 


Gatun Stage: Port Limon. 
Drillia bocatoroensis, n. sp. Plate 5, figure 6 


Shell of medium size, solid and strongly sculptured with 
ribs and spirals; whorls about 11, the nucleus eroded in the type 
specimen; fasciole occupying about 1-3 of the width of the spire- 
whorl, with a large sutural cord above which the edge of the su- 
ture is finely and regularly beaded or granulated; the surface of 
the fasciole is slightly undulated by the obsolete ends of the ribs, 
otherwise smooth; axial ribs about 8 on the last whorl, straight 
and nearly in line across the face of the spire-whorls to the apex, 
but interrupted and lacking from each fasciole; spiral sculpture 
of the few raised cords with wider interspaces; there are 3 
spirals on the spire-whorls, about 6 on the last, with 9 more on 
the anterior canal; the interspaces in addition carry fine and sub- 
microscopic spiral lines; the base of the last whorl is somewhat 
constricted with a short anterior canal. 


Length 23, diameter 8, last whorl 13.5 mm. 


From the several species of fusoid Pleurotomids in the Gatun 
beds of Panama and Costa Rica, this species differs in its heavy, 
persistent ribs and sharp spiral sculpturing. The fasciole is 
nearly smooth, bordered above by a heavy sutural cord. 


Gatun Stage: Bocas del Toro. 


68 BULLETIN 39 240 


Driffia theobroma, u. sp. Plate 5, figure 11 


Shell subfusiform; whorl 7 (apex broken); fasciole a narrow, 
concave zone, not strongly differentiated from the rest of the 
whorl, spirally sculptured and bordered above on the early spire- 
whorls by a sutural cord; sculpture consists on the spire-whorl 
of about ro low ribs, becoming more numerous and subobsolete 
on the last; the spiral sculpture consists of raised, alternating, 
revolving cords; there are 2 or 3 spiral cords on the early spire- 
whorl, becoming 5 or more on the later; the last whori has about 
10 spirals in addition to those on the anterior canal; between 
each pair of the main or primary spirals there is a smaller, sec- 
ondary spiral; base rounded or contracted tothe slender anterior 
canal. 


Length 21, diameter 7.5, last whorl 13 mm. 


It is possible that this species, belongs to the genus Glypho- 
stoma, as the type specimen seems to show a slightly thickened 
and inflated outer lip. The aperture is however so completely 
filled with a rocky matrix, that this observation cannot be fully 
verified. "The axial sculpture is practically lacking from the 
last whorl, leaving a surface simply marked with the rough 
spirals. 


Gatun Stage: Bocas del Toro. 
Drillia aurantia, n. sp. Plate 5, figure 15 


Shel! small, solid; whorls about 9; the nucleus consists of 
about 114 smooth, convex whorls which through the gradual in- 
troduction of the ribs, followed by the spirals blend in with the 
post-nuclear whorls; the anal fasciole is rather wide (about 1-3 
of the spire-whorl) and shallow; it uppears very early and car- 
ries from the start a strong, sutural cord; the sculpture consists 
of numerous closely spaced, linear ribs which are absent from 
the anal fasciole; these ribs number on the last whorl about 21; 
the spirals are uneven threads which are strongest below and on 
the base: the fasciole also carries one or more fine threads except 


24t Costa RicA MirocENE —OLSSON 69 


on the base of the last whorl; the spirals do not cross the ribs so 
that the summit of the ribs are left nearly smooth; aperture 
small with a short, slightly twisted anterior canal. 


Length 11, diameter 4 min. 


A small species sculptured like the D. elegans Conrad of 
the Chesapeake Miocene of Virginia, but with more numerous 
and more closely spaced ribs. 


Gatun Stage: Port Linon, 


Drillia musacina, n. sp. Plate 5, figures 27, 28 


Shell small, slender; the nucleus of about 2 whorls, the rst 
blunt, convex and smooth, followed by the znd which is strongly 
carinate; there are 7 post-nuclear whorls of slightly convex pro- 
file; the fasciole is a narrow band on which the ends of the ribs 
become strongly deflected to the left; the axial sculpture of about 
16, narrow ribs on the last whorl; the ribs are continuous across 
the spire-whorls to the upper sutures but become bent and 
strongly deflected on the fasciole; on the last whorl, the ribs are 
continued to the base of the canal; the spirals vary somewhat 
in strength but are usually subregular spiral bands of which there 
are 7 or 8 on the spire whorls and about 14 on the last whorl and 
the canal; the back of the last whorl is strongly humped; the 
outer lip was probably thin (broken in all our specimens); canal 
short, straight. 


Length 9, diameter 2.75, last whorl 5.00 mm. 
A small species abundant in the Gatun beds of the Banana 
River. 


Gatun Stage: Hill ra, Banana River. 
Hill 3, Banana River. 


Drillia estrellana, n. sp. Plate Io, figures 31, 32 


Shell very small, solid, porcellaneous; slender with a long 
spire and a short body-whorl; nucleus small of 2 smooth, convex 


70 BULLETIN 39 242 


whorls; post-nuclear whorls about 7; sculpture with strong ribs 
and impressed spiral lines in their interspaces; the anal fasciole 
is a constricted band about the upper 1-4 of the whorl and which 
passing over the ends of the ribs dislocates them asin Zerebra; 
there are 11 ribs on the last whorl which commencing at the up- 
per suture pass across the sutural fasciole but somewhat dimin- 
ished in strength, continue across the face of the whorl ina 
slightly oblique direction to the columellar region; the summits 
of the ribs are smooth but the interspaces are sculptured with 
impressed lines produced by spiral bands of regular width; there 
are 4 such bands on the spiral-whorls and about 12 0n the last 
whorl, including a few on the canal; aperture small, subellipti- 
cal and a short twisted canal. 


Height 7.5, diameter 1.75 mm. 
A small species of Terebroid appearance, the ends of the ribs 


being strongly dislocated as they pass across the small anal 
fasciole. 


Gatun Stage: Coll. 7, Estrella River. 
Drillia limonetta, n. sp. Plate 5, figure 10 


Shell small, solid, porcellaneous; whorls about 8, coarsely 
ribbed and polished; fasciole absent; the sculpture consists of 
moderately strong ribs which on the spire-whorls pass suture to 
suture but are only slightly flexed near the upper suture; on the 
last whorl, the ribs number about 11; the last rib is much en- 
larged and forms the outer lip; the ribs are but shortly continu- 
ed upon the base and do not reach the canal; in addition, the 
surface is covered with fine, irregular threads which are nearly 
obsolete on the upper portions of the whorls but are strong on 
the base and on the canal; aperture subelliptical with a short 
canal, somewhat bent or excavated ventrally; anal sinus simply 
as a vertically directed notch at the junction of the outer lip with 
the body-whorl and is bordered by a tooth-like denticle on its in- 
ner side. 

Length 12.25, diameter 5.00, last whorl 7.25 mm. 


243 Costa RicA MrocKkNE—OLSSON al it 


A small, solid and porcellaneous species, belonging to Dall’s 
Cymatosyrinx. The sutural fasciole being absent, the ribs are 
continuous to the upper suture and the surface is finely striated 
with small, irregular spiral threads. 


Gatun Stage: Port Limon. 
Drijlia carruca u. sp. Plate 4, figure I1 


Shell of medium. size, solid; whorls 7 plus (apex broken), 
rather rapidly tapering; no fasciole; axial sculpture of about 8, 
narrow ribs with deep and wider interspaces; the ribs commence 
at the top of the canal and pass across the whorls to the sutures; 
the ribs are nearly in line across the spire-whorls to the apex; 
spiral sculpture of slightly elevated cords, separated by wide, flat 
interspaces; there are about 7 spiral cords on the spire-whorl; 
with 12 on the last whorl; aperture sub-elliptical, the outer-lip 
somewhat thickened and a short canal. 


Height 18, diameter 7.25, last whorl 11 mm. 


This shell resembles Drz//ia musa in its general sculpture of 
fine, widely spaced, spiral cords. 


Gatun Stage: Coll.4, East Grate Potnt Creek. 
Driiiia limonensis, n. sp. Plate 5, figures 8, 4 


Shell of medium size and in form recalling D. venusta So- 
werby; nucleus of nearly three, smooth convex whorls; post-nu- 
clear whorls 8; anal fasciole scarcely perceptible on the carly 
whorls, becoming on the later, simply a narrow, constricted zone, 
bordering the suture; sculpture of rather numercus riblets, which 
are abruptly bent to the left near the posterior suture on crossing 
the constricted anal fasciole; three ribs number on the last whorl 
about 15; the spiral sculpture consists of very fine threads; the 
space between each pair of adjacent spiral threads is finely and mi- 
croscopically decussated by the raised lines of growth, producing 
a shagreening of the whole surface; in mature,shells, the last rib 
becomes unusually large and heavy, producing a large and strong- 


72 BULLETIN 3g 244 


ly thickened outer lip; canal short, slightly bent inward or out: 
wards; callous on the inner lip. 


Length 20, diameter 7.5, last whorl 11 mm. 


The general form of thissheill is that of Drillia venusta Sow- 
erby, which it also recalls in its constricted anal fasciole. The 
whole surface appears minutely shagreened, an effect produced by 
the fine, sub-microscopic, spiral threads as well as a minute de- 
cussation of their interspace. his species is fairly abundant in 
certain sandy layers found intercalated in the hard and gener- 
ally barren coral-reef limestone along the sea-front at Port 
Limon. 


Gatun Stage: Port Limon. 
Drillia Cristobali, n. sp. Plate 5, figure r 


Shell of the general form of wenusta, rather solid; whorls 6, 
(apex broken in the type specimen); anal fasciole a narrow con- 
stricted zone, bordering the upper suture; no sutural cord; sculp- 
ture of numerous, straight, parallel ribs and fine spirals; ribs on 
the early whorls about 12, becoming about I9 on the last; on the 
anal fasciole, the ribs are shortly deflected or bent; spirals very 
numerous and on slight magnification seen to consist of narrow, 
even raised threads with interspaces of nearly twice the width of 
the spirals themselves; anterior canal rather short and slightly 
bent backwards; aperture sub-ellipitical with a thin outer lip; 
anal sinus small and shallow. 


Length 20, diameter 8.5, last whorl 11.5 mm. 

This species, like the /Zémonensis, has the general form and 
constricted fasciole of Drillia venusta Sowerby, but differs by its 
smaller size, short canal and quite different spiral and axial 
sculpture. JD. limonensis, is more closely related, but that 
species differs in its more slender form and peculiar submicro- 
scopic sculpture. 

The single specimen serving as the type, was collected 
from the Gatun shale, exposed about a mile north of the city 
of Bocas del Toro, on Columbus or Cristobal Island. 


245 Costa Rica MrockNE—OLSSON 73 


Gatun Stage: Bocas del Toro, Panama. 


Drillia bocasensis, n. sp. Plate 5, figure 5 


Shell of medium size, solid; nucleus of about 2 smooth 
whorls; post-nuclear whorls about 10; anal fasciole a narrow con- 
stricted zone, nearly filled with a large, sutural cord; axial sculp- 
ture of large, heavy ribs, about 9 on each whorl; the ribs pass 
from suture to suture and on the last whorl acrcss the base to the 
anterior canal; a large, hump-like rib is developed on the back 
of the last whorl, formed during a resting stage; no spirals; the 
growth-lines produce a minute but characteristic sculpture of 
heavy, raised threads, which pass obliquely across the surface of 
the whorls and ribs, and follow the curve of the anal 
sinus across the fasciole; the sutural cord is finely granulat- 
ed by the raised growth-lines; canal short but straight; outer lip 
large, with a deep anal sinus at its posterior union with the 
body-whorl. 


Length 22, diameter 8, last whorl 12 mm. 


This species should more properly be referred to the genus 
Cymatosyrinx Dall, based on the Pleurotoma lunata H. C. Lea, 
a Chesapeake Miocene fossil. The form and general sculpturing 
of bocasensis is similiar to that of many species of this group but 
it may be recognized at once by its very peculiar, submicroscopic 
sculpture formed by the close, heavy growth-lines. 


Gatun Stage: Bocas del Toro. 
Drillia aquaensis, n. sp. Plate 5, figure 25 


Shell small and solid; nucleus of 1+ whorls (mostly broken 
on type specimen); post-nuclear whorls about 8; anal fasciole 
lacking; axial sculpture of about six, large, heavy and straight 
ribs, which pass from suture and across the base of the last whorl 
onto the anterior canal; each set of ribs is in a straight line 
from the canal across the spire-whorls to the apex; spiral sculp- 
ture of about 8 impressed lines which produce a series of flat but 


74. BULLETIN 39 246 


quite wide, spiral bands; on the last whorl there are about 22 
spiral bands; much finer near the suture, larger and wider on 
the middle of the whorl and on the canal; canal short and slight- 
ly bent to the left; inner lip calloused. 


Length 15, diameter 5.5, last whorl 8.5 mm. 
Characterized by its six axial ribs and finer sculpture of flat, 
spiral bands. It belongs to the genus Cymatosyrinx of Dall. 
Gatun Stage: Water Cay, Panama. 


Drillla musa, n. sp. | Plate 5, figure 26 


Shell about the same size as the preceding but witha 
more slender spire; nucleus of 2 smooth whorls (broken); post- 
nuclear whorls about 8; no fasciole; axial ribs about 9, pass 
from suture to suture and across the base to the anterior canal 
the ribs are in line across the whole series of spire-whorls; spirai 
sculpture quite heavy, consisting of raised threads with wider in- 
terspaces; the spirals number on the spire-whorls 8 or 9, and on 
the last whorl about 22; aperture large and suboval; anterior 
canal short, narrow and heavily calloused to form the inner lip. 


Length 17; diameter 6, last whorl 9 mm. 


Related to the preceding, but differs by its more slender 
form; with g instead of 6 ribs and a much coarser spiral sculp- 
ture. 


Gatun Stage: Banana River, C. R. 
Drillia lithocolletoides, n. sp. Plate 5, figure 18 


Shell small, glassy or translucent; nucleus of about 3 smooth, 
convex and glassy whorls, followed by 6 post-nuclear whorls; the 
whorls are strongly angled in the middle by tubercular-like ribs 
but leaving a concave zone about the upper sutures which forms 
the anal fasciole; on the last whorl, the ribs or tubercles number 
I1; are set obliquely and on the back of the whorl, they are con- 
tinued slightly over the base; the surface is smooth and polished; 
last whorl slightly constricted above the short, straight canal; 
aperture subovate. 


24.7 Costa Rico MioceNE—OLSSON 


“I 
wn 


Height 10.75, diameter 3.25 mm. 

The Drillia lithocolleta Watson is a_ recent deep-water 
species, dredged by the Challenger and the Blake, at several 
stations in the West Indiesand off the Florida coast, from depths 
of 400 to nearly tooo fathoms. ‘The Bocas shell seems to be 
very closely related to the recent species; differing only from 
Dall’s figure in the Blake Report (plate 11, fig. 61), in being 
somewhat more slender, with heavier tubercles and a longer base. 


Gatun Stage: Bocas del Toro. 


Drillia cocosina, n, sp. Plate 5, figure 14 


Shell small, glassy or translucent in texture; spire twice or 
more the length of the last whorl and the canal; whorls 8 plus, 
the earlier ones missing; no anal fasciole; the sculpture consists 
at first of a lower row of small tubercles, but a second or upper 
row soon begins to appear and on the last whorl, this upper set 
of tubercles is very nearly equal to the lower; the last whorl has 
about 13 of these tubercles, in addition the whorl is finely sculp- 
tured with fine, regular, closely spaced spiral threads which cov- 
er the entire whorl and the canal, but leaving the tops of the 
tubercles smooth; aperture ovate, with a short, twisted canal. 

Length 12, diameter 3.5 mm. 

This interesting species from the shales near the city of 
Bocas del Toro, is related to the recent Dril/ia oleacitna Dall 
dredged from rather deep water in the Gulf of Mexico and else- 
wherein the West Indies. In differs from that species in being 
less slender and with a larger body-whorl. 


Gatun Sage: Bocas del Toro. 


76 BULLETIN 39 248 


Genus GLYPHOSTOMA Gabb 
Glpyhostoma dentifera, Gabb. 


Glyphostoma dentifera Gabb, 1872, Proc. Acad. Nat. Sci. Phila., vol. 
24, p. 270, pl. 11, fig. 4. 
Glyphostoma dentifera Gabb, 1873, Trans. Amer. Phil. Soc. vol. 15, p. 


210. ; 
Glyphostoma dentifera Dall, 1889, Bull. Mus. Comp. Zool., vol. 18, 
pt. 2, p. 108. 


Glyphostoma dentifera Cossmann, 1913, Jour. de Conchyliologie, vol. 
61, p. 31, pl. 2, figs. 15, 16, 17. 

Glyphostoma deniifera Maury, 1917, Bull. Amer. Pal., vol. 5, p. 225, 
pl. 9, fig. 46. 

A single specimen of this species was collected from the cor- 
alline phase of the Gatun, near Port Limon. The shell showing 
5 whorls (apex lost), measures 18 by 8mm. The pecular sha- 
greening, characteristic of the genus is strong and easily seen. 

This shell agrees closely with the figure given by Cossmann, 
based on a Dominican specimen. It differs in several respects 
from the specimen figured by Dr. Maury in her Dominican Fos- 
sils from a metatype, sent by Professor Gabb to the Cornell Mu- 
seum. Dr. Maury’s shell is larger, has a longer spire, and the 
microscopic shagreening is much finer. 


Dall in the Blake Report, doubtfully refers to this species, 
an imperfect specimen collected in 15 fathoms of water at Sand 
Keys. 

Gatun Stage: Port Limon. 


Glyphostoma moinica, n. sp. Plate 5, figures 29, 30 


Shell small, solid with a coarse, tuberculate, subreticulate 
sculpture anda thickened, enlarged onter lip; nucleus small, 
pointed and tapering of 3 or more, smooth, convex whorls fol- 
lowed by about 4 post-nuclear; the sculpture consists of about 
'4 narrow ribs continuous from the canal to the upper suture 
and crossed by even, spiral cords; their intersection form small 


249 Costa RicA MrockNE—OLSSON 77 


rounded beds or tubercles; immediately after the nucleus, there 
is but one spiral, soon followed by two and on the penultimate 
there are three; the last whorl has five spirals and three or four 
more on the canal; aperture elliptical with a large expanded out- 
er lip, smooth within and a deep, anal sinus at its upper end; 
canal short and straight. 

Height 5.75, diameter 2.75, last whorl 3.75 mm. 


A small species doubtfully referred to Glyphostoma, having 
the outer lip smooth within and lacking the peculair, submicro- 
scopic structure of typical Glyphostoma. 

Its sculpture is coarse, the intersection of the ribs and spirals 
forming small beads or tubercles. 


Gatun Stage: Port Limon. 


Genus CYTHARA Schumacher 
Cythara terminufa var. costaricensis, n. var. Plate 5, figures 21, 22 


cf. Cythara terminula Dall, 1890, Trans. Wagner Free Inst. Sci., vol. 
BODE IGHP srs On Pls 2e tis. 5: 

The Cythara terminula Dall, to which the present shell ap- 
pears closely related, was described from the Caloosahatchie 
beds of Florida, of Pliocene age. The Costa Rican shell with 
the same number of whorls is somewhat small (typi- 
cal terminula of 7 whorls, 16 mim), (variety costartcensts 7 or 8 
whorls, 13 mm), and has 8 instead of 9 ribs. The details of the 
spiral sculpturing is very similiar, consisting of flattened bands 
separated by sharp channels. The spiral bands carry a central 
incised line, which produce the appearance of being in pairs. 

C. cercadica Maury from Santo Domingo, is a larger species 
with higher spire and simple and not banded spirals. 


Length 13, diameter 5, last whorl 10 mm. 
Gatun Stage: Fill ra, Banana River. 


Cytharelia limata, n. sp. Plate 5, figure 20 


Shell small, nearly smooth and porcellaneous; spire slightly 


78 BULLETIN 39 250 


shorter than the narrow aperture; nucleus of about 3% smooth, 
convex whorls, the last 4% turn being very finely and closely rib- 
bed; there are 3% post-nuclear whorls; sculpture consists of 
narrow, slightly oblique ribs which pass across the spire-whorls 
from suture to suture and on the last whorl follow down on the 
anterior canal to its tip; there are 8 ribs on the last whorl; the 
spaces between the ribs are wide, flat and smooth; the tops and 
sides of the ribs themselves are carved or etched with fine, sub- 
obsolete spiral lines; aperture linear-elliptical, the outer lip some- 
what thickened by the last rib, but smooth within; anterior ca- 
nal long straight. 


Height 9, diameter 3.50, last whorl 7mm. 


Gatun Stage: Port Limon. 


Genus BORSONIA Bellardi 
Borsonia coceensis, n. sp. Plate 5, figures 23, 24 


Shell small, biconic; whorls about 7; nucleus of 2, nearly 
smooth whorls, followed by 5 strongly shouldered post-nuclear 
whorls, the angle of which on the spire-whorls comes just 
above the lower suture; above the shoulder, the whorls 
are flat or slightly concave to the upper suture; sculp- 
ture with a double or triple beaded cord on the peri- 
phery and with alternating beaded spirals both above and below; 
on the last whorl, the primary spirals number 15 or 16, above 
the shoulder 3 or 4; the suture is bordered in front by a strong 
cord; pillar long, slightly twisted in front and with 2 sharp, pli- 
cations; the anal sinus lies on the peripheral angle as in typical 
Turris. 

Length 16, diameter &, last whorl 11 mm. 


The present species is based on two specimens fron: Coco 
Plum about 30 miles east of the city of Bocas del Toro. The 
B. varicosa Sowerby from the Dominican Miocene, is a related 
species, differing in its porportionately longer spire. 

Gatun Stage: Coco Plum. 


25% Costa RicA M10cENE—OLSSON 79 


Genus SCOBINELLA Conrad 
Scobinella Morierei (Laville) Cossmann Plate 4, figures 3, 4 
Euchilodon Morierei Vaville, in Cossmann, 1913, Jour. de Conchylio- 
logie, vol. 61, p. 34, pl. 3, figs 6, 7. 

Shell of medium size, with spire and last whorl of nearly 
equal lengths; sculpture of numerous, close, bead-like spiral 
cords, 3 or 4 on the spire-whorls, 18 or more on the last whorl 
and canal: the sutural fasciole is rather narrow, shallow and 
concave and with 4 much finer beaded cords; the suture is bor- 
dered in front by a heavy beaded spiral, which on the early spire- 
whorls, nearly fills the entire fasciole ; columella with four sharp 


folds, the largest above; canal nearly straight; aperture narrow, 
with a lirated outer lip. 


Length 49, diameter 15, last whorl 33 mm. 

This is one of the most elegantly sculptured Pleurotomoids 
in the Gatun beds of Panama and Costa Rica. Described by 
Laville and Cossmann from Mindi in the Canal Zone, we have in 
addition collected the species from Toro and Water Cay, (where 
it is fairly common), Bocas del Toro and from Limon. The pre- 
dominating spiral sculpture and columellar folds are very sug- 
gestive of Mitra. 


Gatun Stage: Mindi, C. Z. (Laville and Cossmann) - 
Toro aud Water Cays, Panama. 
Bocas del Toro, Panama. 
Port Limon, coral limestones. 


Genus HALIA Risso 


Halia americana, n. sp. Plate 4, figure 7 


Shell large, thin, buccinoid in form; spire elevated with 
shouldered whorls and deep sutures; whorls 3 plus, the earlier 
ones missing; the profile of the whorls is convex with a wide, 
shouldered, flattened or even slightly concave band just in front 
of the suture; the surface of the spire-whorls is smooth and un- 
sculptured but the early ones show very faint spiral bands; the 


80 BULLETIN 39 252 


upper half of the body-whorlis smooth but below, the whorl car- 
ries about 16, low, faint, spiral ribbons like those of J/alea but 
much less strong; these spiral ribbons average about 1.25 mm in 
width; the growth lines cross the face of the whorl from the tip 
of the canal to the shoulder angle in a slightly convex curve, the 
convex side of which is towards the aperture; from the shoulder 
angle to the suture the growth lines are reversed with the con- 
cave side facing the aperture; aperture broadly elliptical; the ca- 


nal was probably short and turned inward or ventrally. 
Height 66, diameter 39 mm. 

This truly remarkable species evidently belongs to the Pleu- 
rotomoid genus Hala, of which the 4 known species are Euro- 
pean and North African in their distribution. Its type species 
flalia Priamwzs Meuschen is living off the coast of Spain and 
North West Africa. According to Cossmann, there are three 
Tertiary speciesin Europe, . precedens Pant. from the Helve- 
tian of Italy, 7. Deshayeseana da Costa in the Tortonian of 
Portugal and Italy, and AH. helicotdes Br. in the Plaisancian of 
the maritime Alps. 


The Huropean species are described as being entirely smooth, 
but the Gatun shell is sculptured on its lower half with ribbon- 
like spirals bands. 


Gatun Stage Mi. Flope OZ 


Genus CANCELLARIA Lamarck 


Cancellaria dariena Toula Plate 6, figure 8 


Cancellaria dariena Toula, 1909, Jahrb. K-K. Geol. Reichsanstalt, 
Wien, vol. 58, p. 31, pl. 25, fig. 13, pl. 28, fig. 2. 

Cencellaria dariena Brown and Pilsbry, I911, Proc. Acad. Nat. Sci. 
Phila., vol. 63, p. 345, pl. 24, figs. 3, 4. 

Cancellaria darienesis Cossmann, 1913, Jour. de Conchyliologie, vol. 
61, p. 51, pl. 4 figs. 9, Io. 
var. trachyostraca Brown and Pilsbry. 

Cancellaria dariena, var. Toula, 1909, Jahrb. der K-K, Geol. Reich- 
sanstalt Wien, vol. 58, p. 37, pl. 28, fig. 11. 


253 CosTA RicA M10ocENE—-OLSSON 81 


Cancellaria dariena var. trachyostraca Brown and Pilsbry, 191t, Proc. 
Acad. Nat. Sci. Phila., vol. 63, p. 345, pl. 23, figs. I, 2. 


This species is most abundant in the Canal Zone. It is dis- 
tinguished from the Barretti Guppy, which it most closely re- 
sembles by its shorter spire, larger body-whorl and a more irreg- 
ular spiral sculpture. A large specimen from Mt. Hope, C. Z., 
has the following dimensions: Length 38, diameter 22; last whorl 
30 mm. Shorter and more globose shells form the variety ¢ra- 
chyostraca Brown and Pilsbry. 

Gatun Stage: Gatun and Mt. Hope, C. Z. 

Water and Toro Cays, Panama. 
Veto Gocles\ Gunes 


Cancellaria Barreitti Guppy Plate 6, figure 6 


Cancellaria Barretti Guppy, 1866, Quart. Jour. Geol. Soc. London, 
vol. 22, p. 289, pl. 17, fig. 11. 


Cancellaria reticulata Gabb, 1873, Trans, Amer. Phil. Soc., vol. 15, p. 
236. In part. 


Cancellaria Barretti Guppy, 1876, Quart. Jour. Geol. Soc. London, 
vol. 32, p. 520. 


Not C. Barretti Maury, which is C. Maury@, n. sp. 


This species is the West Indian Miocene analogue or the re- 
cent Cancellaria reticulata Linné. ‘The most important differ- 
ence to be noted between thetwo shells is that in Barretiz the 
columellar plicee are more anteriorly situated and heavier. It 
is a Miocene fossil in the Bowden beds of Jamaica. 


Gatun Stage: Banana River 


Cancellaria Cossmanni, n. sp. Plate 6, figures 9, II 


Shell of medium size, solid; nucleus of 3, small, smooth 
whorls; post-nuclear whorls 5, sutures deep; the whorls are 
slightly channeled or coronated just below the sutures; sculp- 
ture is evenly but coarsely reticulated by numerous, slightly ob- 
lique, narrow ribs, crossed by heavy, raised, narrow, spiral cords; 
the last whorl show 21 or 22 ribs; the spirals consists of narrow, 


82 BULLETIN 39 254 


raised cords, separated by wide, deep interspaces; there are 4 
spirals on the spire-whorls, with 15 on the last whorl and the 
canal; base of last whorl rounded and contracted to the short, 
beak-like anterior canal; columella with 2, narrow, but strong 
plications, of which the upper is much the strongest; aperture 
sub-elliptical with the outer lip carrying 8, narrow, entering 
liree. 
Length 25, diameter 14.5, last whorl 18 mm. 


This shell should probably be considered as a variety of C. 
Larretti Guppy, but in the large series in our collection, the shell 
is always much smaller and with a much coarser and heaver sculp- 
ture. In some specimens, this sculpture becomes quite sharp 
and harsh, brought about by the crossing of the narrow spiral 
cords and equally narrow axial ribs. 


It is an abundant and characteristic species of the Gatun beds 
along the Banana River, C. R. 


Gatun Stage: Banana River. 
Cancellaria Maurya, n. sp. Plate 6, figure 5 


Cancellaria Barretti Maury, 1917, Bull. Amer. Pal., vol. 5, p. 226, pl. 
10, fig. 1. Not of Guppy, 1866. 

Shell large, ovate; an evenly conic spire in height less than 
the length of the aperture and a large, evenly convex last whorl; 
the nucleus is rather small, of 2 smooth, convex whorls followed 
by about 6 post-nuclear whorls; the profile of the spire-whorls is 
slightly convex with deep, distinct suture; the sculpture is can- 
cellate or reticulate, the spirals and ribs of very nearly equal 
strength; of the axial ribs there are 40 or more on the whorl, 
crossed by 24 spiral cords; on the spire-whorls there are 5 spiral 
cords becoming -7 on the penultimate; the spirals are fairly regu- 
lar, separated by interspaces 1% times their width, in which 
smaller spirals may occasionally appear; faint indications of rest- 
ing marks show as crowding of the ribs or as smooth, slightly 
humped spaces; aperture large, ovate; a thin outer lip with 17, 
long entering lireze; columella with 2 simple plicee, of which the 


255 Costa Rico MrocENE—-OLSSON 83 


upper is much larger; a third is probably developed on the an- 
terior border of the canal; a well marked ridge about the base 
and a strong siphonal fasciole. 


Height 36, diameter 23, last whorl 29, aperture 24 mm. 


This shell was figured by Dr. Maury in her Dominican fos- 
silsas C. Barretti of Guppy, but it differs from that species in 
much shorter spire, more globose shell, larger aperture and by 
its columellar plicze. In the species closely related to the veéz- 
culata, such as Barreiti, Cossmanni and others, the columellar 
plicze are heavy and the posterior one is usually more or less 
bifid. In this species, the posterior plication is large, wide and 
thin, especially in the interior of the shell. 


The Costa Rican collection is limited to a single, imperfect 
specimen, so we have used as the type of this species, the origi- 
nal figured specimen of Dr. Maury from the Miocene of Santo 
Domingo. 


Gatun Stage: Water Cay. 


Cancellaria epistomifera Guppy 
Cancellaria Moorei Gabb. 1872, Trans. Amer. Phil. Soc., vol. 15, p. 
236. Not C. Moorei Guppy, 1866. 
Cancellaria epistomifera Guppy, 1876, Quart. Journ. Geol. Soc. Lon- 
don, vol. 32, p. 520, pl. 28, fig. 9. 
Cancellaria epistomifera Cossmann, 1913, Journ. de Conchyliologie, 
vol. 61, p. 53, pl. 4, figs. 5, 6. 
Cancellaria epistomifera Maury, 1917, Bull. Amer. Pal., vol. 5, p. 227, 
plo ess 35) 47 (Se 
For figures of this species, the reader is referred to Dr. 
Maury’s Dominican Fossils. We have collected a few, small 
aud imperfect specimens at Gatun in the Canal Zone. Small 
shells are easily confused with the common C. darzena, but in gen- 
eral the spire is shorter, and the spiral sculpture is heavier and 
more regular. At maturity, the outer lip develops below a 
peculiar and characteristic sulcus or pout, which is not seen on 
any of the associated species of Cancellaria. 


Gatun Stage: Gatun, C. Z. 


84 BULLETIN 39 256 


Cancellaria Rowelli Dall Plate 6, figure 7 


Cancellaria Rowelli Dall, 1898, Proc. Acad. Nat. Sci. Phila., vol. 19, 
Doze, DIO. fetn 
Cancellaria Rowelli Maury, 1917, Bull. Amer. Pal., vol. 5, p. 227, pl. 
16, fig. 2 
Two specimens of a Cancellaria which seem to be this spec- 
tes, were collected from the Gatun beds of East Grape Point 
Creek. ‘The axial ribs are numerous, oblique and somewhat ir- 
regular. The spaces between the ribs are finely spiralled with 
deep, incised lines, producing narrow and regular spiral bands. 
The columellar plications are large and more or less bifid. Shell 
measures: 
Length 30, diameter 18, last whorl 21 mm. 


The Cornell Museum contains, in the Gabb collection, a 
single specimen which is probably this species. It agrees with 
the Costa Rican shells. This species was described by Dall from 
the Miocene of the Rio Amina, Santo Domingo. 


Gatun Stage: Collection 2, East Grape Creek, C. R. 


Subgenus TRIGONOSTOMA Blainville 
Canceliaria toroensis, n. sp. Plate 6, figure 4 


Cancellaria ( Trigonostoma) afi. Cancellavia bullata Sowerby, Toula, 
1911, Jahrb. der K-K. Geol. Reichsanstalt, vol. 61, p. 504, pl. 30, 
fig. 10. 

Shell with 6 large, expanding whorls; the nucleus of 2% 
whorls is small, closely coiled and smooth; the succeeding 2 
post-nuclear whoris are small, convex and finely reticulated with 
6 spirals and fine ribs; the last two whorls are deeply channeled 
or excavated along the suture; this excavated band is smooth, 
except as crossed by oblique growth lines; the last whorl is 
somewhat contracted above and between this contracted zone and 
the sutural channel is a large, heavy, more or less bifid spirai 
cord. ‘This spiral cord is rendered coarsely and sharply nodulose 
by the axial sculpture; the sculpture consists of about 21, narrow, 
low, oblique riblets crossed by strong spiral cords; the spirals are 


257 Costa Rica MIOCENE —-OLSSON 85 


rendered slightly nodulose by the crossing of the longit udinal ribs 
the penultimate whorl shows 6 or 7 spirals and 14 or 15 on the 
last whorl; the contracted band around the upper part of the whorl 
is nearly free from spirals, but is bordered above as already not- 
ed by the large, bifid and strongly nodulose spiral cord; base 
deeply but narrowly umbilicate, spirally sculptured within; col- 
umella with 3, oblique plications; inner lip with a wide callus, 
finely pustulated; the outer lip is broken in the type. 


Length 29, diameter 20, last whorl 23 mm. 


This is the Miocene analogue of the recent West Coast C. 
tuberculosa Sowerby. It is probably the species figured by Toula 
from the Canal Zone and which he compared with C. dullata 
Sowerby. More recently Pilsbry and Johnson have described as 
C. tnsularis,a Trigonostoma from the Miocene of Santo Dom- 
ingo and which they consider as possibly identical with Toula’s 
Isthmian specimen. In their description they mention but 14 
ribs while toroenszs has 21 and they relate their species with the 
Chesapeake Miocene perspecttva Conrad and the Tampa depressa 
Dall, species which belong to an entirely different group. 

Gatun Stage: 

Toro Cays, Providence of Bocas del Toro, Panama. 


Cancellaria Plummeri, n. sp. Plate 6, figures 2, 3 


Shell thin and delicate, with large, loosely coiled whorls; 
the whorls are prominently shouldered and deeply channeled or 
excavated along their upper sutures; nucleus of three small, 
smooth whorls; post-nuclear whorls 4; the sculpture consists of 
very fine and delicate spiral threads or lines and three rows of 
slightly elevated spine-like elevations or small tubercles: the 
larger of these rows is found along the shoulder of the whorls 
bordering the excavated zone, a smaller on the middle and a 
small scarcely noticeable one below; on the penultimate whorl, 
only two rows show and on the upper spire-whorls but one; the 
umbilicus is deep, but narrow: the interior is concealed in the 
matrix. 


86 BULLETIN 39 258 


Height 34, diameter 26, last whorl 17, aperture 19 mm. 

This is a very distinctive species, characterized by its thin, 
delicate shell and fine sculpture. It may be compared with the 
recent 7. bullata Sowerby from the Pacific coast of Central 
America, which differs in its larger, more expansive body-whorl 
and coarser sculpture. The single specimen was collected from 
the lower part of the Gatun formation on Toro Cays, in the east- 
ern part of the Chiriqui Lagoon. 

It is named for Mr. Fred Plummer of the Royal Dutch Oil 
Company of Hague, Holland. 

Gatun Stage: Toro Cays. 


Subgenus APHERA H. and A. Adams 
Cancellaria islacolonis Maury Plate 6, figure 12 


Cancellaria tessellata Gabb, 1873, Trans. Amer. Phil. Soc. vol. 15, p, 
236. Not of Sowerby, 1832. 

Cancellaria islacolonis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 229. 
DI Noy Mies 2 haeaD) 

The C. zslacolonis was selected by Dr. Maury, as the guide 
fossil for the lower of her Miocene formations, the Cercado. It 
is a very rare fossil in Costa Rica and we have collected it only 
along Red Cliff Creek, in the eastern part of the Republic. 
There it occurs in the upper part of the Uscari and in the lower 
part of the Gatun. 

It is closly related to the recent West Coast C. ¢esselata 
Sowerby. 

Uscavi formation, Coll. 6, Red Cliff Creek. 

Gatun formation; Coll. 4, Red Cliff Creek. 


259 Costa RicA MrocENE—OLSSON 87 


(B) SUPER-FAMILY RHACHIGLOSSA 


Genus GLIVA Bruguiere 


Olives are among the most common fossils in the Miocene 
beds of Panama and Costa Rica. ‘The various species, however 
show so few distinguishing characters, that their identification 
becomes at times, both difficult and uncertain. The following 
key has been prepared showing the characters which have been 
most relied upon for their separation. ‘This key should be used 
in conjunction with the figures. 


A. Aperture narrow and of about the same width along its 
whole length. Inner lip finely and regularly crenulated 
above. 

B. Sutures deep or channelled, with the edge of the 
whorl projecting slightly above; spire of medium 
height, length 35 mm. 

O. sayana vat. immortua Pilsbry & Brown 

BB. Sutures not channelled and with the upper edge of 
the whorl beveled or appressed. 

C. Spire generally low, with concave profile and 
usually a projecting and globular nucleus. 
Length 35 mm. O. drevispira Gabb 

CC. Spire high or short, with conic sides; nu- 
cleus or protoconch not large or prominent. 

Di Spire’ high) jandiconie.) shell) wlansxe! 
Length 40 mm or more 
O. cylindrica Sowerby 
DD. Spire short, shell usually small. Length 
rarely over 30 mm. 
O. gatunensts Toula 
AA. Aperture wide and usually expanded and flaring in 
front; inner lip is not crenulated above. (Agaronia) 
B. Spire long, about 1-3 of the length of the shell; 


88 BULLETIN 39 261 
inner lip not strongly calloused. Lenght up to 
45 mm. O. mancinella, n. sp. 
BB. Spire shorter, about 1-4 of the length of the 
Shell; inner lip usually strongly callused. 
Length about 40 mm. 
O testacea vat. costaricensts, 1. var. 
Oliva cylindrica Sowerby Plate 7, figure 1 


Oliva cylindrica Sowerby, 1849, Quart Jour. Geol. Soc. Londou, vol. 
6, p. 45. 

Oliva reticularis Guppy, 1866, Quart. Journ. Geol. Soc. London, vol. 
22, p. 288,. Not of Lamarck. 

Oliva cylindrica Gabb, 1873, Trans. Amer. Phil. Soe., vol. 15, p. 


215. 

Oliva cylindrica Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 
BOY Os \ ESPXO) 

Oliva cylindrica Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, pt. 
6, p. 1583. 


Miva cylindrica Cossmann, 1913, Journ. de Conchyliologie, vol. 61, p. 
SP lenS iu tis Seu Dui at 

Oliva cylindrica Maury, Bull. Amer. Pal., vol. 5, p. 231, pl. 10, figs. 
TAN Aal, 

A single, large specimen from Costa Rica, is doubtfully re- 
ferred to this Dominican Miocene fossil. It differs from the more 
common 6revispira by its larger size and longer, more conic 
spire. This shell is also closely related to the recent O. avaneosa 
Lamarck of the West Coast. 


Length 47, diameter 19 mm. 
Gatun Stage: Upper Gatun beds, 1 mile south of shore, alton: 
Old Man Sam Creek, C. R. 


Oliva brevispira Gabb Plate 7, figures 2, 3, 4 


Oliva brevispira Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 
215. 

Oliva brevispira Maury, 1917, Bull. Amer. Pal., vol. 5 p. 232, pl. 10, 
figs. 16, 17. 


26k Costa Rica MroceNE—OLSSON 89 


variety Giraudi Cossmann 


Oliva Giraudi Cossmann, 1913, Journ. de Conchyliologie, vol. 61, p. 
56, pl. 5, figs. 4-8. 


A fairly common species in Cost Rica. Typically the shell is 


short; moderatly convex, a short projecting spire and concave 
spire-whorls. 


Unusually broad forms as illustrated by figure 4 may be sep- 
arated by the name of Girvaudz Cossmann. 
A large typical shell measures: 
Length 35, diameter 17 mm. 


Gatun Stage: Coll.6, Red Cliff Creek, C. R. 
fleadwaters of Middle Creek, C. R. 
Cols Grape Point CreckanGs ies 


Oliva gatunensis Toula Plate 7, figure 5 


Oliva gatunensis Toula, 1909, Jahrb. der K-K. Geol. Reicesanstaldt, 
vol, 58, p. 702, pl. 25, fig. 12. 


Oliva reticularis gatunensis Brown and Pilsbry, 1911, Proc. Acad. Nat. 
Sci. Phila., vol. 63, p. 348. 


Oliva gatunensis Cossmann, 1913, Jour. de Conchyliologie, vol. 61, p. 
58, pl. 5, figs. 9-12. 

A common species at Gatun. The largest specimen in our 
collection measures 28 mm. in length but Brown and Pilsbry men- 
tion shells of an length of 35-38 mm. Oltva gatunensis, resem- 
bles somewhat the drevirsozra of Gabb, but has a more even con- 
ic spire and lacks the projecting nuclear tip. The sutures are 
appressed. Brown and Pilsbry, place this shell as a variety of 
the larger, recent Oliva reticularis Lamarck. 


Gatun Stage: Mt. ope, | C.)Z, 


Oliva sayana var. immortua Pilsbry and Brown Plate 7, figs. 6, 7 


Oliva sayana var. immortua Pilsbry and Brown, 1917, Proc. Acad. 
Naty Set Phtlar on 3a ui ple 15, fganG: 


The Oliva sayana Ravenel (Oliva literata of most writers) 


go BULLETIN 3g 262 


is acommion recent species along the eastern coast of United 
States, becoming replaced in the West Indies by the Oliva reiz- 
cularis Laniarck. These two species are principally distinguish- 
ed from each other, in that the sutures of O. sayana are quite 
deeply channelled, above which projects the upper edge of the 
whorls while in reticularis the sutures are merely deep with 
rounded or beveled shell margins. 

O. sayana var. tmmortua was described by Pilsbry and 
Brown, from Gatun beds in the vicinity of Cartagena, Columbia. 
What appears to be the same shell is common in the exposures 
found along the lower part of the Banana River. Like the drev- 
estria it is rather darkly colored, but no trace of a pattern is pre- 
served. Our specimens averaged in length about 36 mm. The 
largest specimen measures 41 by 19.5 mm. 


Gatun Stage: Banana River, C. R. 
Oliva testacea Lamarck, var. costaricensis, n. var. Plate 7, figs. 12, 13 


Shell rather solid, with a moderately projecting spire, about 
1-4 or less of the total length of the shell; whorls about 5, the 
last, large, convex and widest about the middle; the spire-whorls 
appear as if mesially divided by an encircling weak callus about 
the lower halt; aperture broadest below, with the inner lip heav- 
ily calloused above near its junction with the outer; the inner lip 
is non-denticulated above, along its lower part with 3 or more, 
irregular oblique, plaits; a broad band, arises from about the mid- 
die of the inner lip, passes obliquely downward over the back of 
the shell to the lower end of the outer lip. 

Length 42, diameter I9 mm. 

‘This Oliva is abundant in the Gatun beds of the Banana 
River, C. R. It is closely related to the recent O. festacea Lam- 
arex but seems to differ in being wider, shorter and with a lower 
spire. 

The Oliva testacea Lamarck (in part O. hiatula of some au- 
thors) is abundant along the Pacific coast of Panama, and in the 
Manuals, its range is given as the West Coast. Recently a close- 


263 Costa Rrco MiocENE—OLSSON 91 


ly related if not identical form, was collected by us from the 
Pleistocene at Almirante, Panama, and several broken specimens 
from the beaches at Bocas del Toro, Panama, and from Manzan- 
illa, C. R. This recent Atlantic form differs only from the typ- 
ical West Coast /estacca in being somewhat more slender and ap- 
parently differently colored. 


Tryon in the Manual of Conchology unités /estacea with 
hiatula Gmelin from West Africa. 

Gain Stage: Banana River, Coll 520 of, \Redp- Clif, 
(Cgaliox (ON IKE 


Oliva mancinella, n. sp. Plate 7, figures 8, 9 


Shell slender and more delicate than the preceding; spire 
long and pointed and about 1-3 of the total length of the shell; 5 
whorls; sutures deep; the spire-whorls have their lower half 
thickened by a smooth, encircling band of callus; aperture broad- 
est about the lower half; inner lip smooth above, with only a 
small and weak callus at its junction with the outer lip; the out- 
er lip carries below, 3 or 4 heavy and very oblique plaite; the 
outer lip is smooth and sharp. 


Length 48, diameter 11 mm. 
This species differs from O. ¢estacea and its variety costaricen- 


sts by its much more slender and delicate shell. The porpor- 
tions of the spire are about 1-3 of the total length of the shell. 


Gatun Stage: Coll. 4, East Grape Potnt= Creek, C. R. 


Genus OLIVELLA Swainson 


A. Shelllarge, length 20 mm. or more; spire long and about 
¥% the total length of the shell. 
Olivella goliath, n. sp. 
AA. Sheli smaller; spire long or short, usually less than % 
of the total length. 
B. Shell short and stubby, spire rather short, 


92 BULLETIN 39 2604 


bluntly pointed; 1-3 or less the total length. 
Olivella limonensts and variety bocas- 
énsts, 1. sp. and var. 

BB. Shell with a longer spire, oftensharply point- 
ed, 

C. Shell rather broad, with a heavy cal- 
lus about the upper part of the inner 
lip and extending partly over the pen- 
ultimate whorl. Length 12 mm. plus. 

Oltvella muticotdes Gabb. 

CC. Shell slender, with a long, pointed 
spire; callus about the upper of the 
inner lip, slight. 

Olivella Boussact Cossmann, var. 

Gliveila goliath, n. sp. Plate 7, figures 22, 23 


Shell large, with a long, pointed spire of about % of the to- 
tal length of the shell; whorls about 6, with plain, nearly straight 
profile; sutures linear; last whorl with the greatest convexity 
about the middie; aperture subelliptical, broadest in front; in- 
ner lip with a thin, flat callus and with 3 small plaits below. 


Length 26, diameter 10, last whorl 20, spire 13 mm 
23 8.5 18 A roaven 


An unusually large species, represented by two specimens 
from Red Cliff Creek and Banana River. Its general form is that 
of O. indevisa Guppy and O. Boussaci Cossmann, but nearly 4 
times as large. 


Gatun Stage: Banana River. 


Call ia wkied | Olapm@necton C.F. 
Olivella muticoides Gabb Plate 7, figures 11, 14, 17, 18 


Oliva muticoides Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 
Ais 

Olivella mutica variety muticoides Dall, 1890, Tarns. Waguer Free, 
Inst. Sci., vol. 3, pt. I, p. 45- 


265 Costa RicA MioceENE—OLSSON 93 


Olivella muticoides Maury, 1917, Bull. Amer. Pal., vol. 5, p. 232, pl. 
TATe io Se ATs 
A. broad, chubby species with a spire of moderate length. A 
large, thick callus is formed about the upper part of the inner 
lip, which spreads partly over the back of the penultimate whorl, 
giving to it a hump-back appearance. Measurements of our 
Costa Rican shell run as follows: 


Length 17, diameter 7, last whorl 14, spire 6.5 mm. 
13 6.5 II 5-5 mm. 
Gatun Stage: Coll. 5, Red Cliff Creek. 
Coll. 4, East Grape Point Creek. 


Olivetla Boussaci Cossmann, variety Plate 7, figures 15, 16 


Olivella Boussact Cossmann, 1913, Journ. de Conchyliologie, vol. 61, 
p. 60, pl. 5, figs. 16-19. 


The Olivella tndivisa Guppy (Proc. U.S. Nat. Mus., vol. 
19, p. 308, pl. 30, fig. 10) from Jamaica and Olivella Boussact 
Cossmann from Martinique, belong to a closely related series 
characterized by their high and sharply pointed spire. The 
main difference seems to be that of size. The zzdivisa hasa 
length of 6.5 mm., the Boussaci of 9 mm. The Costa Rican shells 
are generally larger and when full-grown, often 13 or more mm. 
in length. 


Often very common. 
Length 13, diameter 5, last whorl 10, spire 6.5 mm (6 
whorls) 


Gatun Stage: Hill ra, Banana River. 
fiill 3, Banana River. 


Olivella limonensis, n. sp. Plate 7, figures 19, 20 


Shell short and plump, witha small, broad, conic spire; 
whorls about 5, separated by deep sutures; last whorl very large, 
broadly cylindrical in form and slightly contracted in the middle; 
the inner lip has a wide callus, somewhat heavier above, with 5 


94 _ BULLETIN 39 266 


fine, denticles in the middle and few, more oblique plaits below; 
the base is obliquely encircled by a lighter colored band; outer 
lip sharp, smooth within. 
Length 1o.5, diameter 4.75, last whorl 9, spire 2.25 mm. 

This is a common shell in the coralline phase of the Gatun 
at Port Limon. The middle of the last whorl is usually dark col- 
ored, with the spire and the encircling basal band and callus on 
the inner lip white. 

Gatun Stage: Port Limon. 


Olivelia limonensis var.bocasensis, n. var. Plate 7, figures 24, 25 


A much smaller shell than the /monenszs and possibly a dis- 
tinct species. The spire is somewhat higher and the shell less 
cylindrical and pump. Dimensions as follows: 


Length 6.5, diameter 2.75, last whorl 6, spire 1.5 mm. 
ah BuDi5e 6.5 1.75 mm. 
Gatun Stage: Bocas del Toro, Panama. 


Genus ANCILLARIA Lamarck 
Ancillaria aquaensis, n. sp. Plate 7, figure Io 


Shell of medium size, solid; spire elevated, composed of about 
5 whorls, the sutures of which are concealed by a wide band of 
callus this band of callus commences on the body-whorl, a 
short distance below the suture, and extends to the apex; where 
this band covers the sutures, it forms just above, a depressed or 
constricted band on the middle of each whorl; the last whorl is 
large, with a basal band of callus, which commences near the 
upper end of the inner lip, descends obliquely across the back of 
the last whorl to the anterior tips of the outer and inner lips; a 
short distance above this basal band, the shell carries a single, 
incised line; a small umbilical pit is found just behind the mid- 
dle part of the inner lip; aperture broadly sub-elliptica] with a 
sharp outer lip. 


Length 18.5, diameter 7.5, last whorl 12.5, spire Io mm. 


267 Costa Rica M1ocENE—OLSSON 95 


The Ancillaria pinguis Guppy, from Jamaica is probably a 
closely related species. Guppy’s figure in the Geological Maga- 
zine, vol. 1, Decade 2, isvery poor and insufficient. Our shell 
seems to differ by its much longer spire and more slender shell. 


The Ancillaria chipolana Dall, figured on Plate 41, fig. 3, of 
the Wagner Institute, is larger and has a longer spire. No um- 
bilical pit is shown in the figure. 


Gatun Stage: Water Cay, (sla de Aqua). 


Genus MARGINELLA Lamarck 
Marginella MacDonaldi Dall Plate 6, figures 14, I5, 17, 19 


Marginella MacDonaldi Dall, 1912, Smith, Misc. Coll., vol. 59, No. 
2aRD STE 
This large M/arginella is one of the most common and char- 
acteristic species of the Gatun beds of Costa Rica. It is extreme- 
ly valiable in size and general from and heaviness of its shell, as 
may be seen in the accompanying figures of the more common 
varieties. 


Typically the shell is oblong-cylindrical with a small spire of 
about 4 whorls, usually completely covered in front by a broad 
mass of callus, which spreads over the whole base of the shell 
and the outer lip, leaving an elevated ridge along each side as 
frequently seen in Cyprga, From the back, the spire may be 
seen lying in the mass of callus. 


The Marginella mindiensis Cossmann is a smaller related 
species from the Canal Zone. It differs also in having its outer 
lip finely denticulated, while in /acDonaldz the outer lip is us- 
ually smooth. 


Length 28, diameter of base 15, vertical diameter 10 mm. 


24 14 g mim. 
25 13 9.5 mm. 
ola) II Gjois\ioab ol. 


Gatun Stage: Banana River. 


96 BULLETIN 39 268 


Marginella mindiensis Cossmann Plate 6, figures 16, 23 


Marginella mindiensis Cossmann, 1913, Journ. de Conchyliologie, vol. 
6f, p. 61, pl. 5, figs. 13-15. 

This shell is very closely related to MW. MacDonaldi. It dif- 
fers only in being smaller and usually with the outer lip more or 
less denticulated. The outer lip of WZ. MacDonaldi is generally 
smooth but occasional specimens occur in which the outer lip is 
faintly denticulated. It isa species of the Canal Zone. 


Length 19.5; basal diameter 11, vertical diameter 8 mm. 
16 9 7 mim. 
Gatun Stage: Mindi, (Cossmann) 


Gatun Gok 


Marginella latissima Dall 
Marginella latissima Dall, 1896, Prac. U. S. Nat. Mus., vol. 19, p. 308, 
pli2o hos 
This is a short and very broad species, described by Dall 
from Moen Hill, C. R. Dall remarks:‘‘This is perhaps the short- 
est and widest American species.’’ 


Length 11, diameter 8.5 mm. 
Gatun Stage: Moen, Costa Rica, (Gabb). 


Marginella latissima, var. pilstryi, mn. var. Plate 10, figures 1, 2 


Marginella coniformis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sct. 
Phila., vol. 63, p. 348, pl. 24, fig. 12. Not of Sowerby, 1849. 

Like M7. latissima Dall, but less broad, more pointed anteri- 
orly, a slightly higher spire and a less heavy outer lip. In /afzssv- 
ma, the two posterior plications are shown as being very oblique, 
while in the present form they are nearly transverse as is seen in 
the recent cizcta. ‘The outer lip is finely granulated. 


Length 14, basal diameter 9.5, vertical diameter 6.75 mm. 
Us II 9g mim. 


This shell is figured asthe Marginella contformis Sowerby 
a common Dominican fossil, by Brown and Pilsbry. The A/arg- 


269 Costa Rica’ MiockENE—OLSSON 97 


inella coniformis has been figured by Guppy in the Quarterly 
Journal and similiar shells were collected in abundance by the 
Maury expedition to Santo Domingo. It isa larger and more 
cylindrical form. 


It isan abundant shell in the quarries west of the Gatun 
locks at Gatun. A single, large shell figured as figure 1 was 
collected from the Banana River beds. 


Gatun Stage: Gatun, C. Z. 
Banana River, C. R. 


Marginella avena Valenciennes © Plate 7, figures, 21, 28 


Marginellaavena Valenciennes, 1814, in Kiener, Coq. Viv. Marginella, 
PAL Zip le Oy ao 2Ay 

Marginella avena Gabb, 1881, Journ. Acad, Nat. Sci. Phila., vol. 8, 
2ud series, p. 355. 

A common, recent species along the Caribbean coast of 
Panama and Costa Rica. As a fossil, it occurs plentifully in the 
coralline phase of the Gatun at Port Limon and Bocas del Toro. 
The fossil shells do not differ materially from their recent repre- 
sentative. 


The shell is elongate-cylindrical in outline, with a low, but 
slightly projecting spire. The aperture is norrowly linear, 
slightly wider in front. Outer lip thick, smooth within. The 
columella is provided with 4, obliquely descending plications. 

A series of specimens from Limon and Bocas measure as fol- 
lows. 

Length 11.75, diameter 4.25, spire 2. 
I1.50 4.25 1.15, 
12.50 4.50 a, 

Gatun Stage: Port Limon, C. R. 

Bocas del Toro, Panama. 


Coll. 4, Red Cliff Greek. 


Marginalla collina, n. sp. Plate 7, figures 26, 27 


Shell of the general form and size of JZ. avena, but more 


98 BULLETIN 39 270 


solid and with a lower and less differentiated spire; whorls 4 or 
more with sutures scarcely distinguishable under their glaze or 
coat of callus; aperture linear-elongate, widest in front and 
slightly contracted in the middle; outer lip thickened, smooth 
within; pillar with 4, oblique plaits of nearly equal strength. 


Length 11, diameter 4.5, spire 1.5 mm. 
TO 4.25 I mm. 
Ein 25 5. I mim. 


Closely related to the MWarginella avena Val., this species 
differs in being more solid, less slender and with a lower spire. 
In avena the spire is clearly differentiated, with well marked su- 
tures to its spire-whorls. In the present shell, the spire is much 
lower, and the sutures are more thickly covered with glaze. The 
thick outer lip is carried across the end of the lip and firmly 
joined to the spire without any preciptible line of demarcation. 
In avena, the outer lip and spire are clearly differentiated from 
each other. 


Abundant in the Gatun beds of the Banana River. 
Gatun Stage: Hill ra, Banana River. 


Marginelia leander Brown and Pilsbry Plate 6, figure 22 


Marginella leander Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. 
Phila., vol. 63, p. 347, pl. 24, fig. 13. 

Not known from Costa Rica, and the figure here given is 
from a specimen in our collection from Gatun. It is a broader 
and more cylindrical species than colliza, and with a much low- 
er and scarcely elevated spire. Our specimen measures 9 by 4 
mm. 

Gatun Stage: Gatun, C. R. 


Marginella musacina, n. sp. Plate 10, figure 25 


Shell small and slender, with a high spire of about 1-4 of 
the total length of the shell; whorls about 5, with sutures light- 
ly concealed under a thin coat of glaze; last whorl narrowly cylin- 


271 Costa Rico MiocENE—OLSSON 99 


drical, widest about the upper half; aperture sublinear, with a 

slightly thickened lip, somewhat contracted in the middle, smooth 

within; inner lip smooth, or with only a thin wash of callus and 

with 4, very oblique plications at its extreme anterior end. 
Length 9.25, diameter 3.25, spire 2.75 mm. 

A small, slender species of unusual form. Two specimens 
were collected in.Costa Rica, the type from the Banana River, 
the other from Old Man Sam Creek, near Manzanilla Point. 

Gatux Stage: Banana River, 

Along Old Man Sam Creek, one mile south 
of the beach, C. R. 


Genus VO@LUTA Linnzeus 
Voluta alfaroi Dall Plate 8, figure 2 


Voluta alfarvoit 1912, Smith, Misc. Coll., vol. 59, No. 2, p. 8. 


This fine species described by Dr. Dall, from the Banana 
River, isa common and very characteristic fossil or the Gatun 
beds in Costa Rica and Western Panama, but it still remains to 
be recorded from the Canal Zone. Allied to the recent West 
Indian V. musica Linneeus it differs most importantly in its 
much smaller nucleus. 


The shell is heavy and in its typical form the whorls are 
shouldered and carry about 12 ribs, which may be quite sharp 
and high on the shoulder angle. The whorls are sometimes 
smooth, without ribs and a shoulded angle. Spiral threads usu- 
ally occur on the lower one-quarter of the last whorl and on the 
canal, but in some cases on the spire-whorls as well. A large 
specimen from Water Cay measures; 


Height 59 mm. diameter 34 mm. 


Gaiun Stage: Water Cay Panama. 
Coll. 3, Red Cliff Creek; Old Man Sam Creek, 
r mile from shore, Coll. 2, QOuttana Creek; 
Comadre Creck, Soust creek; Coll. 5, Estrella 
River; Banana River, Rio Blanco, Port Limon. 


100 BULLETIN 39 272 


Genus SCAPHELLA Swainson 
Scaphella costaricana,n. sp. Plate 15, figure 13 


Type fragmentary consisting of the nucleus and part of the 
three succeeding whorls; the sheil is subfusiform; nucleus large, 
smooth and mamumilate at its apex; the three following whorls 
are slightly contracted about the upper sutures and strongly sculp- 
tured with coarse subregular, spiral threads separated by inter- 
spaces as wide or a little more; the spiral threads and their in- 
terspaces are crossed by fine lines of growth; sutures distinct; the 
columella with four plaits, the posterior one being the strong- 
est. 


Length 29, diameter 18, diameter of nucleus 5.75 mm. 


The unique type is unfortunately fragmentary and consists 
of the large, smooth, mammilate nucleus and part of the 3 suc- 
ceeding whorls. ‘The shell is subfusiform and the whole surface 
is strongly sculptured with coarse, spiral threads. The columel- 
la is provided with 4 strong plaits. The type specimen was found 
in the Dentalium zone of the Upper Uscari shales of Cocles 
Creek? 


Uscart Stage: Cocles Creek. 


Genus MITRA Lamarck 
Mitra Swainsoni Broderip, var. fimonensis, n. var. Plate 6, fig. I. 


ef. Witra Swainsont Roderip, 1835, Proc. Zool. Soc., p. 193. 

ef. Mitra Swainsoni Reeve, Conch. Icon., (Vitra, pl. 1, fig. 4. 

ef. Mitra Swainsont, var. antillensis Dall, 1589, Bull. Mus. Comp. 

Zool., vol. 18, p. 158, pl. 38, fig. 7. 

Shell large, of about 6+ whorls; the spire whorls are slight- 
ly convex, the last distinctly shouldered above, depressed or 
slightly concave about its middle; sculpture consisting on the 
spire-whorls of about 6, heavy, spirals cords with sculptured 
interspace as seen on Mitra Henekeni and longa, on the later 
whorls the spiral cords become more widely spaced and on the 


273 Costa Rica MrocENE—OLSSON IOI 


last are in the form of broad, subobsolete, smooth bands without 
the sculptured interspaces; there are about 15 spiral cords on 
the last whorl in addition to those of the anterior canal; anterior 
canal of moderate length, strongly twisted; columella with 4 
plaits, heaviest above; aperture narrow. 


Length 74 (apex broken), diameter 23, last whorl 52, 
spire 35 mm. 

A single, large shell with rudely sculptured whorls was col- 
lected from the coralline limestone near Port Limon. It agrees 
closely with Reeve’s figure of A%ttra Swainsont, except that the 
spiral bands are larger and heavier. 


The Mittra Swatnsont is a West Coast shell belonging to the 
Panamic province. It is distinguished by its large size, rudely 
sculptured whorls and dark colored epidermis. Dall has des- 
cribed as the variety azftillensis, specimens dredged off of Cape 
Lookout, N. C., Colombia and Yucatan. 

Gatun Stage: Port Limon. 


Mitra longa Gabb Plate 6, figure ro 


Mitra longa Gabb, 1873. Trans. Amer. Phil. Soc., vol. £5, p. 219. 
Mitra longa Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., vol. 
Osh pa346. pls 240 hig. Ta. 
Mitra longa Maury, 1917, Bull. Amer. Pal., vol. 5, p. 238, pl. 11, figs. 
TATA 
A long, slender species, sculptured with strong, revolving 
cords and fine, neatly engraved interspaces. We have collected 
this Dominican species only inthe Canal Zone. The figured spec- 
imen has the following dimensions: 


Length 4o, diameter 10.5, last whorl 26, spire 21 mm. 
Gatun Stage: Gatun, C. Z. 


Mitra dariensis Brown ond Pilsbry Plate 6, figure 25 


Mitra dariensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., 
vol. 60, p. 346, pl. 24, fig. 11. 


102 BULLETIN 39 274 


A single specimen from the Island of Bocas (Columbus or 
Colon Island) is here figured. It differs from the W/tra longa, 
in being shorter and broader. ‘The sculpture is less elegant. 


Length 21, diameter 7, last whorl 14, spire 11 mm. 


Gatun Stage: Gatun, C. Z. 
Bocas del Toro, Panama. 


Mitra aff. rudis Gabb | Plate 6, figure 13 


Mitra rudis Gabb, 1873, Trans. Amer, Phil. Soc., vol. 15, p. 220. 


Shell subfusiform, biconic; whorls about 7, coarsely sculp- 
tured with strong, revolving cords, separated by interspaces of 
about twice their width; These interspaces are finely longitudi- 
nally sculptured with raised threads above, obsoletely so below; 
the spire-whorls have 5 cords, the last whorl with 11 and several 
smaller ones on the canal; aperture subelliptical, with a thin out- 
er lip; columella with 4 plaits, largest above; anterior canal of 
medium length, strongly twisted below. 


Length 33.5, diameter 12.5, last whorl 24, spire 16 mm. 


It is with much uncertainty, that I have identified this rare 
species with Gabb’s unfigured Mitra rudis from Santo Domingo. 
Gabb,s descriptions and measurements indicate a shell of much 
the same characters as well as dimensions. 


Gatun Stage: Coll. 3, Hone Walk Creek, C. R. 
Mitra poas, n. sp. Plate 6, figures 20, 21 


Shell small, nearly smooth and columbelloid in appearance; 
whorls 8, with straight sides; last whorl widest just above the 
base, which is contracted to the short anterior canal; the early 
spire-whorls have 5, low, smoothish spiral bands, with longitudi- 
nally sculptured, narrow interspaces; these spirals quickly be- 
come obsolete, leaving the whorls smooth except for one or two 
spirals bordering the upper sutures; the last whorl has about 4 
spirals around the base and smaller ones on the canal; aperture 
subellipical, with a thin outer lip; columella with 4 plaits, larg- 


275 Costa RicA M1ocENE—OLSSON 103 


est above; anterior canal of medium length and slightly bent to 
the left 


Length 21, diameter 7.5, last whorl 13, spire 11 mm. 


A smooth species of Columbelloid aspect. It is fairly abund- 
ant in the coral limestones near Port Limon. 


Gatun Stage: Port Limon, 


Mitra Almagrensis Toula var. coralliophila, n. var. Plate 6, figures 18, 24 


of Mitra Almagrensis Toula, 1911, Jahrb, der K-K Geol. Reichsanstalt, 
VOl VOID. 40m ple 23, fei 13: 

Shell small with a long spire and a shorter anterior canal; 
whorls about 8, with straight or slightly convex profile; sculp- 
ture of low, slightly elevated spiral cords, separated by inter- 
spaces of about their own width; these interspaces are finely 
sculptured by regular, raised, longitudinal threads; the spire- 
whorls have about 5 spiral cords, the last whorl with about 15; 
and smaller ones on the anterior canal; aperture subelliptical; a 
short canal, slightly bent to the left; pillar with 3 folds and a 
very small faint one below. 


Length 17, diameter 6, last whorl 11, spire 10 mm. 


This is a common species in the coral limestones of Limon. 
Toula’s Mitra Almagrensis was described from Tehuantepec. 
Our shells differ from Toula’s figure in being more slender and in 
having the body-whorl less contracted below. 


Gatun Stage: Port Limon. 


Genus FUSUS Lamarck 
Fuscs miocosmius, n. sp. Plate 8, figure 5 


Shell long and slender, with the spire and canal of nearly 
equal length; whorls about 11 plus, very gradually tapering from 
the small nucleus to the body-whorl; whorls convex with indis- 
tinct, appressed sutures; the sculpture consists of large, swollen 
ribs, separated by equally wide interspaces, and the whole 


104 BULLETIN 39 a7 


crossed by strong spirals with finer threads in between; on the 
last whorl there are 7 ribs, which commence on the base, just 
above its union with the long canal, and continue across the 
whorl to the suture; the early spire-whorls have 6 or 7 strong 
spirals with 1, 2 or 3, fine threads between; on the penultimate 
whorl, the earlier spirals have become a primary set of 6 or 7, 
with their intervals occupied by asecondary set, nearly as large 
as the primary, and still smaller tertiary threads; the last whorl 
has about 18 strong spirals in addition to those on the canal; on 
the canal the division into primary and secondary spirals is bet- 
ter shown, there being about 16 primary and the same number of 
secondary spirals; on the extreme tip of the canal the spirals are 
very small and numerous; aperture small, rounded, with the out- 
er lip internally lirated and three or more oblique plaits on the 
columella. 


Height roo, diameter 24 mm. 


This fine species is closely related to the recent Fusus eucos- 
mtus Dall, from the Florida coast and the West Indies. The spire 
of ezcosmius is more slender, has 8 ribs instead of 7, and the out- 
er lip is always smooth, while it is strongly lirated in szocos- 
meus. 


Gatun Stage: Bocas del Toro, Panama. 


Fusus honensis, nu. sp. Plate 8, figure 6 


Shell resembling in general features the /. mzocosmius, and 
with the same number of ribs; the spire is somewhat longer than 
the anterior canal; the whorls are somewhat more convex than 
those of mzocosmtius due to the more strongly knobbed ribs; the 
ribs are crossed on the penultimate whorl by about 6, strong,even 
spirals, there being no secondaries and only occasionally a small 
thread may appear in their intervals; the last whorl, exclusive 
of the canal shows 10 or i1 spirals to which are added 17 or 18 
on the canal; the canal is relatively short and not perfectly 
straight; the aperture is probably rounded (largely broken on the 
type specimen) and with its outer lip internally lirated. 

Height 63, diameter 20 mm. 


277 Costa Rica MrocekNE—OLSSON 105 


Similiar in general form and in the number of its ribs to the 
fF. miccosmius from Bocas, this species differs in its smaller, 
heavier shell, and in the greater coarseness of its sculpture. The 
spirals are heavier and primary in character throughout and the 
ribs are more knobbed and sharp on their crests. From the Do- 
minican /, Hlenekent Sowerby, it differs in being much more 
slender and with fewer, persistent ribs. 


Gatun Stage: Coll. 4, Home Walk Creek. 


Genus FASCIOLARIA Lamarck 
Fasciolaria Gorgasiana Brown and Pilsbry Plate 8, figure 9 


Fasciolaria gorgasiana Brown and Pilsbry, 1912, Proc. Acad. Nat. 
Sci. Philia., vol. 64, p. 506, pl. 22, fig. 5. 

This large Fasciolaria is fairly abundant in the Gatun beds 
of the Canal Zone and is recognized by its shouldered whorls and 
and strong knob-like ribs. The figured specimen is a large shell 
from the Gatun of Rio Betey, Costa Rica. 

Gatun Stage: Gatun, C. Z. 

Water Cay. 
Rio Betey. 


Fasciolaria tulipa Linnzeus, variety Plate 13, figure 4 


Murex tulipa Linnzeus, 1758, Syst. Nat.,ed. 10, p. 754. 
Fasciolaria tulipa Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 
2nd series, p. 354. 
Fasciolaria tulipa Dall, 1890, Trans. Wagner Free Inst. Sci., vol. 3, 
DEIN LOL apl yy 7 uuies on 
The Tulip shell is the common, recent /asctolaria, in the 
Caribbean area, and it is distinguished from its more northern re- 
lative, the dzstans, by its darker coloration and sulcated sutural. 
band. It occurs as fossil in the Pliocene of Florida. 
From the coralline limestones, near Port Limon we collected 
three young specimens and a portion of the spire of a large in- 
dividual of Fasctolaria. ‘The young shells show the nucleus and 


106 BULLETIN 39 27% 


early sulcated whorls of typical ¢ulipa, as fignred by Dr. Dall. 
They however become smooth somewhat earlier and the fragment 
of the spire shows the succeeding whorls to be smooth and lack- 
ing the sulcated sutural band as in dsfans, 


The fasciolaria semtstriata Sowerby is an allied but distinct 
species from the Miocene of Santo Domingo, and likely to be dis- 
covered in the Gatun beds of Costa Rica and Panama. ‘This as 
may be seen from the new figure of Dr. Maury, (Bull. Amer. 
Pal., vol. 5, p. 244, pl. 13, fig. 1), differs conspicuously from the 
distans and tulipa, with which it was united by Gabb, by its 
deep sutural depressed band, which gives to the whorls, a very 
convex or even shouldered appearance. The early whorls are 
spirally sulcated as in ¢u/ija, but have in addition knob-like ribs, 
like those seen on /. gorgastana. 


Useart Stage: Port Limon. 
Fasciolaria MacDonaldi, n. sp. Plate 8, figure 1 


Shell large, (type specimen imperfect, with only the last two 
whorls preserved); number of whorls unknown; the spire-whorls 
are angled about the middle, forming a board, sloping shoulder 
above; the last whorl has the shoulder about the upper third; the 
sculpture is predominantly spiral, consisting of numerous, fine, 
alternating threads, which are slightly roughened by the growth 
lines; the angle of each whorl is longitudinally undulated by 12 
faint ribs; anterior canal is nearly straight with two faint plice; 
outer lip sharp. 


Length (2 whorls) 70, diameter 37 mm. 


Of this large and elegant species, only a single imperfect 
specimen with less than two complete whorls preserved, was col- 
lected from the Gatun beds of the Banana River, Costa Rica. It 
differs from the more common /. Gorgaszana, by its persistent 
and rough spiral sculpture and less heavily tuberculated should- 
er. 


This species is named in honor of Dr. D. F. MacDonald, well- 
known for his geologic work on the Isthmus, during the con- 


270 Costa Rico MrocENE—OLSSON 107 


struction of the Panana Canal, and former Chief Geologist of 
the geological force of the Sinclair Oil Corporation in Panana and 
Costa Rica. 


Gatun Stage: Hill No. 2, Banana River. 


Genus LATIRUS Montfort 


Latirus infundibulum Gmelin, variety Plate 8, figure Io 


Latirus infundibulum Gmelin, Lamarck, Anim. sans, Vert. (ed, Desh. ) 


vol. 9, p. 386. 

Latirus infundibulum Guppy, 1866, Quart. Journ. Geol. Soc. Loudon, 
vol. 22, p. 288. 

Latirus infundibulum Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, 
pe2t7. 


Latirus infundibulum Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. ' 
8, 2nd series, p. 354. 
Latirus infundibulum Maury, 1917, Bull. Amer. Pai., vol. 5, p. 246, 
pla he. 3 
A common recent species of the West indian and Costa Rican 
coasts. Only an imperfent specimen was collected from the Gatun 
beds of lower Pumbri Creek, a small tributary of the Estrella 
River. From recent examples of znfundibulum, it differs by its 
shorter canal, more numerousribs (9 instead 6 or 7),and heavier 
spiral sculpture. The specimen is, however, to fragmentary to 
serve as a type for a new variety or related species. 


Gatun Stage; Coll. 7, Pumbri Creek; GC. R: 
Latirus irazu, n. sp. Plate 8, figure 12 


Shell subfusiform; solid, with a long spire and shorter can- 
al; whorls about 9, with heavy, sharp, knob-like ribs and finer 
spiral threads; the last whorl shows 7, sharp, pointed ribs, which 
are continuous from suture to suture, but only feebly across the 
base of the last whorl; the suture is bordered anteriorly by a 

prominent, frilled band or cord, formed by the elevated edges 


108 BULLETIN 39 280 


of the growth lines: spirals consisting of low, raised threads with 
wide interspaces; canal short, straight above but bent below, and 
with three small, smooth folds on the columella above; aperture 
with a thin outer lip. 

Length 46, diameter 17, last whorl 29, spire 24 mm. 

A smaller and more delicate species than the preceding ZL. zz- 
fundtbulum Gmelin. In the present shell, the ribs are sharp- 
er and more pointed, and the canal is shorter and more strongly 
reflected backwards. The strongly frilled sutural band is lacking 
or only imperfectly developed on znfundibulum. 


Gatun Stage: Port Limon. 
Latirus taurus, 0. sp. Plate 8, figure 4 


Shell large, slender, solid with the spire more than % the 
height of the shell; whorls convex, with indistinct sutures; sculp- 
ture of very regular, narrow ribs with deep interspaces, and cross- 
ed by strong, ridge-like spiral cords; whorls about 9; axial sculp- 
ture on last whorl with about 11 ribs; the ribs commence well 
down on the base and continue to the upper suture and are 
nearly in line acrossthe spire-whorls to the apex; the spirals 
consist of ridge-like cords and finer spiral threads; there are 
2 principal spirals on the spire-whorls which cross the middle 
of the whorls; there are 4 principal spirals on the last whorl 
above the edge of the base, below which there are a few small- 
er threads; the anterior canal has 2 strong cords and finer 
threads are scattered over the whole shell; the anterior canal 
is straight and stocky and carries at its base a deep but narrow 
umbilical pit; the columella has 4 small folds; aperture subo- 
vate, anteriorly extended to form the long narrow canal; out- 
er lip smooth within. 


Height 60, diameter 25, aperture 32 mm. 


A large, solid and strongly sculptured shell. The straight, 
narrow ribs are spaced regularly over the whorls of the shell and 
are continuous from the base to the upper suture. They are 


a8x Costa RicA MiocENE—OLSSON 109 


crossed by heavy, ridge-like, spiral cords, which on crossing the 
deep interspaces between the ribs, form small, sunken pits, so 
that the sculpture appears coarsely trellised. 


Gatun Stage: Toro Cays. 


Genus PTYCHOSALPINX Gill 
Ptychosalpinx ? dentalis, n. sp. Plate 15, figures 14, 18 


Shell buccinoid with convex, cancellated whorls; whorls 
about 6, with channelled sutures so that the whorls appear nar- 
rowly shouldered above; the sculpture is predominantly spiralled, 
the penultimate whorl with 7 strong spiral cords with interspaces 
2 or 3 times their width; small spiral threads may appear in the 
interspaces; the last whorl has 14 or more spirals and the wide 
interspaces with one or more finer threads; the spirals are cross- 
ed by coarse wavy lines of growth, producing a sub-cancellate 
sculpture; the aperture is subovate; a well-developed siphonal 
sinus; the columella carries a single sharp plication at its lower 
end; the outer lip is slightly thickened and bears within about 
12, small, narrow liree. 


Height 28, diameter 11, aperture 16 mm. 

The generic relation of this interesting species is very much 
in doubt. It has the general form of Comznella, but its colum- 
ella is provided with a strong anterior plication. In this regard 
it agrees with Pfychosalpinx of which there are several species in 
the Chesapeake Miocene. The apex of dentalis, appears to have 
been pointed and sharp, although all of our specimens have the 
tip of the spire very much weathered. True Ptychosalpinx is 
characterized by a large, convex and blunt nucleus and the outer 
lip is thin and smooth within. The outer lip of deztalzs is slight- 
ly thickened and internally finely hrated. . 


The P ? dentalis, is a very characteristic fossil of the Denta- 
lium zone of the Upper Uscari formation. 


I10 BULLETIN 39 252 


Uscari formation. Coco Plum, Panama. 
Rio Cocles. 


Comadre Creek, etc. 


Genus PERISTERNIA Morch 
Peristernia insuia, n. sp. Plate 8, figure 11 


Shell small, with a sharp pointed spire, a little longer than 
the aperture; the general form and sculpture of the shell is like 
that of Uvosalpinx cinereus Say; nucleus of 2 small smooth 
whorls, followed by 7 post-nuclear; the profile of the spire-whorls 
is convex and strongly sculptured with ribs and sharp spiral cords; 
the last whorl has 8 ribs which are nearly lacking from the base 
of the whorl; the spiral sculpture consists of 2 principal cords 
about the middle of the earlier whorls above which lie smaller 
threads about the suture; on the later whorls, the spirals are 
somewhat heavier about the middle but irregular with finer, in- 
termediate threads in between the principal ones; base contract- 
ed; aperture subcircular; outer and inner lips crenulated or den- 
ticulated; canal short and bent. 


Height 23, diameter 13, aperture 11 mm. 


This and the following fortugera seem correctly referable to 
the genus FPeristernta Morch. The P. insula is somewhat like 
young specimens of //icata Conrad, from the Chesapeake Mio- 
cene of eastern United States, but has a longer and more pointed 
spire, and heavier sculpture. 


Gatun Stage: Water Cay. 
Peristernia tertugera, n. sp. Plate 8, figure 13 


Shell elevated with a spire much longer than the aperture; 
nucleus of about 2% small, smooth whorls; the post-nuclear 
whorls about 7; sutures distinct; whorls strongly convex, should- 
ered; sculpture of narrow, heavy ribs, widely spaced and num- 
bering on the last whorl about 9; the rib are continued across the 
whorls from suture to suture and for a short distance down on 


283 Costa Rica MIocENE—O1S¢sON II 


the base; the spirals consists of irregular sharp cords; the early 
spire whorls carry 2 main spiral cords but bordered above and be- 
low by finer threads; the 2 principal spirals are continued on the 
later whorls but are nearly equalled in strength by the other 
spirals and hence lose their prominence; a short anterior canal; 
base strongly contracted. 


Height 29, diameter 14.5, aperture 12 mm. 


Distinguished from the preceding zmsau/a by its very much 
longer spire, more strongly contracted base and different spiral 
sculpture. 


Gatun Stage: Port Limon. 


Genus XANCUS Bolton 
Xancus scopulus, n. sp. Plate 11, figure I 


Shell large, solid and heavy; spire nearly as long as the aper- 
ture; spire-whorls 6 plus (the tip broken), strongly coronate 
above and with large, wide, persistant ribs; the earlier spire- 
whorls are simply shouldered or angled about the middle, but the 
area about the upper sutures rapidly deepens and on the later 
whorls is a deep, excavated sutural zone, above which project 
the ends of the ribs; there is a strong sutural cord and a rather 
wide, ribbon-like band just above, forming at first a strongly ap- 
pressed suture; the last whorl has about 7 large, wide ribs, the 
areas between appearing as troughs or depressions; the sutural 
excavated zone carries several irregular spiral threads which are 
crossed by large growth-lines so that the resulting sculpture is 
more or less cancellate, more particularily on the earlier whorls; 
the growth-lines cross the sutural cord and upon the ribbon 
above, become much crowded and strongly bent forwards; the 
young shell was sculptured over the whole shell with strong 
spirals, but with maturity, the spirals become obsolete and the 
shell is smooth and polished; columella with 3, strong plicee as in 
the recent scolymus; a long anterior canal, with a narrow, deep 
umbilicus behind the spreading inner lip. 


m2 BULLETIN 39 284 


Height 265, diameter 136, aperture 158, last whorl 195 
mm. 

This very remarkable species is an extreme development of 
the X. scolymus stock in which the upper portion of the whorl 
becomes a wide, deep, excavated sutural zone. Above this ex- 
cavated zone, project the high, rounded or appressed ribs and the 
carinate edge of the whorls. X. scolymus Gmelin, a recent spec- 
ice found plentifully along the north Panama coast, has the 
whorls simply shouldered, often merely rounded. 


The Santo Domingan Miocene contains X. validus Sowerby, 
which has been identified by some with scolymus. In validus, the 
ribs are more numerous (about 10) and are sharper and more tub- 
ercular in form. 


Gatun Stage: Banana River. 


Genus MELONGENA Schumacher 
Melongena consors Sowerby Plate 9, figure 1 


Pyrula consors Sowerby, 1849, Quart. Journ. Geol. Soc. London, vol. 6, 
Pp. 49. 

Melongena melongena Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p- 
205. Not of Linneeus. 


Pyrula melongena Guppy, 1874, Geol. Mag., vol. 11, p. 438. 

Pyrula melongena Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 
32, P- 523. 

Melongena consors Dall, 1900, Trans, Wagner Free Inst. Sci., vol. 3, pt 
1) 169) bo 


Melongena consors Maury, Bull. Amer. Pal., vol. 5, p. 249, pl. 14, 
fig. 5. 
The Melongena consors is not a common fossil in Costa Rica. 
It is closely related to the recent 7. corona Gmelin of the West 
Indies, the fossils shells differing mostly in having a longer spire 
and somewhat different sculpture above. 
The Costa Rican examples are exactly like Miocene specti- 
mens from Santo Doingo. As a fossil it occurs in the Miocene 
of Jamaica, Santo Domingo and Venezuela. 


285 Costa Rica MrockENE—OLSSON 113 


Gatun Stage: Coll. 5, Old Man Sam Creek. 
Cocles Creek. 


Genus SOLENOSTEIRA Dall 
Soilenesteira Dalli, Brown and Pilsbry Plate 8, figure 8 


Solenosteira dalli, Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. 
Phila., vol. 63, p. 348, pl. 24, fig. 14. 

Coralliophila gatunensis Toula, 1911, Jahrb. der K-K Geol. Reichsan- 
stalt, vol. 61, p. 502, pl. 30, fig. 9. 

A common species in the Canal Zone and figured here for 
comparision with the following species. The whorls are strong- 
ly shouldered, a feature accentuated by the few, but high,angled 
ribs. The spirals are heavy, raised cords, over and between 
which are finer secondary and tertiary threads. 


Length 34, diameter 23, last whorl 27, spire 15 mm. 
Gatun Stage: Gatun, Mt. Hope, C. Z. 


Solenosteira Vaughani Dall, var. medioamericana, n. var. Plate 8, fig. 7 


cf. Solenosteira Vaughani Dall, 1903, Trans. Wagner Free Inst. Sci., 
vol, 3, pt. 6, p. 1633, pl. 60, fig. 18. 

Shell solid, with a small pointed nucleus of 2 smooth whorls, 
and 6 post-nuclear; spire conic, of about % the total length of 
the shell; whorls convex, or bluntly shouldered about the mid- 
dle ; sculpture of numerous low ribs which are rounded, and 
heaviest on the shoulder angles; There are 7 to Io ribs on the 
last whorl; spirals of raised cords, with finer threads on the in- 
terspaces; the spire-whorls have about 6 primary spirals, the last 
whorl with 18, including those of the short canal; canal short 
with a deep but narrow umbilical pit; aperture oval, a thick in- 
ner and outer lip, the outer with about 10, long, entering lire 
and the inner with three small denticles above. 

Length 42, diameter 24, last whorl 34, spire 17 mm. 
34 2S 28 13 mim. 


The Solenosteira Vaughani is a Chesapeake Miocene species 


T14 BULLETIN 3G 286 


from Jackson Bluff and Coe’s Mill, Florida. Dall’s figure in the 
Wagner Free Institute, isso like our shell from western Panama 
and Costa Rica, that there can be but little doubt of their close 
affinities. 

The present shell is larger than the S. Dalli, of the Canal 
Zone, with more numerous, obtuse and not sharp ribs, and with 
a wider and deeper umbilical pit. A canal is developed at the 
posterior angle of the aperture asin the recent S. pallida Brod. 
while in S. Dadli, the posterior portion of the aperture is round- 
ed and there is no canal. 

Gatun Stage: Toro and Water Cay, Panama. 

fiill ra, Banana River, C. R. 


Solenosteira chiriquiensis, n. sp. Plate 8, figure 3 


Shell large, heavy; spire about % the height of the shell; 
whorls about 7, strongly angled about the middle and carrying 
heavy knob-like ribs, crossed by heavy spiral cords; the last 
whorl has 8 ribs which are developed only on the middle of the 
shell, being lacking from the upper slope and from the base of 
the last whorl; the tops of the ribs are crossed by 2 strong,spiral 
cords with a wide, trough-like interval between; above the 2 
principal spiral cords, there are 4 smaller cords on the upper 
slope and on the base and canal ro or 11; base of the last whorl 
contracted to form the straight canal which carries a deep, narrow 
umbilius; aperture subelliptical. 

Height 57, diameter 34, aperture 34 mm. 

It is possible that this species belong to the genus Cymza 
rather than Solenostetra, but its aperture is so completely filled 
with a hard sandstone matrix that the presence or absence of a 
columellar fold cannot be determined. Its sculpture however is 
more like Solenosteiva than Cymia. ‘The species will be recog- 
nized by its large size and characteristic sculpture. 


Gatun Stage: Water Cay. 


287 Costa Rico M1ocENE—OLSSON II5 


Genus METULA H. and A. Adams 


Metula cancellata Gabb Plate 10, figure 12 


Metula cancellata Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, Pp. 
205. 
Metula cancellata Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 2nd 
series, p. 351. 
Metula cancellata Dall, 1903, Trans. Wagner Free Inst. Sci., vol, 3, pt. 
6, p. 1584. 
Metula cancellata Maury, 1917, Bull. Amer. Pal., vol. 5, 249, pl. 14, 
fig. 19. 
Less common in Costa Rica and Panama than the following 
species and from which it is easily distinguished by its usual 
smaller size, more delicate shell, and fine sculpture. 


The spiral threads on the last whorl number about 37 and 
are crossed by nearly as fine longitudinal ribs. The resulting re- 
ticulate sculpture is fine and neat. Our largest shell, a specimen 
from Bocas with 6 whorls measures: 


Length 19, diameter 6.5, last whorl 14, spire 9, aperture 
IO mm. e 


Gatun Stage: Bocas del Toro, Panama. 
fiill ra, Banana River, C. R. 


Metula Gabbi Brown and Pilsbry Plate 10, figure 11 


Metula Gabbi Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., 
vol. 63, p. 351, pl. 25, figs. 4, 8. 


This beautiful species is fairly common in the Canal Zone, 
but usually in a fragmentary condition. It related to the W/etula 
cancellata Gabb, differing mainly in its larger size and heavier 
sculpture. Ona speciman of 8 whorls from Gatun, the spirals on 
the spire whorl number about 8 and 39 on the last whorl and an- 
terior canal. The 2 upper spirals are noticeably stronger than 
the others. Aperture long and narrow and finely denticulated 
along the interior of the outer lip. 


116 BULLETIN 39 288 


The figured specimen from Gatun has the following meas- 
urements: 


Length 29, diameter 10.5, last whorl 22, spire 12.5, aper- 
ture 16 mm. 


Gatun Stage: Gatun and Mt. Hope, C. Z. 
Metula Harrisi, n. sp. Plate to, figure Io 


Shell solid and more coarsely sculptured then Wetula Gabéz, 
and with a longer and broaded spire and shorter aperture; whorl 
6 plus, (apex lost); sculpture of spiral cords and finer, curved 
longitudinal ribs; the spire whorls with 8 spiral cords which are 
finest next to the lower suture, become progressively stronger 
above; on the last whorl and canal there are about 25 spirals; 
the spirals are slightly nodulated by the ribs of which there are 
about 40 on the last whorl; certain of the spirals on the middle of 
the whorl are double; aperture ovate-elliptical with a strongly 
thickened outer lip, internally with about 19 short lire; inner 
lip smooth; canal short and twisted. 


Length 26.5, diameter 11, last whorl 19.5, aperture 13.5, 
spire 14 mm. ¢ 
A solid and more coarsely sculptured species than the A/etula 
Gabbi. ‘The spire is broad and porportionately longer, due to the 
shorter aperture. In Gaddz the spirals are of nearly equal 
strength over most of the shell surface with exception of those 
immediately bordering the upper suture. In the present species 
the spirals are finer below and progressively become stronger post- 
eriorly. 
The type specimen was collected in the lower Gatun sand- 
stones of Coco Plum, Panana. 


Gatun Stage: Coco Plum, Bocas del Toro, Panama. 
Metula Harrisi var. limonensis, n. sp. Plate 10, figures 5, 6 


Shell more slender; a narrow and longer aperture more 
nearly the length of the spire; sculpture coarsely reticulate, 


289 Costa RicA MrocENE—OLSSON 117 


with ro spirals on the penultimate whorl, about 26 on the last; 
aperture long, with a thickened outer lip, internally lirated in 
unison with the external spiral cords; inner lip smooth witha 
long, bent, beak-like canal. 


Length 27, diameter 9.50, last whorl 19, aperture 14, 
spire 13 mm. 

Several specimens from the coral limestones of Port Limon. 
They are most closely related to Harrisz, in their heavy shell, 
coarse sculpture, and thickened outer lip. They differ in being 
more slender, a longer aperture and in minor details of their 
sculpture. 


Gatun Stage: Port Limon, 


Genus PHOS Montford 


Phos gatunensis Toula Plate 9, figures 4, 5 


Phos gatunensis Toula, 1909, Jahrb. der K-K Geol, Reichsanstalt, Wein 
VOlNsop. 7Ol, pl-i28, tig 6: plo 25 (hee T, 

Phos gatunensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila. 
vol. 63, p. 349, pl. 25, figs. 1, 2. 

The Phos gatunensis is possibly the most common fossil in 
the Gatun beds of the Canal Zone and the adjacent sedimentary 
areas in the Province of Colon, but it still remains to be collected 
outside of this, its type area. 


The shell is typically rather slender, with a sub-reticulate 
sculpture formed by the intersection of its numerous, straight 
longitudinal ribs, and the nearly equal, regular, strap-like, spiral 
cords. 


A typical specimen measures as follows: 
Length 30, diameter 13, last whorl 20, spire 15 mm. 
Gatun) Stage: (Gatun, CZ. 


Phos mexicanus Bose Plate 9, figures Io, II 


Phos mexicanus Bose, 1906, Bol. de Inst. Geol. de Mexican, numero 
22, p. 38, pl. 4, figs. 18-21. 


118 BULLETIN 39 290 


Shell more broad and solid than gatumensis of the Canal 
Zone; whorls about 8, those of the spire with nearly straight or 
only slightly convex sides; last whorl large, broad, convex and 
widest about the middle; the sculpture is much more coarsely re- 
ticulate than in gatunens?s, with the intersection points of the 
ribs and spirals more or less subnodulose; the last whorl has 
about 18 ribs, which on the back of the outer lip are more close- 
ly spaced than elsewhere; the spiral cords are not so heavy as the 
ribs, separated by wider interspaces which carry one or some- 
times two very, fine, spiral threads; in addition, the spiral inter- 
spaces are very finely and neatly sculptured with fine, raised, 
longitudinal lines, corresponding to the growth-lines; on the 
spire-whorls there are 5 spiral cords, 13 on the last whorl and 3 
more on the canal; canal short, twisted; aperture sub-ellip- 
tical, with a moderately heavy, but not thickened outer lip and 
internally lirated. 


Length 30, diameter 15.5, last whorl 21, spire 14.5 mm. 
A broader, more solid and more coarsely sculptured shell 
than the Phos gatunensts of the Canal Zone, 


It was described from the Miocene of Paso Real, near Tux- 
tepec, in the State of Oaxaca, Mexico, by Dr. Bose, whose excel- 
lent figures agree exactly with specimens from Water Cay. The 
several species of Phos, are often extremely abundant but local 
in their distribution. Such is the case of Phos gatunensts of 
the Canal Zone, the Phos Mooret var. costaricenis of the Banana 
River. Likewise in the present instance; the mextcanus, is a very 
abundant fossil of the Gatun beds of the islands of the Chiriqui 
Lagoon and the adjacent Valiente Peninsula. A few specimens 
have also been collected in Costa Rica. 

Gatun Stage: Water Cay. Toro Cay. Coco Plum, ete. 

Rio Cocles, C. R. 


Phos Moorei Guppy, var. costaricensis, n. var. Plate 9, figures 8, 9 


Shell of medium size, solid; spire long about one-half the to- 
tal length of the shell; whorls 10, of which the first two belong 


2QT Costa Rica MrocENE—OLSSON 119 


to a small smooth nucleus; succeeding whorls are sculptured with 
heavy persistent, straight ribs (13 or 14) on the last whorl; the 
ribs are crossed by even, raised, spiral cords; the spire-whorls 
have 4 spiral cords and a small one close to the upper suture; an- 
terior canal short and strongly twisted; aperture subelliptical, 
the outer lip with about Io, strong and regular internal lire. 


Height 27, diameter 12, last whorl 17, spire 14 mm. 


This is the common /hos of the Banana River. Although 
closely related to Mooret, from Jamaica, the Costa Rican fos- 
sils are smaller and more slender, as compared with Guppy’s 
nearer they Ouarterly, Journal, volw22s\plario. ig.) 1: 


Dr. Maury’s Phos Moorez, from the Miocene of Santo Do- 
mingo probably represents a distinct species, differing by its 
more slender spire, and in sculpture. The spiral intervals of 
costaricensis are smooth, while in the Santo Domingan shells, 
they carry fine spiral threads, which are best seen on the spire- 
whorls, sometimes becoming obsolete on the later. 


Gatun Stage: Banana River, 
Old Man Sam Creek, r mile from shore or 
beach. 


Phos elegans Gupp, var. limonensis, n. var. Plate 9, figures 12, 13 


Shell usually smaller, more slender and delicate than the pre- 
ceding species; the spire is long, composed of about 9 whorls: 
the nucleus consists of 4 whorls, the first 2 are small, 
smooth and convex, the last 2, smooth, but sharply keeled about 
the periphery on the last 1-4 of the last nuclear whorl, faint 
curved lines appear above the peripheral keel, they gradually in- 
crease in strength and on the Ist post-nuclear whorl become the 
longitudinal ribs; the sculpture consists of fine, longitudinal ribs 
(23 or 24 on the last whorl); occasionally a rib may become 
greatly enlarged, forming varix-like thickenings corresponding 
to resting stages; the ribs are overrun by primary, secondary and 
tertiary spiral threads; the later spire whorls have about 6 spirals, 


120 BULLETIN 39 202 


with about 17 on the last whorl andthe canal; aperture suboval, 
with a heavy but not thickened outer lip, internally lirated. 
Length 27, diameter 11, last whorl 17, spire 4.5 mm. 

This shell is related to the Phos elegans Guppy of the Mio- 
cene of Jamaica and Santo Domingo. Our shells differ from 
typical examples of elegans from Santo Domingo, in being larg- 
er, and more coarsely sculptured. 

Varix-like thickenings occur scattered along the spire-whorls. 
The protoconch of typical elegans and limonensts are practically 
identical. The recent West Coast Phos Veraqguensits Hinds isa 
related but more finely sculptured species. 


Gatun Stage: Port Limon. 
Bocas del Toro. 


Phos estreilensis, n. sp. Plate 9, figures 17, 18 


Shell small, slender, solid; spire much longer than the ap- 
erture, pointed; whorls of the spire 6 or more, convex in profile 
with deep sutures; sculpture of narrow, straight ribs which com- 
mence on the columellar area and pass across the whorl to the up- 
per suture; there are 12 or 13 ribs on the last whorl which are 
evenly and widely spaced except on the back of the outer lip 
where they are smaller and more numerous; the spirals consist 
of a primary set of threads or cords with a single, smaller second- 
ary thread in each interspace except on the base and the canal; 
the primary threads crossing the ribs form small, whitish tuber- 
cles; the spire-whorls have 4 primary spirais and there are 11 on 
the last whorl, exclusively of the canal; aperture sub-elliptical, 
produced anteriorly to form the straight anterior canal; outer lip 
thickened and internally lirated. 


Height 20, diameter 8, aperture 9 mm. 


A small, slender species, coarsely sculptured like the larger 
and broader Phos Moorei variety costaricensis. It is limited in 
our collections to the Gatun beds of the Estrella River. 


Gatnu Stage: Coll. 6, 7, Estrella River. 


293 Costa Rica MrockNE—OLSSON 121 


Phos subsemicostatus Brown and Pilsbry Plate 9, figure 15 


Phos subsemicostatus Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. 
hilary Vols) 6351 p41350.) ple 25) figai 3. 
A larger and finer sculptured species than the Dominican 
Phos semicostatus of Gabb. A single imperfect specimen was col- 
lected from Zone E of Saury Creek. 


Gatun Stage: Gatun (Brown and Pilsbry) 
Zone E,\ Saury (Creeks Oak. 


Phos Veatchi, n. sp. Plate 9, figures 2, 3 


Shell large and heavy, with a moderately long, coarse spire 
of about 9 whorls; nucleus small, of 2 smooth whorls; whorls of 
the spire moderately convex, with strong ribs continuous from 
suture to suture; sutures appressed and bordered below by a 
small spiral; the last whorl is widest about the middle and con- 
tracted below to the short anterior canal; sculpture of heavy, 
persistent ribs and regular raised spirals; the last whorl has 8 
ribs; the spirals consists of raised threads, separated by wide, in- 
terspaces; an occasional secondary spiral will appear in the in- 
terspace but they are generally smooth or only obsoletely sculp- 
tured by the growth lines; there are 8 spirals on the spire-whorls, 
16 or 17 on the last whorl; aperture ovate, with the outer lip 
thickened by the last rib, and internally with 9, long, entering 
liree; inner lip formed by a raised, spreading callus, armed above 
by a tooth-like denticle and a strong fold below on the edge of 
the canal. 


Length 43, diameter 22, last whorl 28, spire 25 mm. 


A large, robust species from Hill 1a of the Banana River. It 
differs from the szbsemzcostata by its strong persistent ribs and 
coarser spirals. 


It is named for Dr. A. C. Veatch, Director of the Explora- 
tion work of the Sinclair Oil Corporation, under whose direction 
the Costa Rican collections were made. 


122 BULLETIN 39 204 


Gatun Stage: Hill ra Banana River. 
Phos beteyensis, n. sp. Plate 9, figure 6 


Shell of the type of subsemicostata; whorls 3, moderately 
convex; sculpture with the axial element very weak but present 
on the penultimate whorl as fine, sub-obsolete, narrow ribs or ir- 
regular wrinkles on the shell surface; they are absent from the 
last whorl; the spiral sculpture consists of a broad zone, about 
the upper 1-3 of the spire whorl on which are irregularly distri- 
buted about 5 raised spiral threads; below this zone, the sculp- 
ture is largely in the formof broad uneven bands formed between 
incised spiral lines; on the last last whorl, this later zone occup- 
pies the middle of the whorl; the zone of raised spirals above the 
more smooth area of banded spirals gives to the whorls a slightly 
coronated appearance. 

Length 27 (2% whorls), diameter 15, last whorl 21 mm. 

The type specimen is very incomplete, but its characters are 
so distinctive and unlike any other species of Phos which we 
have seen that its discription is advisable. Like the swédsemzcost- 
atus the longitudinal ribs or coste fade out on the later 
whorls. 


Gatun Stage; Betey Creek, C. R. 


Phos meiuloides Dall Plate 9, figure 16 
Phos metuloides Dall, 1896, Proc. U. S. Nat. Mus., vol. 19, p. 310, pl. 
28, fig. 15. 
Phos metuloides Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., 
vol. 63, p. 350. 
Phos metuloides Maury, 1917, Bull. Amer. Pal., vol. 5, p. 252, pl. 14, 
fig, 17. 


A small species with a neat sculpture recalling that of J/e- 
tula cancellata and gabéi. It isnot rare in the Gatun beds of 
Mt. Hope in the Canal Zone. Dall also records it from the Mio- 
cene of Santo Domingo. 

Length 19, diameter 9, spire whorl 13.5, spire 10 mm. 

Gatun Stage: Mt. Hope. 


295 CosTA Rico MiockENE—OLSSON 123 


Genus ALECTRION Montfort 
Alectrion ranuncula, n. sp. Plate 9, figures 20, 22 


Shell of medium size, with a large body-whorl and a sharp 
pointed spire; the general form and sculpture recalls the recent 
A. acuta Say; spire about the same length as the aperture and 
canal and composed of about 7 whorls, of which the first 2 belong 
to the small, smooth nucleus; the profile of the spire-whorls is 
convex with deep, excavated sutures; sculpture of narrow ribs, 
the tops of which are crossed by spirals and separated by wide, 
smooth interspaces; the last whorl has about g ribs which are 
large and humped on the ventral face, finer and more numerous 
on the back; there are 3 spirals on the spire-whorls and 7 on the 
last with several more on the short canal; aperture subcircular 
with a large, heavy outer lip, internally denticulated. 

The general form and sculpturing recalls the recent East 
Coast <Alectrion acuta Say, but the Costa Rican shell is much 
broader as indicated by the following comparative measurements 
with 4. acuta Say, of the same height. 


Height 14.00, diameter 6.25, last whorl 8, aperture 6 mm. 
(Alectrion acuta Say) 

Height 14.50, diameter 8.25, last whorl 9g, aperture 7.5 mm. 
(Alectrion ranuncula, n. sp.) 

It is not uncommon in the Gatun beds of the Banana 

River. 
Gatun Stage: Hill ra, Banana River. 
Coll. 7 Estrella. 


Alectrion losquemadica Maury Plate 9, figure 21 


Alectrion losquemadica Maury, 1917, Bull. Amer. Pal., vol. 5, p. 255, 
pls wigsy 225 23% 
The Costa Rica fossil is similiar in its sculpturing and form 
to the common recent West Indian A. ambigua Montfort. As 
Maury has pointed out in regard to the Dominican examples, the 


124 BULLETIN 39 296 


difference between losguemadica and ambigua, liesin the proto- 
conch, which in typical amdéigua is short and blunt, but pointed 
and composed of several whorls in /osquemadica. Our shells are 
larger than typical lJosquemadica from Santo Demingo. 
Height 11.50, diameter 7.00 mm. 
Gatun Stage: Port Limon. 
Coll. 7, Estrella River. 


Alectrion merenda, n. sp. Plate 9, figure 19 


Shell small, a large body whorl anda sharp pointed spire; 
nucleus of 2 small, convex whorls followed by 5 post-nuclear 
whorls; sutures distinct but not excavated; the sculpture con- 
sists of about 12 ribs which are variable in their strength and 
spacing; the ribs are crossed by subequal spiral cords separated by 
wide, interspaces in which there may appear a smaller intermed- 
iate spiral; the suture is bordered by a set of 2 or 3 finer spirals; 
there are 3 spirals on the spire-whorls and 8 on the last whorl 
above the contracted zone of the canal; aperture subcircular with 
a thick, heavy outer lip. 


Height 7, diameter 4.5 mm. 
The aperture is completely filled with a hard matrix which 
completely conceals the characters of the inner and outer lips. 
Uscart Stage: Old Man Sam Creek. 


Genus NORTHIA Gray 
Northia northiae Gray, var. miocenica, n. var. Plate 9, figures 7, 14 


ef. orthia northie Gray, 1834, in Griffith’s Cuvier, pl. 30, fig. 2. 
ef. Northia northie Kiener, 1832 Icon., Buccinum, p. 23, pl. 9, fig. 
28. 

The Northia northie Gray, (NV. serrata Dufresne) is a 
common and characteristic West Coast or Pacific species, ranging 
from the Bay of Panana to Guayaquil Ecuador. The fossils rep- 
resented by three incomplete specimens differ only from recent 
examples from Panama in lacking the hump-like thickening in 


2907 Costa Rica MIocENE- OLSSON 125 


back of the outer lip and the corresponding shoulder to its whorls. 
But this is not an importnnt character and recent examples may 
sometimes lack this character. 

But few investigators who have studied the West Iadian 
Miocene have failed to note the large percentage of species, close- 
ly related or even identical with recent Pacific species, a condition 
indicative of the union of the Atlantic and Pacific, permitting a 
free intermingling of their respective faunas. Late during the 
Miocene, this union was brought to a close or largely restricted, 
and since then the Pacific element has gradually given way before 
the encroachment and development of the West Indian. That 
this extinction has been a gradual one is shown by the Pleisto- 
cene deposits of Panama containg several common West Coast 
Species not known from the recent Caribbean. Such species are 
the Pecten veniricosus Sowerby and Norihia northie both record- 
ed by Dall from the Pleistocene of Panama or Costa Rica. 

Gatun Stage: Middle Creck. 

Banana River, Hill No. 3. 
Goll OmiRed Glog Greek: 


Genus COLUMBELLA Lamarck 
Columbelia submereatoria, n. sp. late 10, figures 33, 34 


Shell ovate, solid, a small conic spire and a large body- 
whorl; whorls about 7; spire-whorls but slightly convex so that 
the profile of the spire is nearly plane; last whorl large, broadly 
convex about the upper 1-3 and sloping evenly below to the 
short, produced anterior canal; sculpture of subobsolete spirals, 
there being about 6 on the spire-whorls and about 22 on the last 
whorl; aperture linear with a thickened outer lip, expanded in 
the middle and finely and evenly crenulated throughout; anteri- 
or canal short and armed with 8 strong denticles. 


Length 17, diameter 10.5, aperture 11, spire 5.5 mm. 


Closely related to the recent C. mercatoria Linn., common 
along the northern Costa Rican ccast, but differs in its more 


126 BULLETIN 39 298 


globose form and finer spiral sculpturing. Traces of coloration 
are still preserved as faint blotches of yellow scattered over the 
back of the shell. 


Typical Columbella has heretofore not been recorded from 
beds older then the Pliocene either in America or Europe. 


Gatun Stage: Zone 5, Red Cliff Creek. 


Genus STROMBINA Moerch 


Strombina ambigua Guppy Plate 10, figure 9 


Columbella ambigua Guppy, 1866, Quart. Jour.Geol. Soc. London, vol. 
22, p. 288, pl. 16, fig 8. 


Strombina ambigua Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, 
pt. 6, p. 1584. 

This species was described from the Bowden beds of Jamai- 
ca, of Miocene age. Our shells from Costa Rica agree exactly 
with Guppy’s figure in the Quarterly Journal. 

A large species of Columbelloid aspect due in large measure 
to the slight thickening of its outer lip. The sculpture consists 
of about 25 ribs, which are smooth on the spire whorls and upper 
half of the last. The base of the last whorl and the canal have 
in addition about 14 spiral cords. 


Length 27 (7 plus whorls), diameter 10, spire 15, aper- 
ture 13 mm. 
Gatun Stage: Banana River. 


Strombina Lessepiana Brown and Pilsbry Plate 10, figures 13, 20 


Strombina lessepiana Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. 
Phila., vol. 63, p. 352, pl. 25, figs. 11, 12. 

Columbella (Strombina) gatunensis Toula, 1911, Jahrb. der K-K, 
Geol. Reichsanstalt, Wien, vol. 61, p. 501, pl. 30, fig. 8. 

An abundant species in the Canal Zone and with the excep- 
tion of S. ambzgua, the largest species of Stvombina in the Mio- 
cene beds of Panama and Costa Rica. The spire is long and 
smooth, with the exception that the upper sutural zone is usual- 


299 Costa Rica M1ocENE—OLSSON W277 


ly distantly raised into small, longitudinal plicsee which is most 
characteristic. An average specimen from the Canal Zone will 
measure: 

Length 25, diameter 10, spire 13, aperture I2.50 mm. 


We collected from the lower Gatun sandstones of Cocles 
Creek, near Old Harbor, Costa Rica, several small Strombinas 
which except for size are typical Lessepiana. 

Gatun Stage: Gatun and Mt. Hope, C. Z. 

Cocles Creek, C. R. 


Strombina matima, n. sp. Plate 10, figure 7 


Shell ovate, moderately solid with a long evenly tapering and 
sharp-pointed spire; nucleus small, pointed and of three, smooth 
whorls; post-nuclear whorls 6; suture distinct and bordered an- 
teriorly by a small sutural ridge; the sculpture consists of about 
18, smooth, narrow ribs, which are lacking from the back of the 
last whorl, which is smooth; the ribs are oblique in direction and 
along the upper sutures are slightly knobbed by the sutural ridge; 
the base of the last whorl and canal carries about 12 spirals; aper- 
ture subelliptical with a heavy outer lip internally provided with 
3 or 4 denticles about the middle; anterior canal straight and 
smooth within. 

Height 12, diameter 5.25, aperture 6 mm. 

This is a species witha long, evenly-tapering spire and 
small nucleus. The spire-whorls are sculptured with numerous, 
small, smooth riblets. The riblets are obsolete on the back of 
the last whorl which is entirely smooth except for the spirals 
about the base and canal. .S. matima resembles S. Basst Maury, 
from the Miocene of Santo Domingo but is more slender and has 
a great many more ribs. 

Gatun Stage: Hill ra, Banana River. 

Coll. 7, Estrella River. 


Strombina cricamola, n. sp. Plate 10, figure 18 


Shell ovate, solid, with a large, convex and spirally sculp- 


128 BULLETIN 39 300 


tured body-whorl and a long attenuated spire; whorls 6 plus 
(apex broken so that at least one complete whorl is lost from the 
type specimen); the first 4 whorls are smooth, after which a 
few incised spiral lines appear just below the upper suture; these 
spirals gradually increase in strength and in number and on the 
following whorls have become strong, raised spiral cords; the pen- 
ultimate whorl has 5 spirals, the last about 24 (including those 
of the anterior canal); the upper half of the last whorl and some- 
times the penultimate, are longitudinally plicated with 18 rib-like 
costz; aperture narrow, with a thickened outer lip, notched 
above and obsoletely denticulated below; the inner lip is smooth; 
canal short, twisted. 

Length 16, diameter 8.5, aperture 9, spire 7 mm. 


An unusually short and broad species, distantly related to 
the C. pulcherrima Sowerby, recent from the Gulf of Dulce. Two 
specimens were collected from the Gatun sandstones of Water 
Cay. 

Gatun Stage: Water Cay. 


Strombina costaricensis, u. sp. Plate 10, figures 8, 17 


Shell solid, ovate, with a long attenuated spire; last whorl, 
large, flattened or contracted about the middle; whorls about 9%; 
the first 6 whorls are smooth and form the long attenuated spire; 
faint longitudinal plicee appear on the 7th, which increase in size 
and form the rib-like costee on the later whorls; axial ribs persist- 
ent and number on the last whorl about 15; the spirals begin 
shortly after the first appearance of the longitudinal plice as 
faint incised lines near the upper suture; these spiral lines are 
further increased by the introduction of more lines below and be- 
come on the following whorls raised spiral cords; the penultt- 
mate whorl has 5 or 6 spirals, the last with about 20; the spirals 
vary in strength on individual shells and on some specimens may 
be lacking from the middle of the whorl, forming a smooth me- 
dian band; aperture linear, small and coutracted and with a cal- 
loused inner and outer lip ; outer lip strongly thickened, notched 
above and feebly dunticulated below; canal short, straight. 


301 Costa Rica Mr1ockNE—OLSSON 129 


Length 15.5, diameter 7.5, spire 8, aperture 8 mm. 

A species with an Azachis-like sculpture, the ribs and gen- - 
erally the spirals being persistent. The body-whorl is flattened 
or slightly contracted about the middle, but less so than in the 
following variety. A long, tapering and nearly smooth spire as 


in the preceding S. cricamola but much more slender and at- 
tenuated. 


Gatun Stage: Headwater of Middle Creek, C. R. 
Strembina costaricensis, var. musanica, n. var. Plate 10, figures 26, 27 


Shell ovate, with a long, conic spire of 8 or more whorls; 
last whorl strongly contracted or indented about the middle; the 
first 3 whorls or more (apex broken) are smooth or sculpturless; 
on the following 3 whorls, the sculpture is largely of numerous, 
straight rib-like coste, the spirals being only very faint and in- 
distinct; on the last whorl the ribs number about 14; spirals 
about 9 on the penultimate whorl as faint raised threads and on 
the last whorl may be lacking from the middle or contracted 
zone; aperture linear-ovate, a thickened outer lip with 3 faint 
denticles below the upper notch; canal straight. 

Length 15, diameter 6.5, spire 13, aperture 8 mm. 

Probably a distinct species from the preceding and differing 
in its more evenly tapering and more strongly sculptured spire, 
and deeply contracted body-whorl. It resembles somewhat the 
figure of S. carfagensis Brown and Pilsbry from Colombia, but is 
smaller and has more numerous ribs. ‘The periphery of S. car- 
tagensis is rounded. 

Gatun Stage: Banana River, C. R. 

East fork of Red Cliff Creek. 


Strombina sincola, n. sp. Plate 10, figures 19, 21 


Shell small, with a large, conic spire and a dorso-ventrally 
flattened body-whorl; whorls about 8, the first 3 belonging to the 
small, smooth, pointed nucleus; sculpture of the post-nuclear 
whorls of numerous, straight, strong, longitudinal ribs, which are 


730 BULLETIN 39 302 


persistent to the penultimate whorl, but are laking from the 
last whorl which is smooth and polished; the penultimate whorl 
has about 15 ribs; the ribs are generally beaded about their up- 
per extremities; sutures distinct; last whorl large and strongly 
laterally flattened between a prominent ridge or hump on the left- 
hand side of the whorl and the large outer lip; aperture sub- 
elliptical with a large, expanded outer lip, deeply indented be- 
hind; canal short and beak-like and spirally striated with about 
8 threads on the back. 
Length 7, diameter 3.25, spire 3.50, aperture 3.75 mm. 

A small species abundant in the Gatun beds of Hill 1a, of the 
Banana River. The last whorl is rather small and strongly flat- 
tened dorso-ventrally. The spire is large and about % of the to- 
tal length of the shell. 


Gatun Stage: Hill ra Banana River. 
fiill 3, Banana River. 


Strombina chiriquiensis, n. sp. Plate 10, figures 14, 24 


Shell small and in genral characters like the preceding but 
with a larger body-whorl and a shorter, more conic spire; whorls 
about 8, of which the first 3 are smooth and belong to the small 
pointed nucleus; the succeeding whorls until the penultimate are 
sculptured with fine,straight, longitudinal ribs,there being 19 on 
the penultimate whorl; the last whorl israther large, smooth and 
dorso-ventrally flattened between a hump-like ridge on the left 
side and the large, expanded outer lip; canal short, beak-like and 
spirally striated; aperture linear-ovate, deeply indented behind; 
inner and outer lips are both heavily calloused, especially about 
their posterior portions; over thiscallous and between the outer 
and inner lips passes the small posteriorly directed canal; lower 
half of the inner and outer lips denticulated with 8 or 9 strong, 
denticles, heaviest above. 

Length 7.50, diameter 4.25, spire 2.50, aperture 4.50 
mm. 


Of much the same size and general characters as S. s¢zcola of 


303 Costa Rico MiocENE—OLSSON 13k 


Costa Rica, but with a larger and broader body-whorl, and a 
shorter, more conic spire. ‘The porterior portion of the inner 
and outer lips are heavily calloused, a feature not developed in 
S. sincola. 


Gatun Stage: Water Cay. 


Genus MUREX Linnzus 


Murex messorius Sowerby 


Murex messorius Sowerby, 1840, Proc. Zool. Soc. London, p. 137. 


Murex rvecurvirostris Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 
201. Not J. recurvirostris Broderip. 


Murex messorius Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila. 
vol. 63, p. 353. 

Murex messorius Maury, 1917, Bull. Amer. Pal., vol. 5, p. 265, pl. 16, 
figs. 1, 2. 

This is the common Murex in Costa Rica. It is generally 
small, seldom exceeding 40 mm in height. Each whorl carries 
three primary varices, between which are 3 or 4 smaller ribs or 
costz. It is arecent species in the West Indies. 


Cammy Stage: Gatun, (GZ. 
Water Cay, Panama. 
Zone 5, Red Cliff Creek. 
Coll. 7, Pumbri Creek. 


Banana River. 


Subgenus PHYLLONOTUS Swainson 


Murex cornurectus Guppy 


Murex (Chicoreus) megacerus Gabb, 1873, Trans. Amer, Phil. Soc., 
vol. 15, p. 202. Not WZ. megacerus Sowerby. 

Murex cornurectus Guppy, 1876, Quart. Journ. Soc. London, vol. 32, 
(OS) aig jolly keh pavers ite 

Murex (Phyllonotus) cornurectus Maury, 1917, Bull. Amer. Pal., vol. 
St DOV pI DOs Sey TO: 


132 BULLETIN 39 304 


A large species, possibly identical with the recent Murex 
brevifrons Lamarck. ‘The varices are large and cross the spire 
whorls in a spiral direction. Between the varices, there is gen- 
erally a single rib-like elevation in the middle of the whorl. 


Gatun Stage: Coll. 4, East Grape Point Creek. 
Coll si ibed \Gligy Greer: 
Rio Betey. 
Soust Creek. 


Genus TYPHIS Montfort 
Typhis alatus Sowerby Plate 10, figure 15 


Typhts alatus Sowerby, 1849, Quart. Journ. Geol. Soc. London, vol. 6, 
p. 48, pl. 10, fig. 4. 

Typhis alatus Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 203. 

Typhis alatus Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 32, 
Pp. 522. 

Typhis alatus Brown and Pilsbry, 1911, Proc. Acad. Nas. Sci. Phila., 
vol. 63, p. 354. 

A very characteristic species distinguished by the large wing- 
like expansion of its outer lip. It is the largest species in the 
Gatun and measures about 23 mm in height by 17 mm in diame- 
ter. Itisfound also in the Miocene of Santo Domingo and in 
the Bowden beds of Jamaica. 


Gatun Stage: Gatun, C. Z. (Brown and Pilsbry) 
Toro Cays, Panama. 


Typhis linguliferus Dall, var. costaricensis, n. var. Plate r0, figs 22, 29 


cf. Zyphis linguliferus Dall, 1890, Trans. Wagner Free Inst. Sci., vol. 

By EP ta Ps r5 25 ployee mio 
This is a smaller and more delicate speciesthan the 7. Gabdz 
Brown and Pilsbry from the Canal Zone. They agree with the 
Chipolan “inguliferus Dall in their general form but differ in their 
uniformly smaller size and in nearly lacking the spine-like pro- 


305 Costa RicA MrocENE—OLSSON 133 


cess on the shoulder of primary varices. It is fairly abundant in 
the Gatun beds of the Banana River. 


Gatun Stage; Ffill ra, Banana River. 


Genus S!STRUM Montfort 
Sistrum nodulosum C. B. Adams Plate 10, figure 23 


Purpura nodulosa ©. B. Adams, 1845 Proc. Bost. Soc. Nat. History, 
Dens 

Ricinula nodulosa Tryon, 1880, Manual of Conch., vol. 2, p. 190, pl. 
59, fig. 275. 

Sistrum nodulosum Dall and Simpson, tgo1, Bull. U. S. Fish. Com., 
VOI Te Penal. 

Worn specimens of the species occur in the transitional beds 
of Red Cliff Creek. They cannot be separated from recent ex- 
amples from the present coast. It is very common as a recent 
species on the rocks and reefs along the northern Costa Rican 
and Panama coast, where it occurs about high-water level, asso- 
ciated with the equally as abundant Lxgina turbinella Kiener 
and Phasianella affinis C. B. Adams. 


Uscari and Gatun transitional beds; Zone 5 and 6, 
Red Cliff Creek. 


(C) SUPER-FAMILY T4NIOGLOSSA 


Genus DISTORTRIX Link 


Distortrix similfima Sowerby 


Triton simillimus Sowerby, 1849, Quart. Journ. Geol. Soc. London, 
vol. 6, p. 48. 

Persona simillima Guppy, 1866, Quart. Journ, Geol. Soc. London, vol. 
22, p. 288, pl. 17, fig. 13. 

Distortia simillimus Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 
QU2 10% 

Fersona simillima Guppy, 1874, Geol. Mag., p. 439. 


134 BULLETIN 39 306 


Persona simillima Guppy, 1876, Quart. Journ. Geol. Soc, London, 
vol. 35, p. 522. 

Distorsio ( Distortrix, Persona) gatunensis Youla, 1908, Jahrb. der K-K 
Geol, Reichsanstalt, Wien, vol, 58, p. 700, pl. 25, fig. 10. 


Distorsio gatunensis Brown and Pilsbry, 1g1t, Proc. Acad. Nat. Sci. 
Phil., vol. 63, p. 356, pl. 29, fig. 8. 


Distortrix simillima Maury, 1917, Bull. Amer. Pal., vol. 5, p. 271, pl. 
17, Hes. 4,5. 
Not common in Costa Rica. It also occurs in the Miocene 
of Jamaica and Santo Domingo. 


Gatun, Stages Gatun, Ga Ze 
Banana River. 
Port Limon. 


Genus BURSA Bolten 
Bursa crassa Dillwyn Plate 15, figure 15 


Ranella crassa Dillwyn, Reeve, Conch. Icon.,fig. 18. 

Ranella crassa Guppy, 1866, Quart. Journ. Geol. Soe. London, vol. 22, 
p. 288, pl. 18, fig. 9. 

Bursa crassa Gabb, 1873, Trans, Amer. Phil. Soc., vol. 15, p. 212. 

Ranella crassa Guppy, 1874, Geol. Mag., p. 438. 

Ranella crassa Guppy, 1876, Quart. Jouru. Geol. Soc. London, vol. 32, 
Pen522: 

Bursa crassa Maury, 1917, Bull. Amer. Pal., vol. 5, p. 272, pl. 17 
figs. 6, 7. 

This is a recent species in the West Indies. It occurs also 

as a Miocene fossil in Jamaica and Santo Domingo. 


Gatun Stage: Banana River. 
Saury Creek. 


Genus CASSIS Lamarck 
Cassis sulcifera Sowerby 


Cassis sulcifera Sowerby, 1849, Quart. Journ. Geo 1. Soc. London, vol 
OF P47 supe ow sion i 


307 Costa Rica MiocENE—OLSSON 135 


Cassis sulcifera Maury, 1917, Bull. Amer. Pal., vol. 5, p. 274, pl. 18, 
TONERS, gh Ad ey 
This is a large species very common in Miocene rocks of 
Santo Domingo but very rare in Costa Rica. Our record is bas- 
ed on a single imperfect specimen from the Banana River and for 
a figure of this species the reader is referred to Dr. Maury’s Do- 
minican Fossils. 


Gatun Stage: Flill No. 3, Banana River. 
Cassis flamimea Linnaeus 


Buccinum flammeum Vinneeus, 1758, Syst. Nat., ed. 10, p. 736. 

Cassis filammea Reeve, 1848, Conch. Icon., vol. 5, pl. 5, fig. 12. 

Cassis flammea Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 2nd 
series, p. 356. 

This is a recent species recorded by Gabb, from his so-called 
Pliocene clay beds of Costa Rica. Under this term, Gabb con- 
fused two formations, one composed largely of coralline lime- 
stones but with subordinate amounts of clays and sands equiva- 
lent in large measure to the Gatun formations,and an overlying, 
mostly sand and clays of true Pliocene age. Itis most likely 
that the single example of Cassis fammea collected by Gabb, 
was obtained from the upper or Pliocene formation. 


Subgenus PHALIUM Link 


Phalium moniliferum Guppy Plate 12, figure 11 

Cassis monilifera Guppy, 1866, Quart. jouru. Geol. Soc. London, vol. 
224) pi 287, pla 17, fg. 8. 

Casidea granulosa Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 222. 
Not C. granulosa Bruguiere. 

Cassis reclusa Guppy, 1874, Geol. Mag., London. pp. 434, 439. 

Cassis reclusa Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 32, 
Pp. $25. 

Cassis monilijera Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 
32, P- 525. 

Phalium moniliferum Maury, 1917, Bull, Amer. Pal.. vol. 5, p. 274, pl. 
DOM MUSSe AS pl TOs) fol. 


136 BULLETIN 39 308 


The Costa Rican specimens like those of Santo Domingo are 
extremely variable in their sculpture. The typical mzonzleferum 
of Jamaica carries on the shoulder of the body-whorl one or two 
rows of large tubercles or granules and Guppy’s figure shows the 
spirals below as finely granulated. 


In Costa Rica, the shells may vary from those with nearly 
smooth spirals, through transitional forms in which only the su- 
perior bandsare granulated, to the extreme in which all the spirals 
are finely granulated over the whorl shell. A typical specimen 
will average: 


Length 35, diameter 22 mm. 


Gatun Stage: Middle Creek. 
Rio Betey. 
Lone 5, Red Cliff Creek. 


Genus SCONSIA Gray 
Sconsia laevigata Sowerby, var. Gabbi, n. var. Plate 12, figure 3 


ef Cassidaria laevigata Sowerby, 1849, Quart. Journ. Geol. Soc. Lon- 
don, vol. 6, p. 47, pl. Io, fig. 2. 

cf Cassidaria sublaevigata Guppy, 1866, /demm, vol, 22, p. 287, pl. 27, 
fig. 9. 

Cassidaria laevigata Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 
2nd series, p. 356. 


cf Sconsia sublaevigata Bose, 1906, Bol. Inst. Geol. de Mexico, Numero 
22, p. 36, pl. 4, figs. 9, Io. 

Sconsia laevigata Prown and Piisbry, 1911, Proc. Acad. Nat. Sci. Phila. 
vol. 63, p. 356. 

cf Sconsia laevigata Maury, 1917, Bull. Amer. Pal., vol. 5, p. 275, pl. 
19, fig. 2. 

As Gabb noted years ago,the Sconsias of Costa Rica are not 
the typcal Jaevigata of Santo Domingo but appear to approach 
more closely in their striated whorls, the Jameican sublaevigata 
_of Guppy and the recent s¢vzata of Lamarck. 


Typical /aevigata, has generally a distinct shoulder to its 


309 Costa Rica MioceNE-—-OLSSON Way 


body-whorl and a lower spire. The spire-whorls are striated but 
the spirals are lacking from the generally polished body-whorl. 
The aperture of /aevigata is subrectangular in outline, rather 
wide anteriorly and flaring posteriorly, while in Gaddz, the aper- 
ture is narrow and but little larger anteriorly. In daevigata, the 
callus of the inner lip is large and spreads widely over the colu- 
mellar area, but its outer edge is raised and shelf-like, instead of 
rounded and appressed as in Gaddz. Mature shells of laevigata, 
have three large, heavy varices, marking the resting stages but 
such varices are small and sometimes entirely lacking from the 
Costa Rican shells. 


Typical /aevigata will probably be found to be largely limit- 
ed to Santo Domingo. ‘The Jamaica sud/acvigata (possibly a var- 
iety) has more strongly sculptured whorls, a short spire and less 
complete varices. Its aperture is that of true /aevigata. 


The Sconsia laevigata was found by Dr. Maury, in her work 
on the Paleontology and Stratigraphy of Santo Domingo to be 
limited to her upper or Gurabo formation (the-Sconsza laevigata 
zone). In Panama and Costa Rica, the Sconsza laevigata variety 
Gabi, is frequently very abundant in the Gatun beds, but it is 
apperently lacking from the underlying Uscari shales where it is 
replaced by the new Scozsza cocleana and probably dbocasenszs. 


Carn Stage Gatun, CZ, 
Toro Cay, Water Cay, Panama. 
Comadre Creek near Cahutta C. R. 
TRAD: SEU er Tao (Os, SKE: 


Sconsia bocasensis, n. sp. Plate 12, figures 12, 13 


Shell like that of /aevigata,but narrow and with coarsely sculp- 
tured whorls; spire more evenly conic, the apex of which is not 
produced or attenuated; whorls coarsely sculptured with heavy, 
persistent spirals threads, of which there are about g on the spire- 
whorls and nearly 50 onthe last whorl; sutures appressed and 
bordered anteriorly by a strong ridge-like cord or by stronger 
spirals; the varices are but slightly developed; aperture subellip- 


138 BULLETIN 39 310 


tical, widest about the anterior 1-3; outer lip thickened and with 
about 18, fine, liree-like denticles ; the inner lip with a thin spred- 
ing callus and with about 16 long, narrow rugations. 

Length 35 (last 2 whorls only) diameter 25 mm. 

Differs from the variety Gadd of laevigata by its more narrow 
shell, coarser sculpture and larger aperture. The suture is bord- 
ered by a cord-like ridge. 

Gatun Stage: Bocas del Toro. 


Sconsia cocleana, n. sp. Plate 12, figure 7 


Shell globose; spire low of about 7 whorls, the apex slightly 
projecting; sutures distinct and deep, but not appressed; surface 
coarsely sculptured with large, irregular, but narrow spirals 
bands separated by deep interspaces which may in addition carry 
one or more finer spiral threads; there are 7 or 8 spirals on each 
spire whorl; on the last whorl the spirals are nearly twice as wide 
as the upper third of the whorl and more anteriorly; no varices; 
the outer lip is broken from the single specimen; inner lip is a 
wide but thin wash of callus, without rugations or denticles of 
any sort. 


Length 46, diameter 32 mm. 


The Uscari shales which everywhere in Costa Rica appear to 
underlie the Gatun sandstones is very rich in the smaller types 
of foraminifera, but mollusks are rare, fragmentary and difficult 
to collect. Fragments of Scozsta, were noted by us at several 
places which we believe belong to this species or to the docas- 
Ensts. 

his species will be recognized by its broad, globose shell 
and small spire. It lacks the sutural cord so strikingly develop- 
ed on docasensis. 


Uscari Stage: Rio Cocles near Old Harbor, C. R. 


gir Costa Rico MrocENE—OLSSON 139 


Genus DOLIUM Lamarck 


Subgenus MALEA Valenciennes 
Matea elliptica Pilsbry and Johnson Plate 12, figure 2 


Malea elliptica Pilsbry and Johnson, 1917, Proc. Acad, Nat. Sci. Phila., 
vol. 69, p. 169. 

The true Malea camura Guppy, so common in the Miocene 
of Jamaica and Santo Domingo still remains to be collected in 
Costa Rica. It is usually a smaller species (height 60 mm) with a 
heavier shell and high, narrow, special bands numbering about 
16 or 17 rather widely separated. 


The 1/. elliptica is larger (height 60-80 mm or more), more 
globose and thin. The spiral bands numbering about 22 are 
closely spaced and ribbon-like. The coiling is regular, while in 
camura the coiling is irregular and descending in the latter 
stages of growth. 


The Malea elliptica was described but not figured by John- 
son and Pilsbry from the Miocene of Santo Domingo. 
Height 63, diameter 54 mm. 
Uscart Stage: Red Cliff Creek, 
Gatun Stage: Bocas del Toro. Estrella River. 


fill ra, Banana River. 


Genus CYPRAEA Linné 
Cypraea parisimina, n. sp. Plate 12, figure 10 


Shell small, more or less depressed and with expanded, an- 
gulated sides; dorsal surface convex, but with a deep pit or 
depression at its posterior end, beyond which project the 2 
short, pouting ends of the lips of the aperture; ventral sur- 
face flat, with a narrow aperture which is straight except near 
at its posterior end where it is slightly curved; lips strongly 
but regularly crenulated (about 21 on each lip) posterior sinus 


140 BULLETIN 29 312 


small and curved to the left; anterior sinus small and round- 
ed. 


Length 39, basalt diameter 25, height or vertical diame- 
ter 17 mm. 
A small shell with strongly depessed and angulated sides. 
Collected from the coral limestones of Port Limon. 
Gatun Stage: Port Limon, 
Cpyraea cf. dominicensis Gabb 
Cypraea Dominicensis Gabb, 1873, Amer. Phil. Soc., vol. 15, p. 
236. 
Cypraea dominicensis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 280, 
pl. 19, fig. 11 
Our collection contains a single, fragmentary shell from the 
coralline limestones near Limon. ‘This specimen agrees closely 
with the Dominican shell figured by Dr. Maury as C. dominicen- 
sts ?, It is characterized by its very thin shell, cylindroid form 
and straight, evenly denticulated apertrue. 
Length 28, height 12 mm. 
Gatun Stage: Port Limon. 
Cypraea cinsrea Gmelin Plate 12, figure 8 
Cypraea cinerea Gmelin, 1792, Syst. Nat., p. 3404. 
Cypraea cinerea Reeve, Icon., vol. 3, pl. 22, fig. 124. 


Cypraea cinerea Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 2nd 
series, p. 360. 


The Cypraeca cinerea is a common, small species of Cowry 
shell along the north coast of Panama and Costa Rica. Several 
small Cypraeas were collected from the Gatun sandstones of 
the Rio Blanco which cannot be distinguished from recent ex- 
amples of cinerea. 

Gatun Stage: Rio Blanco. 


Cypraea almirantensis, n. sp. Plate 12, figure 9 


Shell large, heavy and rather high; the outlines of the shell 


313 Costa Rica MIocENE—-OLSSON 141 


from below, the side and from above is nearly rectangular, but 
with the anterior extremity more pointed; the shell is high (about 
¥% that of the length), with nearly flat or slightly convex sides; 
dorsal surface convex, a deep depression or pit about the posteri- 
or 1-4 and low, but large tubercles on each side of the posterior 
sinus; aperture narrow, curved, with the lips coarsely but regu- 
larly dentate (about 20 on each lip); posterior sinus is long and 
vertical; anterior sinus small and rounded; basal surface flat. 
Length 60, basal diameter 39, vertical diameter 30 mm, 

This is a species but distantly related to the C. Henkeni So- 
werby, differing by its longer and higher shell, straight sides and 
smaller dorsal tubercles. The aperture is strongly curved and 
very deep on the posterior extremity, ending in the long, vertical 
sinus. 

Gatun Stage: Water Cay, Panama. 


Genus STROMBUS Linnzeus 


Strombus gatunensis Toula Plate 13, figures 5, 6 


Strombus gatunensis Toula, 1909, Jahrb. der K-K Geol. Reichsanstalt 
Wien, vol. 58, p. 673, pl. 25, fig. 7. 

Strombus gatunensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. 
Phila., vol. 63, p. 355, pl. 26, figs. 3, 5. 

But two species of Stromdbus are known from the Gatun beds 
of Panama and Costa Rica, while five species occur in the synch- 
ronous Miocene strata of Santo Domingo and four in Jamaica. 
Other species may therefore be expected with further explora- 
tion work in Costa Rica. 

In S. gatunensis, the coiling of the whorls follow closely the 
shoulder angle so that the last whorl appears very large and the 
spire low but wide. Young shells occasionally have the shoulder 
armed with few, large tubercles which later become nearly cov- 
ered by the close coiling and appressed sutures. The last whorl 
is large, nearly smooth and with a rounded shoulder angle. 


Occasionally traces of the original coloration is preserved 


142 BULLETIN 39 BIA 


and consists of narrow, yellow V-shaped lines, the acute base of 
which are directed backwards or away from the aperture and lie 
along the center of the whorls. Similiar markings may be seen 
on Veatcht and Guppy* has described the same markings for 
pugiloides from Jamaica. Length 63, diam. 42 mm. 
Gatun Stage: Gatun (OF 
Water Cay, Panama. 
fleadwaters of Middle Creek, C. R. 


Strombus pugifoides Guppy Plate 13, figures I, 2, 3 

Strombus pugilis Guppy, 1866, Quart. Geol. Soc. London, vol. 22, p. 
287. 

Strombus pugilis Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.- 
233" 

Strombus pugiloides Guppy, 1874, Geol. Mag., new series, vol. I, p. 
433. 

Strombus pugilis Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, p. 
340. 


Strombus pugilis Bose. 1906, Boll. de Inst. Geol. de Mexico, numero 
22, p. 35, pl. 4, figs. 1-6. 

Strombus pugiloides Maury, 1917, Bull. Amer. Pal., vol. 5, p. 284, pl. 
20, fig. 6. 

A smaller species than the gatunensts, from which it also 
differs by its longer spire and more strongly sculptured shell. 
The coiling follows below the shoulder of the whorls, which is 
either simply rounded or bears several large, spine-like tubercles 
asin the recent pugilzs. Resting stages develop large, hump- 
like areas irregularly distributed on the spire-whorls. Varieties 
occur which may be spirally sculptured over the whole shell or 
the last whorl may be largely smooth. ‘The original coloration 
is still preserved on some shells and consist as in gatumnenszs of 
narrow, yellow, V-shaped lines, the apex of which lie about the 
middle of the shell and directed backwards. It is very distinct 
from the recent West Indian pugz/7s,which does not appear to de- 
scend below the Pliocene. 


* Proc. Sci. Assoc. Trinidad, 1873, vol. 2, p. 82,(reprint: Bull. Amer 
Pal., vol. 8, p. 214). 


OLSSON 143 


315 Costa RicA MIocENE 


Uscari Stage: Sapote, C. R. (Gadd) 
Gatun Stage: Banana River. 
Zone FE, Saury Creek. 
Zont 5, East Grape Point Cretk. 


Genus CREPITACELLA Guppy 
Crepitacella limonensis, n. sp. Plate 15, figure 17 


Shell small, melanoid in form,with a large body-whorl and 
shorter conic spire; nucleus of 2 small, smooth whorls followed 
by 5 post-nuclear whorls; sutures distinct; the whorls are slight- 
ly coronated and crossed by oblique and slightly curved riblets 
which become headed on the coronate edge of the spire-whorls 
just below the upper sutures; on the last whorl, the ribs do not 
pass below the middle; the surface is covered with faint spiral 
threads, most distinct on the base of the body-whorl; aperture 
elliptical, with a thin, oblique outer lip and a subobsolete anter1- 
or canal; pillar straight. 

Height 8, diameter 4.5, aperture 4 mm. 

The Costa Rican shell, although possibly not mature,is much 
smailer than the C. cepula Guppy from the Miocene of Venezuela 
and Santo Domingo of the same number of ribs. It differs also 
in having but 2 nuclear whorls, while cepuda has 3. The corona- 
tion of the whorls which is such a striking feature of the Domin- 
ican shell, is very much less and inconepicuous. The longitud- 
inal costee are more numerous and produce a fine circle of beads 
as they cross the small coronate edge of the spire-whorls. 

Gatun Stage: Port Limon, 

Boras del Toro. 


Genus CERITH!UM Adanson 
Cerithium costaricensis, n. sp. Plate 10, figure 28 


Shell with a long, slender spire of about 12 whorls (tip brok- 
ed); sutures deep but somewhat appressed; the profile of the 


144 BULLETIN 39 316 


spire-whorls is convex with an occasional hump-like resting mark; 
the sculpture consists of primary spiral threads between which 
lie 1, 3, or more finer secondaries; the earlier spire-whorls have 
heavy ribs but they become replaced on the later by simple gran- 
ulations on the primary spirals; a narrow, sloping band borders 
the anterior side of the suture and carries finer, nou-beaded spiral 
threads: below this band, the later spire-whorls bear about 4 
beaded, primary spirals and finer secondaries; the last whorl shows 
6 primaries together with other spirals on the base and canal; 
aperture subcircular, with a moderately expanded outer lip, obli- 
quely produced anteriorly. 
Height 28, diameter 10.5, aperture 8 mm. 

The general form and sculpture of this species is that of the 
recent West Indian C. eburneum Bruguiere. ‘The fossil shell is 
larger, more slender and differently sculptured. 


Gatun Stage: Coll. 6, Red Cliff Creek. 


Genus CLAVA Martyn 
Clava costaricana, n. sp. Plate 10, figures 3, 4 


Shell of medium size, cerithoid; whorls about 15, the early 
spire-whorls very small; the last whorl carries a large hump on 
the ventral side of the shell to the right of the aperture, so that 
the shell appears flattened on this side; the sculpture consists of 
strong spiral cords which are coarsely granulated by small, close- 
ly spaced longitudinal cost; the upper spiral cord bordering the 
suture is somewhat larger than the other; there are 3 spirals be- 
tween the sutures of the spire-whorl and 6 on the last whorl with 
additional smaller ones on the base of the shell; the outer lip is 
broken on all our specimens; the columella carries 2 strong 
folds. 

Height 40, diameter 15 mm. 
Al, I4 mm. 

This species bears some resemblance to the C. caloosaénsts 

Dall from the Pliocene of Florida. The Costa Rican shell is 


317 Costa Rica MiroceNE--OLSSON 145 


very much larger, broader and the longitudinal costze are more 
rib-like, continued across the whorls, crossing the interspaces as 
well as the spiral cords. 


Gatun Stage: Banana River. 
Clava alajuela, n. sp. Plate 10, figure 30 


Shell small, slender; whorls 12 plus, the apex broken; the 
sides of the whorls are straight and form the narrow, tapering 
spire; sutures indistinct; the sculpture consists of nearly equal 
ribs and spiral threads; the early spire-whorls have 3, equal, 
spiral threads, separated by interspaces as wide as the spiral 
threads and deep; on the later whorls fine, intermediate threads 
appear in the interspaces and a fourth large spiral in the lower 
suture; the spirals are crossed and strongly beaded by straight or 
slightly curved, longitudinal ribs, which number on the last 
whorl about 25 or 24; heavy and thickened resting marks ap- 
pear on the later whorl on each volution; the aperture is broken, 
the columella straight and witha single small fold. 


Height 17 mm, diameter 6 mm. 
A small and finely sculptured species, represented by the 
single type specimen. 
Gatun Stage: Middle Creek. 


Genus SERPULORBIS Sassi 


Serpulorbis papulosa Guppy Plate 12, figure I 


Vermetus papulosus Guppy, 1866, Quart. Journ. Geol. Soc. London, 
vol. 22, p. 292, pl. 17, fig. 3. 
Vermetus papulosus Guppy, 1876, Quart. Journ. Geol. Soc. London, 
vol. 32, p. 519. 
Sespulorbis papulosa Maury, 1917, Bull. Amer. Pal., vol. 5, p. 291, pl. 
22, fig. Io. 
Considerable variation may be seen in the Costa Rican shells, 
from specimens in which the sculpture is quite coarse and with 
the interspaces between the main cords carrying only one strong 


146 BULLETIN 39 318 


intermediate cord, to more delicately sculptured forms with finer 
tuberculated cords separated by wide and finely sculptured inter- 
spaces. 


This species occur also in the Bowden beds of Jamaica and 
in the Gurabo formation of Santo Domingo. 


Gatun Stage: Fiill 1, 2, Banana River. 


Genus PETALOCONCHUS H. B. Lea 


Petaloconchus sculpturatus, H. C. Lea Plate 14, figures 10, 15 
g 


Petaloconchus sculpturatus A. C. Lea, 1845, Trans. Amer. Phil. Soc., 
Vol./o,)\p-)2230 pl) 34 hig. ia! 

Petaloconchus domingensis Sowerby, 1849, Quart. Journ. Geol, Soc. 
London, vol. 5, p. 51, pl. 10, fig. 8,a be. 

Patalocouchus sculpturatus Gabb, 1875, Trans. Amer. Phil. Soc., vol. 
25, Pp. 240. 

FPataloconchus sculpturatus Guppy, 1876, Quart. Journ. Geol. Soc. Lon- 
don, vol. 32, p. 5109. 

Vermetus (Petaloconchus) sculpturatus Dall, 1892, Trans, Wagner 
Free Inst. Sci., vol. 3, pt. 2, p. 305. 

Vermetus (Petaloconchus) pulcher Bose, 1906, Bol. Inst. Geol. de Mex- 
ico, Numero 22, p. 32, pl. 3, figs. 22, 23. 

Petaloconchus domingensis Brown and Pilsbry, 1911, Proc. Acad. Nat. 
Sci. Phila., vol. 63, p. 359. 


Petaloconchus domingensis Pilsbry and Brown, 1917, Proc. Acad. Nat. 
Sci. Phila., vol. 69, p. 37. 


Pctaloconchus domingensis Maury, 1917, Pull. Amer. Pal., vol., 5, p. 
2O2s Pl, 22s) ite ME. 

Pataloconchus domingensis Hubbard, 1920, New York Academy of 
Science! Scei)(Sur. Py Ray voluaa pt 2 ip ail39: 

The Antillean fossil was described by Sowerby as domingen- 
sis. It has been by some writers, considered synonymous with 
sculpturata, described a few years earlier by H. C. Lea from the 
Chesapeake Miocene of the eastern United States, while others 
have given domingensis the rank of a full species. A close com- 
parative study of the Costa Rica fossils with several sets of typi- 


319 ' Costa Rico MrocENE—OLSSON 147 


cal sculpturata reveals no distinction between them. Both are 
variable in their coiling from closely wound, narrow cylinders to 
more open, umbilicate coils. The sculpture is much the same. 


In the United States, this fossil is limited to the Upper 
Chesapeake Miocene. In the Antillean and South American re- 
gions, itis found in the Bowden beds of Jamaica, the Gurabo 
formation of Santo Domingo, the Quebradillas limestone of Por- 
to Rico, the Springdale beds of Trinidad, fromm the Gatun beds 
near Cartagena, Colombia and Mexico. 

Gatun Stage: Gaiun Stage, GC.) Z. 

Water Cay, Panama. 
Coll. 4, 5, East Grape Point Creek. 
Comadre Creek. 


Genus SILIQUARIA Lamarck 


Siliquaria modesta Dall, var. limonensis, n. var, Plate 12, figure 4, 5, 6 


ef Siliquaria modesta Dall, Mus. Comp. Zool., vol. 9, p. 39. 


ef Siliguaria modesta Dall, 1889, Bull. Mus. Comp. Zool., vol. 18, p. 
260, pl. 26, fig. 4. 

This shell is very abundant in certain marly and sandy beds 
in the coralline limestones of the Limon Peninsula and elsewhere. 
Specimens in perfect condition are very difficult to obtain, the shell 
being very fragile and delicate. There are no longitudinal threads 
and the shell is smooth, except for the growth lines which are 
sometimes a little irregular. 


We have associated this shell with the .S. szodesta of Dall, a 
recent species and recorded by Dr. Dall from several stations in 
the West Indies and the Gulf of Mexico, in waters ranging from 
94 to 805 fathoms in depth. The fossils differ in being more 
loosely coiled and at the later stages long drawn out. 


Dr. Maury’s S. guarabensts from the Miocene of Santo Do- 
mingo is more irregular in its habitus of growth and the surface 
of the shell carries faint longitudinal threads. Gabb has des- 


148 BULLETIN 39 320 


eribed as sculpturata a Siliquaria from the Limon Peninsula. 
This is united by Dall with the recent West Indian sguamata 
Blainville. I have seen no spicimens and Gabb may have collect- 
ed it from the true Pliocene beds of the Limon Peninsula. 
Gatun Stage; Island of Bocas del Toro. 
Port Limon. 


Genus TURRITELLA Lamarck 
Turritelia gatunensis Conrad Plate 14, figures 12, 13 


Turritella gatunensis Conrad, 1857, Pacific R. R. Report, vol. 6, p. 72, 
PLS whe20: 

Turritella conradi Toula, 1909, Jahrb. der K-K Geol. Reichsanstalt, 
vol. 58, p. 694, pl. 25, fig. 4. 

Turritella gatunensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci- 
Phila., vol. 63, pl. 27, figs. 4, 5, 9. 

The Zurritella gatunensis is a medium sized shell seldom 
over 40 mm in length. The base of the whorl overhangs the next, 
and each whorl is encircled by a median concave or constricted 
band. The whole surface it finely sculptured with small spiral 
threads. The spirals defining the median band are usually slight- 
ly heavier as well as a few about the upper part of the base. 


The 7. gatunensts is a common and widely distributed 
species in the Gatun beds, but always less abundant than the 
T. altilira or its varities. Like the altdliva, zones of 7. gatun- 
ensis are frequent in some localities. 


The 7. atacta Dall of the Tampa Silex beds, the 7. acropo- 
ra fossilin the Pliocene of Florida and recent along the east 
coast of United States and the West Indies, are related 
species. 

Gatun Stage: Gatun, C. Z. 

Water Cay, Panama. 
Boucary Creek, C. R. 


32K Costa RicA MrocENE—OLSSON 149 


Turritella mimetes Brown and Pilsbry Plate 14, figure 5 


Turritella mimetes Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. 
Phila., vol. 63, p. 357, pl. 27, fig. 1. 


Turritella (Haustator) aff. Hanleyana Reeve- 7. lineolata (Kiener) 
Toula, 1911, Jahrb. der K-K Geol, Reichsanstalt, vol. 61, p. 
401, pl. 30, figures 6a and 6b, 
A large, robust species in form and sculpture like the 7. 
variegata Linnzeus, recent in the West Indies. 


The whorls are straight and usually slightly overhanging 
the lower sutures. Surface sculptured with primary, secondary 
and tertiary spiral threads. The primary threads are irregularly 
disposed and usually number about 7 or 8 to each whorl. Be- 
tween the primaries are the finer secondaries and tertiaries. 


Small shells may be mistaken for the gazunensis, but the 
whorls are flat, without the median concave and constricted zone 
and the sculpture is more coarse. The figured shell of 14 whorls 
has a length of 70 mm and a diameter of 18 mm. A larger but 
more imperfect specimen of but 4 whorls measures: 

Length 50, diameter 20 mm. 


Gatun Stage: Gatun, C. Z. 
Turritella altilira Conrad, and varities. 


Turritella attilira Conrad, 1857, Pacific R. R. Reports, vol. 6, p. 72, 
pl. 5, fig. 19. 

Turritella gabbi Toula, 1909, Jahab. der K-K Geol. Reichsanstalt, p. 
695, pl. 25, fig. 5. 

Turritella altilira Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. 
Phila., vol. 63, p. 358, pl. 27, figs. 2, 3. 

Of the species of 7urritella in the Gatun beds of Costa Rica 
and Panana, the 7. adé/zra of Conrad and its varieties is the most 
common and characteristic. The 7. fornata Guppy, from the 
Miocene of Trinidad and Venezuela is a closely related species. 


These magnificent 7urrztelle are preminently characteristic 
of the West Indian and Caribbean Miocene, but probably deriv- 
ed from earlier forms in the Oligocene. The stock continued in- 


150 BULLETIN 39 322 


to the Pliocene where its acme of development was reached in 
the elegant 7: preattenuata Heilprin of Florida. It has left no 
descendants in the recent fauna. 


The shell is long and tapering, each whorl sculptured with 2 
prominent beaded spiral cords, between which the surface is con- 
cave or deeply channeled and may be smooth or with finer 
spiral threads. The upper spiral cord is simple or double. Several 
varieties may be recognized which differ in details of their sculp- 
ture, size and form of shell. 


T. altitira Conrad, typical Plate 14 figures 6, 7 


This, the most elegant form is limited to the small Gatun 
sedimentary basin of the Canal Zone and the adjacent Province 
of Colon. 


The shell is long, slender and with numerous whorls; each 
whorl is strongly sculptured by the two, primary spiral cords, 
each of which is bordered on adjacent faces by a smaller 
spiral, between which lies a deeply channelled groove; small- 
er beaded spirals occur in the median channel and are best devel- 
oped on the spire whorls; the beading or granulation of the 
spirals is produced by the arcuated lines of growth, the sinus of 
which lies a little above the middle of the shell; the upper spiral 
cord is larger than the lower and more strongly beaded and in 
general becomes double on the later whorls. 


Gatun Stage: Canal Zone and Province of Colon. 
Var. chiriquiensis, n. var. Plate 14, figures 4, 8, 9, 14 


Shell larger, less attenuated and more coarsely sculptured 
than typical altliva; the 2 spiral cords are less strong, of more 
nearly the same size and usually remaining simple; the 2 spiral 
cords enclose a wide, concave area of about % the width of the 
whorl; this concave band usually carries 2, 3, or more fine spiral 
threads; the beading or granulation of the spiral cords is feeble 
and best developed on the earlier spire whorls and in certain large, 


323 Costa RicA MiocENE—OLSSON 151 


gerontic shells the spiral cords may become nearly smooth. 


This is the common form on the islands of the Chiriqui La- 
goon, and of eastern Costa Rica. Like the typical a/tzlira it is 
often a very abundant fossil, forming zones of Turritella in the 
Gatun beds. 


The sculpture is much less elegant than that of true al¢zzra, 
and consists mainly of the 2, primary spiral cords, between which 
lies a wide concave band. This band is but feebly sculptured by 
small, spiral threads, or in some cases, is smooth. 


In small matters of detail of sculpture and in the form and 
taper of the shells, there is considerable variation. Such varia- 
tion is usually of but local signifieance and in general the shells 
collected at the same or nearby localities will agree closly with 
each other but differ more or less from those of more distant 
areas. 


The 7. fornata Guppy from the Miocene of Venezuela and 
Trinidad should probably be considered asa variety of alézlzra, 
most closely allied to the chirzguensts. Its sculpture is like chzr- 
Zquensts but with stronger spiral threads on its median band. 


Gatun Stage: Water Cay. 
Grape Point and Red Cliff Creeks, C. R. 
Banana River. 
Rio Blanco. 


Var. costaricensis, n. var. Plate 14, figure II 


Shell small; whorls less numerous and more rapidly increas- 
ing so that the shell is porportionately broader and shorter; su- 
tures deep; the 2 primary cords are but weakly developed, be- 
tween which lies, the concave band as in chiriquiensis,; the band 
carries 3, 4, or more strong, irregular spirals, but slightly weak- 
er than the primaries; the granulation of the cords is weak and 
irregular. 

Much smaller than the preceding varieties (usually about 4o 
mm) and with wider and more rapidly increasing whorls. The 


152 BULLETIN 39 324 


cords on mature shells are small and but slightly heavier than the 
spiral threads on the concave band. 


This type replaces the chiriguiensis in the Gatun sandstones 
in the area of Upper Hone Creek and Boucary Creek. It is often 
extremley abundant, associated with the Turritella gatunen- 
STS. 

Gatun Stage: Upper Hone and Boucary Creeks. 


Turritella exoleta Linnzeus, var. limonensis, n. var. Plate 14, figs 2, 3 


Shell turreted, the whorls increasing more rapidly in diame- 
ter than typical evolefa; the nucleus is small, immediately follow- 
ed by whorls carrying a median carina which gradually becomes 
more anterior in position and later the lower carina of the adult 
sculpture; the whorls are deeply concave or excavated about the 
middle between 2 principal carinze; on the last whorl there is 
seen to be a third carina which is concealed in the suture; the su- 
tures are excavated, becoming generally deeper with age; the 
surface is nearly smooth, except for very faint spirals and rough 
growth lines which may be raised and sublamellose on the medi- 
an concave zone. 

Height 4o (6 later whorls) diameter 17.50 mm. 
Height 35 (10 earlier whorls) diameter 10 mm. 

This is probably a distinct species from the recent West In- 
dian Turritella exoleta Linnzeus, differing in its more rapidly ex- 
panding whorls. Faint spirals may generally be seen covering 
the entire shell and the growth lines become raised and sublamel- 
lose in the concave medial zone. It is acommon shellin the cor- 
alline phase of the Gatun at Port Limon and elsewhere. 


Gatun Stage: Port Limon. 
Bocas del Toro. 


Turritella oreodoxa, n. sp. Plate 14, figure 1 


Shell large, solid and very slender; whorls numerous and 
each but slightly larger than the preceding, slightly concave or 
constricted about the upper 1-3, above which the edge of the 


325 Costa Rica Mrocene#—OLSSON 153 


whorl is enlarged, forming a cord-like expansion about the upper 
suture; sutures indistinct and appressed; sculpture of fine, ir- 
regular and somewhat wavy spiral lines and indistinctly alter- 
nating in strength; growth lines as typical for Haustator (not 
visible on the base of our specimen), commence at the lower 
suture and pass straight and parallel to the longitudinal axis 
of the shell to the middle of the whorl where they become 
bent™and continue obliquely to the left or towards the aper- 
ture to the upper suture. 
Length (2% whorls), 49, diameter 23 mm. 

A very rare species and represented in our collection by 
only two imperfect specimens. It appears to be closely related 
to the 7. cartagensis Pilsbry and Brown from the Gatun of 
Colombia. The full-grown Costa Rican shell was probably 
longer and more attenuated and each whorl is prominently 
elevated or ridged just in front of the lower suture. 


Gatun Stage: Rio Banana beyond Hill No. 5. 


Genus ALABA A. Adams 


Alaba turrita Guppy, 1896, Proc. U. S. Nat Mus., vol. 19, p. 321, pl. 
28, fig. 7. 

Guppy’s figure of his type specimen from Jamaica agrees 
very closely with our specimen from the Estrella River. The 
whorls are finely spiralled with low ridges which are obsolete 
from the upper parts of the whorls. Each whorl carries 2 or 
more broad, smooth varices formed during resting stages. 
The last whorl is strongly angulated by these varices. 


Height 5, diameter 2.25 mm. 
Gatun Stage: Coll. 7, Estrella River. 


154 BULLETIN 39 326 


Genus ARCHITECTONICA Bolton 


Architectonica granulata Lamarck Plate 13, figure 10 ,1I, 12 


Solarium granulatum Wamrack, 1822, An. sans. Vert., vol. 7, p. 3, 


Architectonica prespectiva Tuomey and Holmes, 1857, Pleioc. Fos. S. 
Ce pe i2ouple26s sige: 

Solarium granulatum Dall, 1892, Trans. Wagner Free Inst. Sci., vol. 
BPE Pa sZOe 

Solarium Villarelloi Bose, Inst. Geol. de Mexico, Numero 22, p. 30, 
pl. 3, figs. 4-11. 

Solarium gatunense Toula, 1908, Jahrb, der K-K Geol. Reichsanstait, 
Wien, vol. 58, p. 693, pl. 25, fig. 3. 

Solarium granulatum Maury, 1917, Bull, Amer. Pal., vol. 5, p. 295, 
pl. 23, fig. 3. 


The Costa Rican fossils are usually quite typical and not 
to be distinguished from recent examples of granulata. ‘They 
vary somewhat in their height and persistence of their spiral 
sculpturing. The beaded and granulated character of the spir- 
als generally becomes obsolete on the later whorls so that the 
spiral bands appear smooth. The umbilicus is deep but nar- 
row. 


This is a stable species ranging from the lower Miocene 
to the recent. Its recent distribution extends from Cape Hat- 
teras on the north, southward through the West Indies and on 
the Pacific side from the Gulf of California to Peru. 


Height 20, greater diameter 43 mm. 
19, 29 mm. 
12.75 20 mm. 
Gatun Stage: Gatun, C. Z. 
Water Cay, Panama. 
Goll; 2 "RaolGocles: 
Banana River. 


Rio Betey. 


327 Costa Rico MIOCENE——OLSSON 155 


Genus NATICA Scopoli 
Natica canrena Linnzeus Plato 13, figure 9 


Natica canrena Vinneeus, 1758, Syst. Nat., ed. 10, p. 776. 


Natica canrena Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 2nd 


series, p. 357. 

Natica canrena Brown and Pilsbry, 1912, Proc. Acad. Nat. Sci. Phila., 
p. 508, 

Natica canrena Maury, 1917, Bull. Amer. Pal., vol. 5, p. 298, pl. 23, 
fig. Io. 


This is a recent species but also common as fossil in the Mi- 
ocene of the West Indies and the States. It is a warm water 
species so that in the Chesapeake Miocene of the eastern United 
States, it is found only in the warm, upper or Duplin stage. We 
collected several specimens at Gatun but none at Costa Rica. 
Gabb however has recorded it as abundant from the Limon Pen- 
insula but whether his specimens were obtained ,from the Mio- 
cene or Pliocene is not known. 


The species is distinguished by its strong, tangential, su- 
tural plice, but the main surface of the whorl is otherwise 
smooth. 


Gatun Stage: Gatun, C. Z. 
Natica Youngi Maury, var., cocleana, n. var. Plate 13, figure 8 


cf. Natica Youngi Maury, 1917, Bull. Amer. Pal., vol. 5, p. 29y, pl. 
22h ISS wells 2s 

Shell subglobular with 5 rounded and convex whorls; sur- 
face smooth but with irregular, coarse growth lines about the su- 
tures; the surface is also faintly straited with weak spiral lines 
as may be seen on several otherwise smooth Naticas; aperture 
semilunar with a narrow callus on the inner lip, slightly thicker 
about opposite the middle of the umbilicus; umbilicus small, 
with the umbilical callus small and but poorly developed. 


Length or height 25, diameter 34 mm. 


156 BULLETIN 39 328 


A single specimen from the Gatun of Cocles Creek. It agrees 
almost exactly with Dr. Maury’s type specimen of Watica 
Young? from the Miocene of Santo Domingo in size and form, 
but differs in its smaller umbilicus and smaller umbilical callus. 
The Costa Ricam shell has the aspect of a Meverita, 


Gatun Stage; Cocles Creek, near Old Harbor, C. R. 


Section STIGMAULAX Moerch 


Natica Guppyana Toule Plate 13, figures 13, 14, 15 


Vv 


Natica (Stigmaulux) Guppyana Toula, 1909, Jahrb. der K-K Geol. 
Reichsanstalt, Wien, vol. 58, p. 696, pl. 25, fig. 6. 

Natica Guppyana Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. 
Phila., vol. 63, p. 360. 

This species stands intermediate in its sculpture and form 
to the 4. canrena Linneeus and JV. sulcata Born, both recent and 
fossil Miocene species. In Guppyana, the sculpture consists of 
even, wide and deep sulcations, which cross the face of the whorl 
from the upper suture to the umbilicus. They are in fact the 
continuation of the tangential sutural plicee of VV. canrena. With 
further growth these sulcations may become obsolete on the mid- 
dle of the whorls until they are lacking from the greater part of 
the shell except in the immediate vicinity of the upper sutures 
and on the umbilical angle. This change in sculpture is not seen 
to the same extent on all shells and some spetimens may remain 
strongly grooved and sulcated even when large and mature. In 
other cases (in general with shells from the Banana River) 
only very young shells have the sulcation continuous across 
the face of the whorl, but soon become smooth, leaving the plica- 
tions only about the upper suture and on the umbilical angle. 
Such shells resemble closely MV. canrena, but in addition to 
the grooving of the umbilical angle, have a larger and more 
exanded aperture. 


The WV. sulcata Born still remains to be found in Costa Rica 
and Panana. It is abundant as a Miocene fossil in Santo 


329 Costa Rica MiIocENE—OLSSON 157 


Domingo, Jamaica and Venezuela. It differs from the Guppyana 
by its closer and more irregular longitudinal sculpture crossed by 
faint spirals, producing a cancellated appearance. The umbili- 
cus is larger and more ample. 
Gatun Stage: Gatun, C. Z. 
Water Cay, Panama. 


Banana River, C. R. 


Genus POLINICES Montfort 


Polinices subclausa Sowerby Plate 13, figures 16, 17 


Natica subclausa Sowerby, 1849, Quart. Journ. Geol. Soc. London, 
vol. 6, p. 51. 


Natica subclausa Guppy, 1866, Quart. Journ. Geol. Soc. London, vol. 
22, p. 290, pl. 18, fig. 8. 
Folinices subclausa Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. 
Philia., vol. 63, p. 560. 
Folinices subclausa Maury, 1917, Bull, Amer. Pal., vol. 5, p. 300, pl. 
23, fig. 14. 
This is a common fossil in the Miocene of Santo Domingo 
and Jamaica. The Costa Rican shells are small, the largest 
specimen in our collection measuring only 17 mm in altitute. 


When viewed from the dorsal side, the shell has a distinct- 
ly Nerita-like aspect, due to its low spire, close coiling and rap- 
idly enlarging whorls and appressed sutures. The umbilicus is 
of medium size, expanding below. ‘The inner lip is provided 
with a thick, heavy callus which joins above with the outer lip. 
At the upper edge of the umbilicus, the callus carries a deep, 
transverse groove, a most characteristic feature. 


Gatun Stage: Gatun, C. Z. 
flill No. 3, Banana River, C. R. 


Polinices Stanislas-Meunieri Maury Plate 13, figure 7 


Polinices Stantslas-meunierit Maury, 1817, Bull. Amer. Pal., vol. 5, 
Pp. 300, pl. 23, figs. 15, 16. 


158 BULLETIN 39 330 


The Banana River beds contain a large species of Polinices 
which appears to be this Dominican species. It differs from the 
recent mammtllaris Lamarck, of the Costa Rican and Panana 
coast by its much narrow umbilicus. Its surface is lined with 
very fine spiral threads, 

Young shells may be mistaken for the swbclansa which oc- 
curs in the same beds, and careful study is sometimes necessary 
for their separation. The present shell is less heavy, with a 
higher spire and more convex spire whorls. The umbilicus is 
much narrower, anda smaller callus on its inner lip which shows 
but faintly the transverse groove. The surface of sudclausa is 
smooth, but in Stazzslas-Meunteri very finely lined with minute 
spirals. 

Gabb has described from Sapote, Costa Rica, a Matica emi- 
nulotdes which in form resembles young shelis of the S¢anzslas- 
Meuntert. Wis figure shows a different shaped umilicus and no 
transverse groove on the callus of its inner lip. 

Length or aititute 44, diameter 38 mm. 
Gatun) Stage: ul ra 2 > sbanang liven. Cs Re 


Subgenus NEVERITA Risso 
Heverita nereidis Maury 


Neverita nereidis Maury, 1917, Bull. Amer, Pal., vol. 5, p. 301, pl. 23, 
figs. 17, 18. 

Our collection from ths Lower Gatun of Red Cliff Creek, 
contains a fragment of this very interesting species. The J. 
nererdts described by Dr. Maury, from the Cercado formation of 
Santo Domingo, is closely allied to the recent West Coast /V. 
glauca Humboldt. The Miocene \V. zeretdis is less broad, with 
a narrower umbilicus and a larger umbilical callus. 


Gatun Stages) Goll. ol Reds Gla Greer: | 6. fe. 


33% Costa Rica MrocENE—OLSSON 159 


SUB-ORDER SCUTIBRANCHIATA 
A. SUPER-FAMILY RHIPIDOGLOSSA 
Genus NERITINA Lamarck 


Section SMARAGDIA Issel 
Neritina viridemaris Maury Plates 15, figure 22 


Neritina viridis Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 242, 
Not NV. viridis Linneeus. 

Neritina (Smaragdia) viridemaris Maury, 1917,Bull. Amer. Pal., vol. 
5, p. 316, pl. 24, fig. 11. 

The Costa Rican specimens are a little smaller than typical 
viridemaris from the Miocene of Santo Domingo; but are identi- 
cai in other respects. They still retain their original coloration of 
a pale sea-green marked with straight and zigzag black lines. It 
is the Miocene precursor of the common, recent West Indian and 
Caribbean XV. vzrzdzs Linné. 


Gatun Stage-\Coll. 5, feed Clif Creer. 
Middle Creek. 


Genus CREPIDULA Lamarck 


Crepidula plana Say 
Crepidula plana Say, 1822, Journ. Acad. Nat. Sci. Phila., vol. 2, p. 
226, 
Crepidula plana Wall, 1892, Trans. Wagner Free Inst. Sci., vol. 3, pt. 
2, p. 358. 
Crepidula plana Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., 
vol. 63, p. 360. 

This common and well-known species has been recorded by 
Brown and Pilsbry from the Canal Zone. We have a single spec- 
imen from Water Cay, found in the interior of the specimen of 
Solenostetra Vaughani var. medioamericana. This species first 


160 BULLETIN 29 3324 


appears in the Lower Miocene and has continued into the re- 
cent fauna. 


Gatun Stage: Gatun (Brown and Pilsbry). 


Genus XENOPHORA Fischer de Waldheim 
Xenophora conchyliophora Born 


Trochus conchyliophorus Born, 1778, Mus. Caes. Ind., p. 333. 

Trochus agglutinans Lamarck, Anim. sans Vert., vol. 7, p. 14. 

Fhorus agglutinans Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, 
pi24n 

Xenophora conchyliophora Dall, 1892, Trans. Wagner Free Inst. Sci., 
vol. 3, pt. 2, pp. 360-362. 

NXenophora conchyliophora, Maury, 1917, Bull. Amer. Pal. vol. 5, p. 
297. 

One immature shell was collected from the upper Gatun 
beds of Old Man Sam Creek. It is one of the few species which 
has continued pratically unchanged from Upper Cretaceous times 
to the recent. It is rare asa recent shell along the northern 
Panama and Costa Rican coast. Gatun Stage. 


Genus PHASIANELLA Lamarck 
Phasianelfa mollis, n. sp. 


Shell small, solid, ovate with a short, blunt spire anda 
small narrow umbilicus; whorls about 4, somewhat convex: su- 
ture distinct; surface smooth and usually showing no features; 
in some cases, color markings are preserved as brown, wavy 
blotches of flammules and with very faint suggestions of fine re- 
volving lines which are so common on recent small Phasianella. 

Height 3.25, diameter 2.75, aperture 1.50 mm. 

The Phasianella punctata Gabb from the Miocene of Santo 
Domingo, has a longer spire and different color markings. It is 
muuch more like the recent P. ajfinzs than the Costa Rican mollis. 
P. mollis is related to P. umbilicata d’Orbigny but differs in its 
sculpture. 


BAZ CosraA Rica Miocknez—OLSSON 16% 


The smaller forms of Phastanedla, of which there are 2 or 3 
species along the northern Costa Rican coast, are often extreme- 
iy abundant on the rocks and reefs exposed to the heavy, pound- 
ing surf. There they are associated with a host of other gas- 
tropods, principal among which are several species of /Verzta, 
Fissurella, Purpura and Chitons. The fossil shells therefore like 
the preceding Weritina viridemaris and several others which we 
have already called attention to, have drifted from some near-by 
rocky coast. 


Gatun Stage: Coll. 5, Red Cliff Creek. 
Middle Creek. 


Genus TURBO Linnzeus 
Turbo saltus, n. sp. Plate 15, figure 12 


Shell perforate, ovate-conic; spire high of about 6 convex 
whorls, separated by deep, canaliculate sutures; the last whorl is 
large and perfectly convex in form; sculpture consists of irregu- 
lar, spiral cords or liree; the earliest spire-whorls are strongly 
carinated by a single, strong spiral, above'and below which the 
whorl is smooth; a second spiral appears above and the succeed- 
ing whorls of the spire have 2 principal spiral cords or lire, and 
g smaller ones; the last whorl has three spirals that are a little 
stronger and many smaller ones of different sizes; the spirals are 
smooth; aperture perfectly circular, a small, spreading callus on 
the body-whorl just above the perforate base. 

Height 17, diameter 15.5, aperture 7.5 mm. 

Among the recent species of Zurbo the saltus is related to 
the large 7. Spenglerianus Gmelin,on one hand and to the small- 
er J. jilosus Kiener on the other. The shell is perforate,in which 
character it approaches 7. /ilosus, but the sutures are canalicu- 
late or channeled as in the non-perforate 7. Spenglerianus. The 
sculpture consists of irregular, smooth spiral cords or lire of 
which 2 or 3 about the middle of the shell are a little larger 
than the others. 


162 BULLETIN 39 334 
Gatun Stage: Coll. 5, Red Cliff Creek. 
Genus ASTRALIUM Link 

Astralium brevispinum Lamarck, var. basalis, n. var. Plate 15, figs. 4, 5 


The Astralium brevispinum Lamarck is a recent species 
found living along the north coast of Panama and Costa Rica. 
It is most easily distinguished from the West Indian 4. long7spr- 
num Lamarck by having its columellar area brilliantly stained 
with scarlet. 


From upper Old Man Sam Creek in eastern Costa Rica, we 
collected a single specimen of an Astralium, very similiar in most 
of its characters to bvevispinum. It differs most importantly in 
having a more strongly sculptured base. This sculpture consists 
of a wide, flat band, inside of which are 3, heavy, nearly 
equal spiral cords. The whole surface is crossed by squamose 
lines of growth. 


Height 20, diameter 29.50 mm. 
Gatun Stage: Coll. 6, Old Man Sam Creek. 


Astralium caelatum Gmelin, variety 


Trochus celatum Gmelin 1792, Syst. Nat., p. 3531. 

Astralium celatum Pilsbry, 1888, Manual of Conch., first ser., vol. Io, 
p. 224, pl. 57, figs. 45, 46. 

A large Turboid shell related to the recent 4. celatum Gme- 
lin was collected from the coral limestones near Port Limon. It 
is so firmly imbedded in its rocky matrix that only a small! por- 
tion of its surface isexposed to view. Its surface sculpture is like 
that of celatum, but the radial costee on the upper part of its 
whorl are closely spaced and not far apart as in the recent shell. 
Its height measures 55 mm, diameter 54. 


Gatun Stage: Port Limon. 


335 Costa Rico M1ockENE—OLSSON 163 


Genus CHLOROSTOMA Swainson 
Subgenus NEOMPHALIUS Fischer 


Chlorostoma costaricensis, n. sp. Plate 15, figure 16 


Shell conoidal, solid, witha narrow, but deep umbilicus; su- 
ture distinct; the last whorl is strongly carinate or angled below; 
above this angle, the whorl is evenly convex, below, the base is 
nearly fiat; the sculpture consists of beaded spiral cords, there 
being on the last whorl above the angled periphery 7 or 8 cords; 
the interspaces are nearly as wide as the cords, and are finely 
etched by oblique lines of growth and finer spirals; on the base, 
the spirals are irregular, the strongest about the middle; there 
are 2 or 3 cords about the upper part of the umbilicus, but the 
interior wall is smooth. 


Height 10.5, diameter 16.5 mm. 


The transitional beds between the Uscari and the Gatun of 
Red Cliff Creek, contain in addition to the usual fauna, several 
species, as certain Cerithoids, Szstrum nodulosum, Modulus mo- 
dulus, Nitidella and the present species, whose normal habitat 
is on the rocks and reefs of the extreme, upper limit of the lit- 
toral zone. They are always much broken and worn as if they 
had been rolled and washed about by the surf on a rocky or sandy 
beach and then drifted out to the deeper and quiter waters off 
_ shore where depostion was going on. 


Although all of our specimens of C. costaricensis are very 
fragmentary and much worn, we have deemed it best to place 
the species on record, as this subgenus has been recorded only 
very doubtefully from beds older than the Pleistocene. 


Uscari Stage: Zone 6, Red Cliff Creek. 
Gatun Stage: Zone 5, Red Cliff Creek. 


164 BULLETIN 39 336 


Genus CALLIOSTOMA Swainson 
Calliostoma limonensis, n. sp. Plate 15, figures 8, 1 


Shell of medium size, conic; whorls about 8, flat or but 
slightly convex, so that the profile of the shell is a broad cone; 
sutures indistinct; the whorls are closely sculptured with fine, 
beaded spirals of primary and secondary orders; each spire-whorl 
has 7 or 8 primary spirals alternating with 1, 2 or 3 secondary; 
periphery sub-angular; base but slightly convex and closely spir- 
alled with flat or sub-obsoletely beaded spirals (about 20), umbil- 
icus narrow but deep, smooth within. 


Height 19, diameter 18.5 mm. 


Fairly abundant in the coral limestones along the shore west 
of Port Limon. Related to the recent C. jujubinum Gmelin, it 
differs by its more globose form and small, pointed attenuated 
apex. 

Gabb hasalso described a small Callzostoma from the Limon 
Peninsular, as C. Guppyana. This shell of 8 or 9 whorls meas- 
ures but 5 mm in height. It is a higher and more conic form 
with the whorls sculptured with beaded spirals. 


Gatun Stage: Port Limon. 


Calliostoma mancinella, n. sp. Plate 15, figures 9, Io 


Shell of medium size; whorls about 8, with very indistinct 
sutures and a coarse sculpture of beaded spirals; the spire whorls 
are flat sloping evenly from the small projecting nucleus to the 
broadly rounded periphery of the last whorl; sculpture of the 
spire-whorls consists of about 6, strongly beaded spirals the 
largest of which boundsthe upper sutures; between these spirals 
are scattered a few smaller ones, the lower half of the peri- 
phery and the base have about 12 additional spirals; these 
spirals are larger, more widely spaced and are smooth except 
the 4 or 5 around the umbilicus; the interspaces of the basal 
spirals are finely incised by minute longitudinal lines, corres- 
ponding to the growth lines; umbilicus narrow, but deep, 


337 Costa RrcA MroceENE—OLSSON 165 


smooth within; aperture rounded, with the outer lip strongly 
oblique. 


Height 15, diameter 21 mm. 


Several specimens of this well-marked species were obtained 
from the Gatun beds of eastern Costa Rica, in the vicinity of 
Manzanilla Point. In the Santo Dominican Miocene it is relat- 
ed to the C. Grabaui Maury, but the Costa Rican shell is much 
lower and more depressed. 


The C. Grabaui and the mancinella belong to the group 
which contains the large, elegant C. Sayana, described by Dall 
from deep water off the Hatteras coast. 


Gatun Stage: Coll. 4, East Grape Potnt Cree. 
fleadwaters of Middle Creek. 
Coll. 5, Red Cliff Creek. 
zr mile south of shore, along Old Man Sam 
Creek, (Veatch) 


Calliostoma castilla, n. sp. Plate 15, figure 6, 7 


Shell rather large, in form and sculpture resembling the C. 
phulanthropus of the Chesapeake Miocene of eastern United States, 
but umbilicated; whorls 5 plus (apex broken), flat and forming 
a broadly conic spire; periphery of the last whorl, subangular, 
due to a broad, cord-like ridge, which appears as if formed 
form an overlapping of the base; sculpture of the spire-whorls of 
fine, alternating and beaded spirals, there being about 13 on each 
whorl; the base is nearly smooth the spirals being nearly obso- 
lete or as broad low bands; the spirals about the umbilicus are 
slightly stronger and obscurely beaded; umbilicus deep and 
narrow. 


Height 20, diameter 25 mm. 


A large species with an angulated periphery, fine, beaded 
spirals above and a nearly smooth base. Buta single specimen 


166 BULLETIN 39 338 


was collected from the coral limestones near Port Limon, asso- 
ciated with the common /zmonensis. 


Gatun Stage; Port Limon. 


CLASS SCAPHOPODA 
Genus DENTALIUM Linnzeus 


Subgenus DENTAL!UM, s. s. 


Dentalium bocasensis, n. sp. Plate 15, figures 2, 3 


Shell rather larger, solid, gently curved and increasing grad- 
ually in size; the tip is more curved than the rest of the shell and 
strongly hexagonal in cross-section; the sculpture consists at first 
of 6 strong, primary ribs and wide, flat or slightly concave in- 
terspaces; the ribs are further increased by the introduction of 6 
intermediate ribs and still later by 12 more; these ribs are sub- 
equal in size on the later portions of the shell; the interspaces are 
finely transversely lined and faintly longitudinally striated. 

Length 52, diameter 6.25 mm. 
44; 4.75 tam. 

The Dentalium bocasensis is fairly abundant in the shale 
beds of Bocas Island. It is recognized by its strongly hexagonal 
tip and finely transversely lined interspaces,and strong longitudi- 
nal scuipture. 


Gatun Stage: Bocas del Toro. 


Subgenus FISSIDENTALIUM Fischer 
Dentalium uscarianum, n. sp. Plate 15, figure 1 


Shell large, solid and heavy; gently curved, and the taper of 
the shell is slow and gradual; the tips are all broken on our 
specimens, but the later sections of the shell are circular or slight- 
ly compressed dorsal-ventrally; the surface is sculptured with 
about 24, low, subregular and rounded longitudinal ribs with 


339 Costa RicA MrockNE—OLSSON 167 


smaller intermediate ones in the interspaces; the ribs become 
gradually obsolete with age and the later portions of full-grown 
shells are nearly smooth; fine transverse lines of growth cross the 
surface and are best seen on the earlier portions of the shell; 
operture oblique. 


Length 60, diameter 12 mm. 


This large Dentalium is abundant and very characteristic of 
certain conglomeritic zones in the Upper Uscari shales. There it 
occurs associated with a small but very distinctive fauna, includ- 
ing the Ptychosalpinx? dentalis, Sconsia cocleana and Scaphella 
costaricana and other species. When complete, the Dentalium 
uscartanune frequently reached a length of 110 mm or more. 

All our specimens are unfortunately lacking in the apical 
tip. In general sculpture, it recalls the Dentalium floridense 
Henderson dredged from 35 to 110 fathoms of water off Florida 
and in the West Indies, but the taper of the shell is more gradual 
and the sculpture more irregular, becoming obsolete with age. 
Bose* has described a large species as Dentalium rimosum from 
the Miocene of the Tehuantepec Peninsula. In that species, the 
ribs number about31, are very regular with deep interspaces, and 
ribs are finely etched by transverse lines of growth. 

Useari Stage: Coco Plum, Panama. 

Rio Coeles. 

Margarita—Old Harber trail. 
Comadre Creek. 

Pumbri Creek, ete. 


*Boll. Inst. Geol. do Mexico, No. 22, p. 55, pl. 3, figure 1, 1906. 


juL ‘7 Wee 


BULLETINS 
OF 


AMERICAN PALEONTOLOGY 


Vol. 9 


No. 39 


The Miocene of Northern Costa Rica 


With Nores On Its GENERAL STRATIGRAPHIC RELATIONS 
BY 
A. A. OLSSON 


Part 2 
CLASS PELECYPODA 


(Pp. r69-Finis) 


Yun 2T,) 1922 


Harris Co. 
Cornell University, Ithaca, N. ¥. 
Wie Se sek. 


ae f i 


343 Costa Rrca MroceNE—OLSSON 171 


ORDER PRIONODESMACKHA 


Genus NUCULA Lamarck 
Nucula orbicella, n. sp. Plate 28, figures 19, 20 


Shell of medium size, broadly elliptical or subovate in out- 
lines; obliquely truncated at its anterior end; gently convex; the 
surface is covered with close, subregular, concentric wrinkles and 
fine radial threads so that the sculpture is decussated; on the 
posterior dorsal slope, they are larger and pustular-like and their 
course is obliquely downward across the growth-lines; they are 
weak on the poorly defined lunule; the radical threads are main- 
ly between the concentric wrinkles but ventrally they cut into 
the wrinkles as well; interior of the shell, silvery white or faint- 
ly tinged with purple; the ventral margin is finely crenulated; 
chondrophore very small, oblique; anterior set containtng about 8 
teeth, the posterior with about 17. 


Length 9.50, height 7.50, semi-diameter 2.25 mm. 
II.00 8.50 2.50 mm. 


From the recent VV. crenulata A. Adams, this shell differs 
by its larger size and more elliptical and depressed shell. Gabb 
has described from the Pliocene of the Limon Peninsula WVzw- 
cula limonensts and moenensis, but both of these species are per- 
fectly smooth and sculptureless. ue Mucula orbicella is a com- 
mon species in certain marly beds intercalated in the coralline 
limestones of Port Limon. 


Gatun Stage: Port Limon. 
Nucula cahuitensis, n. sp. Plate 18, figures 21-24 


Shell small, solid, obliquely subtrigonal in form; ventral 
margin widely rounded, a nearly straight posterior dorsal margin 


123 BULLETIN 3g 344 


and a widely truncated anterior end; valves gently convex; pro- 
toconch on the tip of the umbo small, smooth and white; surface 
polished; the sculpture of the disk of the shell is developed to a 
variable extent, but the umbos are always smooth; typically the 
lower half of the shell is covered with close, regular, concentric 
wrinkles which are slightly decussated by fine, even, and slight- 
ly raised radial threads; in other cases, the wrinkles may be pres- 
ent only on the extreme anterior or posterior ventral extremities 
and the radial lines showing only as a part of the shell substance 
and not as raised threads; the escutcheon is elliptical and thickly 
covered with raised wrinkles which lie at right angles to the dor- 
sal margin of the shell; they are similiar on the lunule; interior 
of the shell tinted with brown; ventral margin crenulated; chon- 
drophore small, oblique with the anterior set of teeth numbering 
about 7, the posterior about 14. 
Length 3.50, height 3.00, diameter 1.60 mm. 

This is a small species of subtrigonal outlines, suggestive of 
the Pacific VV. extgua Sowerby in general form and size. Onthe 
few specimens of V. exiywa, which I have from the Bay of Pan- 
ama the sculpture is finely decussated over the whole shell with 
fine radial and concentric threads. The most striking features of 
cahutiensts are its strongly sculpture lunule and escutcheon. 


Gatun Stage: Zone G, Saury Creek. 


Nucula tenuisculpta Gabb Plate 18, figure 25 
Nulua tenuisculpta Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 
255. 
Nucula tenuisculpta Maury, 1917, Bull. Amer. Pal., vol. 5, p. 325, pl. 
26, figure 8. 


A species of the size and form of cahuztenszs but the valves 
are entirely smooth and polished. ‘The lines of growth are fine 
and irregular. The radial lines are faint and sub-microscopic. 
Inner margin crenulated. 


Height 3.25, length 4.00, semidiameter .75 mm. 
Gatun Stage: Coll 4, Red Cliff Creek. 


345 Costa RicA MrocENE—OLSSON 78 


Genus LEDA Schumacher 


Leda Balboae Prown and Pilsbry i Plate 18, figure 15 


Leda Balboe Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., 
vol, 63, p. 362, pl. 27, fig. 8. 

This species is fairly common in Costa Rica, but generally 
smaller than in the Canal Zone, where it frequently reaches a 
length of 22 mmior more. Its valves are moderately convex and 
regularly sculptured with even, close, concentric riblets which on 
the center of the shell disk may be slightly appressed. The 
escutcheon is well-defined and similiarly sculptured with the rib- 
lets running parallel to the hinge margin. 


Gatun Stage: Gatun, C. Z. 
Coll. 1, East Grape Point Creek. 
Zome G, Saury Creek. 
Outtana Creek. 
fiill No. 3, Banana River. 


Leda Davilae, n. sp. Plate 18, figure 20 


Shell small, solid, strongly convex; the beaks nearly cen- 
tral; the rostrum is short, pointed and sharply defined from the 
rest of the shell disk by a high, cord-like ridge; the anterior end 
is bluntly rounded and carries 2, low flexures from the beaks to 
the anterior ventral extremity; lunule small and narow, lacking 
the concentric ribs and sculptured with small, pustular-like 
ridges passing at right angles to the hinge-line; the sculpture con- 
sists of about 30 concentric riblets which are somewhat larger 
and with deeper interspaces on the umbos; the interspaces show 
on the umbos and along the anterior slope, fine, radial lines or 
punctations; interior of the shell cavity deep, with a heavy 
hinge; the anterior and posterior sets of teeth number about 13 
or 14, are high and V-shaped. 


Length 5.75, height 4.00, semi-diameter 1.75 mm. 


174 BULLETIN 39 3246 


A small, plump species, coarsely sculptured with even, con- 
centric riblets, separated by deep interspaces. These interspaces 
may carry fine, radial lines or punctations at the base of the con- 
centric ribs and are best seen on the umbones and the anterior 
slope. The ZL. indigina Dall (L. dbisulcata Guppy) from the 
Miocene of Jamaica is similiar but proportionately longer. 


Gatun Stage: Coll. 4, Red Cliff Creek. 
Bocas Island. 


Subgenus ADRANA, H. and A. Adams 
Leda quitanensis, n. sp. Plate 18, figure 19 


Shell thin, elongate but slightly convex; beaks very low and 
scarcely distinguishable from above and situated at the anterior 
I-3; ventral margin an even curve from the pointed extremities; 
dorsal margin straight; surface polished but covered with very 
fine, concentric lines which are crossed on the lower half of the 
anterior 2-3, by even, oblique lines (.Sczssuda-like); these oblique 
lines are spaced about .25 mm apart; interior concealed. 


Height 7.25, lenght 26.00, semidiameter .75 mm. 


A much smaller species than the next (emsznoides), slightly 
nlore convex and with beaks situated more anteriorly. The sur- 
face is finely sculptured about the middle and the anterior por- 
tion by slightly oblique lines, as seen on certain Tellinoids such 
as Scissula. The Manzanilla beds of Trinidad contain Leda 
(Adrana) Guppyi Dall (Cercomva ledeformis Guppy). It is less 
elongate, more contracted posteriorly and has a different surface 
sculpture. Dall has also recorded Leda Guppyi from the Bow- 
den beds of Jamaica. 

Gatun Stage: Quitana Creek. 

Zone F, Saury Creek. 
Coll. 1, Fast Grape Point Creek. 


Leda ensingides, n. sp. Plate 18, figure 12 


Shell rather large, elongate, depressed and subequilateral; 


347 Costa RicA M1ocENE—OLSSON 175 


beaks very small and scarcely distinguishable from above, and 
situated nearly about the middle of the dorsal margin; extremi- 
ties bluntly rounded, with a broad, even curve to its base or ven- 
tral margin; the dorsal side is nearly straight but descending 
slightly towards its extremities; surface nearly smooth, the 
growth lines indistinct, but with fine, even very slightly oblique 
lines on the anterior two-thirds of the shell; interior con- 
cealed. 


Length 54.00, height 15.00, diameter 4.50 mm. 


Differs from the preceding species by its larger size, nearly 
central beaks and by its finer, more concentric sculpture. ‘There 
are several species of this group recent in the West Indies and 
along the Pacific coast of Central America, but none appear at 
all closely related to the Gatun species. 


Gatun Stage: Hill No. 3, Banana River. 


Leda Dalliana, n. sp. Plate 28, figure 17 


Adrana sp ? Dall, 1898, Trans. Wagner Free Inst. Sci., vol. 3, pt. 1 
4, P- 592. 


Shell small, thin, elongate, depressed; the small, scarcely 
differentiated beak is sutuated about the anterior one-fourth of 
the shell; the anterior side is somewhat contracted and shortly 
rounded while the posterior side is produced and 3 times the 
length of the anterior and bluntly pointed at its extremity; the 
rostrum is long and narrow, formed by 2 radial, scabrous threads, 
separated by a wide interspace; concentric sculpture of thin, 
elevated threads or Jamellae, regularly distributed on the lower 
half of the shell, but widely spaced on the umbonal area; the 
escutcheon is long and narrow, sculptured with threads parallel 
to the hinge-margin; interior of shell cavity shallow, the long 
hinge with an anterior set of 20 small teeth, the posterior with 
26 or 27. 


Length 10.50, height 3.50, diameter of left valve .75 mm. 


176 BULLETIN 39 348 


A much smaller species than the preceding and differing in 
its more anterior beaks, depressed shell and heavier concentric 
sculpture as well as in other characters. It is probably the shell 
which Dall refers to as having fragments of from the Pliocene 
of Limon. But a single perfect specimen was collected 


Gatun Stage; Port Limon. 


Genus YOLDIA Moller 
Yoldia bocasensis, Plate 28, figures 21, 22 


Shell small, thin, ovate-elliptical in form; beaks situated 
slightly in front of the middle; vaives moderately convex; both 
the anterior and posterior extremities are broadly rounded and 
connected by the gentle curve of the basal or ventral margin; the 
umbos are very small! but distinct and lie just above the greatest 
convexity of the valves; surface smooth, polished with no other 
sculpture than the incremental growth-lines; chondrophore dis- 
tinct, scarsely oblique and bordered on each side by an equal num- 
ber of V-shaped hinge-teeth; the anterior set contains about 14 
teeth, which are very small next to the chondrophore becoming 
progressively larger distally, the posterior set with 13 or 14; the 
pallial sinus is indistinct, but rather large, broad and extends 
nearly to the middle of the valve. 


Length 13.00, height 7.00, semidiameter 2.25 mm. 
9.50 5.50 1.50 mm. 

A fairly common species in the clay beds of Bocas Island 
where it is associated with a fauna of rather deep water charac- 
ter. It has much the form of certain deep-water Yo/dzella but 
the interior of the valves show an indistinct but deep and wide 
pallial sinus. 


Gabb has described from the Miocene of Santo Domingo, a 
Yoldia ovalis, which has never been figured and its description is 
very meager and insufficient. . 


Gatun Stage: Bocas del Toro. 


349 Costa Rica M10cKNE—OLSSON 177] 


Genus LIMOPSIS Sassi 
Limopsis menilis, n. sp. Plate 26, figures 23, 24 


Shell small, solid, moderately convex and subcircular in form 
but slightly oblique; the hinge-line is straight, the anterior dor- 
salsubmargin very small, the posterior large undifferentiated 
from the rest of the shell ; the umbos are high and prominent, 
lying a little in front of the middle; exterior sculptured with 
strong radial ribs which are largest and strongest on the middle 
of the shell disk, finer on the dorsal submargins; these ribs are 
crossed by high widely spaced concentric lamellae, more closely 
spaced on the umbos; the lamellce on crossing the ribs are raised 
into short, spine-like fringes which are directed ventrally; inter- 
ior of shell deep; the hinge-line with a small cartilage pit at the 
anterior 1-3 of the shell; it is bordered on the anterior side by a 
set of 3 high, projecting teeth and 4 onthe posterior side, the 
ventral and a part of the anterior margin is crenulated, most fine- 
ly in the middle and quite coarsely at the posterior portion. 


Length 1.75, height 2.00, semidiameter .60 mm. 


This is a small, subcircular species with a crenulated inner 
margin and a strong surface sculpture of ribs crossed by distant 
concentric fringes. It is fairly common at its type locality, giv- 
en below. 

Gatun Stage: Old Man Sam Creek, 1 mile south of beach 

(A. C. Veatch, collector). 


Genus GLYCYMERiS, Da Costa 
Glycymeris canalis Brown and Pilsbry Plate 18, figures 2-7 
Glycymeris canalis Brown and Pilsbry, Igt1, Proc. Acad, Nat. Sci. 


Phila., vol. 63, p. 364, pl. 28, fig. Io. 


Glycymeris trilobicostata Pilsbry and Brown, 1917, Proc. Acad, Nat. 
Sci. Phila. 


Very abundant locally in the Gatun beds of Costa Rica, are 
small and medium sized Pectunculids belonging to the group of 


178 BULLETIN 39 350 


G. acuticostata Sowerby of the Santo Dominican Miocene. They 
vary somewhat in size, (20 to 32 mm in height), are usually de- 
pressed and with an outline suggestive of the recent East Coast 
and Upper Miocene G. pectinata Gmelin. Typical acuticostata 
does not seem to occur in Costa Rica and I have adopted Brown 
and Pilsbry name of canals. 


The G. canalis Brown and Pilsbry from the Canal Zone, and 
the frilobtcostata Pilsbry and Brown from Colombia, are both bas- 
ed on young shells 15 mm or less in height. Both forms can be 
duplicated amongst the Costa Rican shells and are but varieties 
of the same species. 


In its usual and more abundant form, the shell is depressed, 
with numerous (about 26), closely spaced rounded or angled ribs. 
These ribs are overrun by even, close, concentric threads. They 
represent the usual caza/zs or its variety frzlobicostata, and aver- 
age in height about 25 mm. More rarely, the shells are larger 
and as illustrated by figure 4 and 5 the ribs may become divided 
into 2 or 3, strong, radial cords. The following measurements 
will show the usual range in size. 


Height 22, diameter 21, thickness 5.5 


24 23 5-5 
22 32 8 
29 29 8 


Gatun Stage: Gatun, C. Z. 
Comadre; Banana River; Coll. 4 East Grape 
Point Creek. 
Old Man Sam Creek, C. R. 


Glycymeris jamaicensis Dall Plate 19, figures 3, 4 


Pectunculus pennaceus Guppy, 1866, Quart. Journ. Geol. Soc. London, 
vol. 22, p, 293. Not of Lamarck. 


Axinea pennacea Gabb, 1873, Trans. Amer, Phil. Soc., vol. 15, p. 255. 


Pectunculus decussatus Guppy, 1876, Qart. Journ. Geol, Soc. London, 
vol. 32, p. 532. Not of Linné. 


351 Costa Rica M1ocENE—OLSSON 179 


Glycymeris jamaicensis Dall, 1898, Trans. Wagner Free Inst., Sci., 
vol. 3, pt. 4, p. 608. 

Glycymeris jamaicensis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 345, 
pl 26; fig! 13) 

This large Pectunculid, externally resembles in its angular 
posterior side the Glycymerts pennacea with which it was confus- 
ed by Guppy and by Gabb, It differs in being somewhat less con- 
vex, less strongly sculptured and most importantly in its central 
beaks, located about the middle of the ligamental area. It is, on 
the other hand, much more closely allied with the recent West 
Indian G. undatus Linné (G. lineatus Reeves) .of which it is 
doubteless the Miocene ancestral form, the main difference being 
in details of its finer sculpture. 


The Glycymerts jamaicensis occurs in the Miocene of Jamaica 
and Santo Domingo but there never seems to reach the large size 
that the species sometimes attains in Costa Rican. A large shell 
in our collection from Old Man Sam Creek measures as fol- 
lows: 


Height 60, lenght 60, diameter of both valves 37 mm. 


Gatun Stage: Water Cay. 
Collis Red (Clip) Oreck. 
Coll. ga, East Grape Point Creek. 
Colls. 5, 6, Old Man Sam Creek. 
Soust Creek of Upper Hone Creek. 


Glycymeris carbasina Brown and Pilsbry Plate 19, figures I, 2, 5 


Glycymeris carbasina Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. 
Phila., vol. 63, p. 363, pl. 28, fig. 9. 

The Glycymeris carbasina was described by Brown and Pils- 
bry from a small individual from Gatun, measuring only 16 mm 
in height. It is closely related to the G. jamatcensis Dall and 
probably represents but a varietal form. Its main differences is 
its more circular and less convex shell, and less number of hinge- 
teeth which number about 1o to 12 on the anterior and posterior 
sides respectively. 


180 BULLETIN 39 352 


All our specimens were collected in the lower Gatun of E. 
Grape Pt., Cliff Creek in Eastern Costa Rica, the largest shell 
measuring as follows: 


Height 56, length 55, diameter of both valves 32 mm. 
Gatun Stage: Coll. 4, East Grape Point. 


Glycymeris castaneus Lamarck Plate 18, figures 13, 14 


Fectunculus castaneus Vamarck, 1819, Anim. sans Vert., vol. 6, p 
53. 

Fectunculus castaneus Reeve, 1843, Conch. Icon., Pectunculus pl. 6, 
fig \/32) 

Our collection from the Banana River contains several small 
Pectunculids which differ from the jamazcensis and its allies by 
their less circular shell, greater convexity and coarser radial 
sculpture. These shells agree however very closely with a recent 
species which is abundant on the beaches of northern Costa 
Rica, which appears to be the Glycymeris castaneus of Lamarck. 
Glycymerts castaneus is distinct from undutus Linné (G. lineatus 
Reeve) with which it is sometimes united, differing by its form 
which is more produced posteriorly but not angulated and by its 
more pronounced radial costeze. Both species are similiarly col- 
ored with chestnut markings. 


Gatun Stage: Hill No. 3, Banana River. 
Quitana creek. 


Glycymeris decussatus Linnzeus Plate 19, figures 6, 7 


Arca decussatus Linneus, 1758, Syst. Nat., ed. 10, p. 694. 

Fectunculus pennacea Vamarck, 1819, An. s. Vert., vol. 6, p. 51. 

Fectunculus pennacea Reeve, 1843, Conch. Icon., /ectunculus, pl. 
5s ten 248 

Fectunculus decussatus Linné. P. pennaceus Wamarck, (Lamy) 1911, 
Journ. de Conchy., vol. 59, p. 119, pl. 3, fig. 7. 

Of this rare shell, more commonly known as the G. penna- 

cea Lamarck we have several excellent specimens collected by 


353 Costa RicA M1ockENE—OLSSON 181 


Dr. A. C. Veatch from the Upper Gatun of Old Man Sam Creek 
in eastern Costa Rica and a few specimens from the coral-reef 
limestones of Port Limon. Although a very distinct species and 
not to be confused with any other Pectunculid recent or fossil, 
its characters have been generally misunderstood and its name 
has often been applied to shells of the G. /éneatus group. 


The shell is of moderate size, strongly convex and inequila- 
teral. The beaks are slightly posterior of the middle, with the 
ligamental area entirely anterior to the beaks as seen in figure 7. 
The surface is sculptured with numerous, moderately coarse, 
radiating threads which are simple on the umbonal area but be- 
come divided by 3 or more finer threads ventrally. 

Gatun Stage: Old Man Sam creek. 

Port Limon. 


Glycymeris Lloydsmithi Pilsbry and Brown Plate 25, figures 8. 9, I0 


Glycymeris Lloydsmithi Pilsbry and Brown, 1917, Proc. Acad. Nat. 
Sci. Phila., vol. 59, p. 39, pl. 6, fig. 6. 

This is an abundant fossil in eastern Costa Rica, where it 
frequently forms zones in the lower and middle Gatun. Itstype 
of sculpture of broad, smooth, rounded ribs, is strikingly like 
that of G. swbovata Say of the Miocene of eastern United States. 
It differs in its higher and narrower umbos and by its nearly 
smooth cardinal area. 

The G.Lloydsmithi was described by Pilsbry and Brown, 
from beds equivalent to the Gatun, near Cartagina, Colombia. 

Gatun Stage: Zone 3, East Grape Point Creek, C. R. 

Coll a Red: \Cligh Greeks (Guipie 
Headwater of Middle Creek, C. R. 


Genus ARCA Linnzeus 
Arca occidentalis Philippi Plate 22, figure I 


Arca occidentalis Philippi, 1847, Abbild. u. Beschr., 3, p. 14, pl. 17b, 
fig. 4a-c. 


182 BULLETIN 39 354 


Arca noe Guppy, Journ. Geol Soc. London., vol. 22, p. 293. 


Arca occidentalis Sheldon, 1916, Paleont. Amer., vol. 1, p. 8, pl. 1, 
figs. 8-11. 


Arca occidentalis Maury, 1917, Bull. Amer. Pal. vol. 5, p.327, pl. 29, 
Tey 
This is a common recent species of the West Indian and 
Caribbean fauna. Asa Miocene fossil it is found in Jamaica 
and Santo Domingo and it is here recorded from the Miocene of 
Costa Rica. 


Gatun Stage: Port Limon. 


Arca umbonata Lamarck Plate 22, figure 2 


Arca umbonata Lamarck, 1819, An. s. Vert., vol. 6, p. 37. 


Arca imbricata Gabb, 1873, Trans. Amer. Phila. Soc., vol. 15, p. 254, 
In part. 


Arca umbonata Sheldon, 1916, Paleeont. Amer., vol. 1, p. 8. pl. 1, 
figs. 12-17. 

Arca umbonata Maury, 1917, Bull. Amer. Pal., vol. 5, p. 327, pl. 30, 
Ag.) TN. 

Arca mubonata Maury, 1920, New York Academy of Science, Sci. Sur. 
Of PAR vols) 3\ipt.) 1s) pii6: 

The Arca umbonata and the preceding occidentalis area com- 
mon as recent shells along the entire northern coast of Panama 
and Costa Rica. The A. umbonata is recognized by its more 
convex and angular shell and finer sculpture of the middle of 
the shell disk. Asa Gatun fossil we have but a single small 
shell from the Hast Grape Point Creek. Dr. Maury records 
the species from the Miocene of Santo Domingo and from 
Porto Rico. 


Gatun Stage: Coll. 1, East Grape Point Creek. 
Subgenus BARBATIA (Gray) Adams 


Arca Mauryae, n. sp. Plate 22, figure 4, 7 


Barbatia cf. Bonaczyi Maury, 1917, Bull. Amer. Pal., vol. 5, p. 329, 
pl. 30, fig. 15. Not of Gabb. 


355 Costa RicA MIocENE—OLSSON 183 


Shell small, thin, convex, subrhomboidal in form; umbos 
wide and full; with the beaks at the anterior 1-3 of the shell; a 
rounded posterior umbonal slope; the anterior extremity is brok- 
en inthe type specimen but from the earlier growth lines is 
simply rounded in form; the posterior side is obliquely truncate 
and meets the dorsal margin in a well-marked angle; the exteri- 
or is sculptured like that of Arca barbata, with the riblets group- 
ed in wide, regular bands which are defined by slightly deeper 
interspaces; the radial riblets are crossed and crudely granulat- 
ed by concentric threads; the sculpture on the posterior submar- 
gins is much heavier and predominantly radial that of the an- 
terior submargin is but slightly heavier than on the middle of 
the shell disk; interior of shell cavity deep, with a straight hinge- 
margin, the teeth arranged asin darbata; cardinal area rather 
high, longitudinally striated posterior of the beaks, plain in 
front. 

Length 24? height 11.50, semidiameter 5.50 mm. 

This species is closely related to the recent Avca barbaia 
Linné. It differs from daréaza in its proportional shorter shell, 
greater convexity, less anterior beak and higher cardinal area. 
It is equivalent to the Dominican shell figured by Dr. Maury as 
Barbatia cf. Bonaczyt Gabb. Gabb in his remarks following his 
his description of Bonaczyi, compares his shell with éarbata, and 
states that in Bonaczyz, the beaks are more anterior and the pos- 
terior end more produced and sloping. ‘This is the reverse of 
the conditions as seen in the present shell. Dall on the other 
hand, considers Gabb’s shell as synonymous with Arca umbonta 
Lamarck. 


Gatun Stage: Port Limon. 


Section CALLOARCA Gray 
Arca cf. candida Gmelin Plate 22. figures 5, 6 


Arca candida Gmelin, 1792, Syst. Nat., p. 3311. 


Barbatia (Calloarca) candida Dall, 1898, Trans. Wagner Free Inst., 
SCi., Vols) 34) pt 4). p. 626. 


184 BULLETIN 39 356 


Arca cadida Shelton, 1916, Palaeont. Amer., No. 1, p. 16, pl. 3, figs. 
The 02) 

We have several small valves of a Calloarca from the Mio- 
cene limestones of Port Limon, the largest of which has a length 
of 1 mm. ‘They belong to the Arca candida group and may 
possibly represent the young of that species. They seem to dif- 
fer in being more finely sculptured, especially about the middle 
of the shell disk. ‘This sculpture consists of fine, beaded or 
granulated radial threads, which become coarser on the anterior 
and posterior submargins. 


The Arca candida is recorded by Dall from the Bowden beds 
of Jamaica, the equivalent of the coral limestones of Limon. Dall 
also mentions from the same locality, the occurrence of a smaller 
and possible distinct species. The Arca candida is a common re- 
cent species of the West Indies and the Caribbean, and is abund- 
ant on the north coast of Panama and Costa Rica. 


Length 18, height 11?, semidiameter 3.75 mm. 
Gatun Stage: Port Limon. 


Subgenus SCAPHARCA Gray 
Arca dariensis Brown and Pilsbry Plate 22, figures 10-13 


Arca oronlensis Dall, 1898, Trans. Wagner Free Inst. Sci., vol., 3, pt. 
4, p. 658. Not of Gabb. 

Arca dariensis Brown and Pilsbry, 1911, Proc. Acad, Nat. Sci. Phila., 
vol. 63, p. 362, pl. 22, fig. 10. 


Arca gatunensis Toula, Dec. 1911, Jahrb. der K-K Geol. Reichsanstalt, 
vol, 61, p. 493, pl. 30, fig. 4. ! 

The common Ark of the Canal Zone. The shell is elongate 
in form with about 30 ribs. The ribs of the left valve are usually 
granulated over the whole shell, while those of the right valve 
are narrow and smooth on the middle of the shell disk. The ribs 
are characteristically divided on the anterior and posterior ends 
of the shell. 


Gabb has described an Avca oronlensts from the black shales 


357 Costa Rica M1ocENE—OLSSON 185 


of Oronli Creek in the Talamanca Valley (probably the Uscari 
shell). Itis like A. dariensis, in form, but with more central 
beaks and umbones and simple ribs. We have not seen this 
species. 
Gatun Stage: Gatun and Mt. Hope, C. Z. 
Woter Cay. 


Arca actinophora Dall Plate 23, figs. 7, 8; Plate 25, fig. 3 


Scapharca (.Scapharca) acitnophora Dall, 1898, Trans. Wagner Free 
Inst. Sci., vol. 3, pt. 4 p. 647, pl. 33, fig. 26. 


Scapharca (Scapharca) actinophora Sheldon, 1916, Paleont. Amer., 
VOl tA Ps) 50) pl. Tn, fees: 

A larger species than the davzenszs, with broad, wide umbos. 
The ribs number 36 to 40, are narrow and separated by inter- 
spaces of a little more than their own width. ‘These interspaces 
are generally finely ornamented by regular, even, concentric 
threads. The ribs are simple and on the left valve, usually strong- 
ly beaded or granulated throughout, becoming wider and smooth- 
er on the posterior portion, or with the granulations only on 
their edges, so that the ribs appear as if mesially depressed. The 
ribs of the right valve are less strongly beaded and may become 
quite smooth, especially on the center of the shell disk. 


Its ventral margin is gently curved to the rounded and not 
pointed posterior extremity. 


It is a common species at Mt. Hepe in the Canal Zone, but 
it also occurs but less abundant in Western Panama and in 
Costa Rica. 


Catun Stage. Mi. ope, C. Z. 
Water Cay, Panama. 
Zoned, hea Clif, Greer iGy ie 
Coll. 3, Hone Walk Creek, C. R. 
Hill t Banana River, C. R. 


186 BULLETIN 39 358 


Arca Henekeni Maury Plate 24s aman ts 
Arca consobrina Sowerby, 1849, Quart. Journ. Geol. Soc. London, vol. 
6, p. 52, pl. ro, fig. 12. Not of d’Orbigny, 1844. 


Arca consobrina Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 
32, P. 531. 

Arca consobrina Dall, 1888, Trans. Wagner Free Inst. Sci., vol. 3, pt. 
4, p. 646. 


Arca consobrina Sheldon, 1916, Paleont. Amer., No. I, p. 49. 

Arca Henekeni Maury, 1917, Bull. Amer. Pal.,vol. 5, p. 331,pl. 29,fig. 2. 

Arca (Scapharca) sobrina Pilsbry and Johnson, 1917, Proc, Acad. Nat. 

Sci. Phil., vol. 64, p. 186. 

Our shells from Water Cay are similiar in form and general 
sculpture to the 4. Heneken? Maury (A. consobrina Sowerby), 
from.the Miocene of Santo Domingo, the only noticeable differ- 
ence being the greater number of ribs on the Panama shell. The 
ribs of Hlenekeni from Water Cay average about 39 in number, 
and 35 to 36 on the Dominican shells. The ribs of the left 
valve are evenly beaded or granulated by concentric threads 
which pass across the interspaces as elevated threads. The 
ribs of the posterior-dorsal submargins are nearly smooth. 
They are more or less divided at the anterior extremity and 
very finely beaded. 


Length 29, height 18.25, diameter 14.15 mm. 
Bile 18, semidiameter of right value 


7.25 mm. 
Gatun Stage: Water Cay. 


Arca costaricensis, n. sp. Plate 25, figures I, 2 


Shell moderately large, elongate, convex, with the beaks sit- 
uated at the anterior 1-3;ventral margin but gently curved to the 
attenuated and pointed posterior extremity; right valve with 
about 30 ribs, of which the most anterior and posterior ones are 
rather wide, the others narrow; interspaces on the center or the 
disk 3 times or more the width of the ribs; the ribs of the right 
valve are obscurely granulated on the anterior extremity, more 
or less smooth on the center but on the posterior extrem- 
ity, they are double and peculiarly granulated along their sides; 


359 Costa RrcA MiocENE—OLSSON 187 


this granulation begins on the sides and on the more anterior 
ribs, the center is smooth; this granulation is in the form of in- 
verted V’s; their acute apices lying along the center of the ribs; 
the ribs of the left valve are more or less granulated or beaded 
throughout and with the ribs on the posterior extremity, double 
and similiar to those of the right valve; hinge-line straight, 
with very numerous small, uninterrupted teeth; a rather wide 
cardinal area with 5 or 6 ligamental grooves; ventral margin in- 
ternally fluted in harmony with the external ribs. 


Length 52, height 28, diameter 34 mm. 
52, Bn 17 mm (right valve). 


The form of this shell with its gently curved ventral mar- 
gin and attenuated posterior extremity is like the Avca actino- 
phora Dall, but is longer and with fewer and heavier ribs. On 
the other hand, this species may be compared with darzenszs 
Brown and Pilsbry, from which it differs most strikingly by its 
larger size, wider, higher and more central umbos and more 
pointed attenuated posterior extremity. 


Gatun Stage: Hill ra, Banana River. 
Fill 2, Banana River. 
Zone 7, Pumbri Creek. 


Arca honensis, n. sp. Plate 22, figures 8, 9 


Shell rectangular, elongote, moderately convex; beaks situ- 
ated at about the anterior 1-4; right valve with about 26, nar- 
row, smooth or irregularly granulated ribs, separated by inter- 
spaces of twice their own width; the left valve with the same 
number of ribs and similiar to those of the right valve; the ribs 
of both valves are simple and undivided and if granulated, most 
heavily on the anterior half of the left valve; interspaces smooth 
or with irregular, raised concentric threads; cardinal area long 
and rather wide and with 5 or 6 linear grooves; hinge-lines 
straight with very numerous, small, uninterrupted teeth; ven- 
tral margin fluted in harmony with the external ribs. 


188 BULLETIN 39 260 


Length 39, height 23.5, semi-diameter 11 mim (lest valve) 
36.5 23 12 (left valve) 
34 23 ie) (Galea wade) 


In general form, quite similiar to the Avca dariensis Brown 
and Pilsbry, but distinguished mainly by its simple, undivided 
ribs. Itis also amore convex shell, with higher and more 
prominent umbos. It differs from Gabb’s figure of A. ovonlen- 
sts in its more anterior umbos. 


It isa common species, occurring in zones in the Gatun 
sandstone of Hone Walk Creek of Eastern Costa Rica. 


Gatun Stage: Hone Walk Creek. 


Arca golfoyaquensis Maury, var. medioamericana, n. var. 
Plate 23, figures 4-6 


ef. drca golfoyaquensis Maury,1917, Amer. Pal., vol. 5, p. 332, pl. 28, 
fig. 5. 

? drca (Andara) consobrina Gabb, 1881, Journ. Acad. Nat. Sci. Phila., 
and series, vol. 8, p. 378. 

The Arca golfoyaquensis was described by Dr. Maury from 
the Gurabo and Cercado formation of Santo Domingo. It is 
closely related to the Arca Henekent Maury, (the Arca consob- 
yina) differing mainly in form. 

The Costa Rican shells are very similiar to Dr. Maury’s 
type specimens of golfoyaquensis, but differ in having 35 instead 
of 38 ribs, a higher shell, fuller umbos and sharper ribs on the 
posterior-dorsal area. The central ribs of the left valve are ele- 
gantly beaded with their interspaces evenly sculptured with 
raised concentric threads. On the right valve, the ribs are more 
smooth but their interspaces are similiarly sculptured to those of 
the left. The ribs of the posterior-dorsal angle of both valves 
become divided ventrally by 2 or 3 fine threads. ‘The posterior 
extremity is straight, meeting the hinge-line at an angle of 
nearly 120 degrees. 


Length 54, height 34, diameter 34 mm. 


361 Costa RicA MiIocENE—-OLSSON 189 


Gatun Stage: Hill 1, Banana River. 
Coll ss ied. Glan Greek 
Rio Blanco. 
Old Man Sam Creek, r mile south of beach. 


Arca Veaichi, nu. sp. Plate 23, figures 1-3 


Shell large, moderately convex; in outline, the shell is near- 
ly square with the height about equal to its length; the anterior 
side is widely rounded to the ventral margin, the posterior side 
_ nearly straight, nearly at right angles to the hinge-line; umbos 
wide, with the unbonal angle widely rounded and with the ex- 
treme tip of the beaks situated at the anterior 1-3 of the total 
length of the shell; right valve with about 45 ribs, of which the 
posterior 14 are on the posterior-dorsal slope; the ribs are rec- 
tangular in section with the interspaces on the center of the 
disk about 3-4 of the width of the ribs; the ribs are finely scul- 
ptured but less strongly on the right valve; on the center of the 
shell disk, the ribs are finely nodulated or granulated but on the 
posterior-dorsal slope and on the anterior extremity, the granu- 
lations of the ribs is gradually overshadowed by the introduc- 
tion of 4 or 5 fine, threads on the top of each rib the interspaces 
on the center of the disk of the left valve are finely sculptured 
with even, raised concentric threads; the left valve is slightly 
larger and overlaps to a small extent the right; hinge-line 
straight with numerous small teeth, larger and higher at the ex- 
tremities; cardinal area that of typical Scapharca, and grooved 
with 3 lines; interior of the shell deep, with the posterior ad- 
ductor scar about twice the size of the anterior; basal margin 
fluted in harmony with the external sculpture. 


Length 51, height 51, diameter 41.5 mm. 


This isa rather large, Cardiwm-like Arca of nearly square 
outlines and numerous finely sculptured ribs. Its form is also 
suggestive of Avgzna, but its cardinal area is that of a true 
Scapharca. Thetype specimen, a finely preserved individual 


1gO BULLETIN 39 362 


with both valves comes from the Gatun of Water Cay. It is 
quite unlike any other known American species. 


It is named for Dr. A. C. Veatch, Director of the Explora- 
tion Department of the Sinclair Consolidated Oil Corporation. 


Gatun Stage: Water Cay. 
Mit TLopen Guz 


Arca auriculata Lamarck Plate 22, figure 3 


Arca auriculata Lamarck, 1819, n. s. Vert., vol. 6, p. 43. 

Arca (Scapharca) auriculata Dall, 1898, Trans. Wagner Free Inst., 
Sci., vol. 3, pt. 4, p. 649. 

Arca auriculata, Sheldon, 1916, Paleont, Amer., vol. 1, p. 50, pl. I1. 
fig. 19, 

Scapharca auriculata Maury, 1917, Bull. Amer. Pal., vol. 5, p. 339, pl. 
28.) fi) 3) 

A common, recent species of the West Indies. Its hinge- 
line is generally produced or auriculated at its posterior extrem- 
ity. 

It occurs in the Miocene of Jamaica and Santo Domingo. 
Dall also records it from near Limon. 

The Costa Rican specimens in our collection are all small 
but otherwise typical. 

Gatun Stage: Coll. 3, Hone Walk Creek. 

Old Man Sam Creek, 1 mile south of the beach, 


Section CUNEARCA Dall 
Arca cacica, n. sp. Plate 24, figure 1 


Shell of moderate size, strongly convex and with high, in- 
flated umbos; beaks and umbos about the middle of the shell; 
the greatest convexity of the shell lying about the anterior 1-3, 
and with a narrow but deep, radial depressed zone extending 
from the beaks to the ventral margin, just anterior to the um- 
bonal angle; the anterior end is slightly rounded, the posterior 


363 Costa Rica MiocENE—OLSSON 19] 


submargins strongly depressed or flattened with a straight post- 
erior margin; the left valve with 30 smooth and strongly nodul- 
ous radial ribs of which 9 are situated on the posterior sub- 
margin; the ribs of the posterior submargin are low, wide and 
nearly smooth; on the rest of the shell, the ribs are nodulose, 
very regularly in the middle of the valves more distantly on the 
anterior ribs; the interspaces between the ribs are narrow on the 
middle of the valves, become wider anteriorly and on the ex- 
treme anterior extremity equal the ribs in width; the cardinal 
area is high, as nearly % of the length. 
Length 39, height 33, diameter of the left valve 15 mm. 

The above description is based on a single left valve from 
the Upper Gatun beds near Cahuita. Distantly related to the 
Arca tncongrua Say and its southern form 6vaszliana Lamarck, 
the fossil species differs in its higher and more convex shell, and 
in its high cardinal area. In this latter feature, this species is 
like the recent West Coast Arca labiata Sowerby, but has a 
greater number of ribs and different form. 

Gatun Stage: Across the divide trom Comadre Creek. 


Groups of Arca Pittiert Dall 


This is a group of small and medium-sized Arks, containing 
3 species, whose general appearance is that of the recent Avca 
Chemnitzt Phil. The Costa Rican shells are usually abundant 
wherever they occur in the Gatun formation. Their relations 
and differences may be summarized as follows: 

A. Both valves more or less similiarly sculptured, that is 
the ribs of the posterior half of the right valve, at least obsolete- 
ly beaded. 

B. Ribs 25 to 28; umbos high and full; posterior-dor- 
sal slope, angular; posterior extremity somewhat produced, 
giving an elongate outline to the shell. 


Area Pittiert Dall 


AA. Valves not similiarly sculptured; posterior half of 


192 BULLETIN 39 264 


the right valve with narrow ribs, separated by wide inter- 
spaces. 

B. The intervals between the ribs on the posterior 
half of the right valve with small (sometimes faint) interstitial 
threads. Ribs about 29. 

Arca Lloydi, n. sp. 

BB. The intervals between the ribs on the posterior 

half of the right valve smooth; ribs 25, shell usually 


larger. 
Arca Flindsiz, n. sp. 
Arca Piitieri Dall Plate 24, figures 2-6 
Arca (Scapharca) Pittieri Dall 1912,Smith. Misc. Coll., vol. 59, No. 2, 
P. 9. 


This isa common species in the Gatun beds of the Banana 
River. The ribs of the 2 valvesare similarly sculptured but 
somewhat less strongly on the right. The umbos are high. 
The posterior extremity is pointed and produced so that the out- 
line of the shell is more elongate than in the following species: 

Length 25, height 25, diameter 26 mm. 
20 28 semi-diameter 14.5 mm. 

Gatun Stage: FHiill ra, Banana River, 3. 

Zone Ei, Saury Creek. 
Rio Betey. 
Goll. 7, Pumbre Creek. 


Arca Llioydi, n. sp, Plate 24, figures 10-12 


Shell small, cordiform, covex, subrectangular in outline but 
with the left valve somewhat more pointed at its posterior-ven- 
tral extremity; anterior side well rounded; posterior side straight 
and meeting the basal margin at an angle of about 75 degrees; 
umbonal angle rounded; the left valve has 29 ribs of which g 
are found on the posterior- dorsal slope; the ribs of the left 
valve are strongly beaded or granulated, and separated by inter- 
spaces of about their own width; on the right valve, the first 10 
anterior ribs are strongly beaded; the next set as far as the um- 


365 Costa Rica MrocENE—OLSSON 193 


bonal angle are smooth, beyond which on the posterior-dorsal 
slope the ribs again become beaded as they approach the dorsal 
margin; the interspaces on the right valve are wide, and on the 
shell disk anterior to the umbonal angle they carry a fine, but 
often faint interstitial thread; the intervals are otherwise smooth 
or finely etched with concentric lines; hinge-lines straight with 
numerous small, vertical teeth and a wide, smooth cardinal 
area. 
Length 18, height 18, semi-diameter 9.5 mm. 

A smaller species than the following (//zzdsz) and easily 
distinguished by the interstitial thread of its right valve. It is 
much less common than the other two species. 

Named for Dr. E. R. Lloyd, formerly of the United States 
Geological Survey, and who was a member of the first Hxplora- 
tion Party of the Sinclair Oil Company in Panama and Costa 
Rica. 

Gatun Stage: Hill 3, Banana River. 

Zone 5, ked Cliff Creek. 
Coll. 7, Pumbri Creek. 


Arca Hindsi, n. sp. Plate 24, figures 7-9 


Shell small or of medium size, cordiform and similiar in out- 
line to the preceding species but larger; umbos high and wide, 
and but slightly anterior to the middle; the left valve has about 
26 strongly nodulated ribs, separated by interspaces of slightly 
greater width; the first 9 or 10 ribs ef the right valve are nodu- 
lated, followed by smooth ones to the umbonal angle, beyond 
which the remaining 7 become nodulated as they approach the 
dorsal margin; interspaces wider than the ribs, smooth and 
without any interstitial thread; hinge-line straight with numer- 
ous, small vertical teeth and a wide, smooth cardinal area; on 
some shells the ligamental area may carry one or two, small 
diamond-shaped grooves. 


Length 24, height 25, semi-diameter 14 mim. 


A much larger species than the Z/oyd7 and without any in- 


194 BULLETIN 3G 266 


terstitial thread in the interspaces of its right valve. It is nam- 
ed ror Dr. Henry Hinds, formerly of the United States Geologi- 
cal Survey and the Exploration Department of the Sinclair Con- 
solidated Oil Corporation. 
Gatun Stage: Hill No. 3, Banana River. 

Zone f, Saury Creek. 

Coll, 2, East Grape Point Creek. 

Coll. 3, East Grape Point Creek. 

Rio Betey. 


Subgenus NOETIA, Gray 
Arca MacDonaidi Dall Plate 25, figures 4-7 


Area (Noetia) MacDonaldi Dall, 1912, Smith, Misc. Coll., vol. 59, No. 
PISO (0) 

This large Voétia, is very common and characteristic of the 
Gatun formation in many parts of Northern Costa Rica, and 
large specimens may reach a length and height of 55 mm or 
more. It was originally described by Dall from specimens col- 
lected by Pittier and MacDonald from the Gatun beds of the 
Banana River, where it is associated with Avca Pittieri and is 
very abundant. 


Two forms or varieties may be recognized. In typical J7Zac- 
Donaldi, the shell is high, trigonal, with elevated, narrow umbos 
and a producted anterior extremity. ‘The beaks are separated from 
the hinge-line by a wide ligamental area. ‘This is the conimon 
form along the Banana River. Frequently associated with typi- 
cal MacDonaldi are shells'with a more rectangular outline, lower 
but wzde umbos. ‘The beaks are close to the hinge line so that 
the ligmental area ts narrow. These two types have a very dif- 
ferent appearance but seem to be connected by transitional forms. 
To the last variety, I propose the name of subreversa. 


Among recent shells, the 47ca MacDonald? finds its closest 
relation with the common Avca reversa Sowerby of the Pacific 
coast of Central America. 


367 Costa RicA MrockNE—OLSSON 195 


Gatun Stage: Hill 1, ra, Banana River, typical and var. 
suberversa 
Coll. 7, Pumbric Creck (typical) 
Coll. 6, Red Cliff Creek (typical) 
Betey (subreversa) 
Zone G, Saury (subreversa) 


Genus PTERIA, Scopoli 


Pteria inornata Gabb Plate 28, figure 9 
Avicula inornata Gabb, 1873, Trans. Amer. Phil. Soc., vol., 15, p. 
253. 
Pteria inornata Dall, 1898, Trans. Wagner Free Inst. Sci., vol. 3, pt. 
4, p. 669. 
Pteria tnornata Maury, 1817, Bull. Amer. Pal., vol. 5, p. 345, pl. 26, 
fig. 14. 


This species is locally common in Costa Rica, but generally 
difheult to collect because of its thin and fragile shell. Its 
valves are as high as long, strongly convex and smooth. The 
anterior ear is of medium size, the posterior much larger and 
produced. 

It was described as a Miocene fossil from Santo Domingo, 
but has also been collected at Bowden. 


Uscari formation. Old Harbor. 
Gatun Stage: Zone 3; Red Cliff Creek. 


Genus OSTREA, Linnzus 
Ostrea megodon Hanley Plate 18, figure 1 


Ostrea megodon Hanley, 1845, Proc. Zool. Soc. London, p. 106. 

cf. Ostrea cerrosensis Gabb, 1869, Geol. Surv. Calif., vol. 2, p. 35, pl. 
II, fig. 61. 

Ostrea megodon Dall, 1897, Trans. Wagner Free Inst., vol. 3, pt. 4, p. 
685. 

Ostrea megodon Maury, 1917, Bull. Amer. Pal., vol. 5, p. 347, pl. 34, 
LGR Be 


196 BULLETIN 39 363 


This is a medium sized and sickle-shaped oyster with 4 or 5, 
large, tooth-like folds on the anterior and ventral margins. The 
beak and ligamental area are directed upwards and sometimes 
strongly to the posterior side. Its external surface, although 
strongly and deeply folded, is plain and nearly smooth. 


The Ostrea megodon isa recent species along the west or 
Pacific coast of Central America. It occurs also as a Miocene 
fossil in Santo Domingo and in the Bowden beds of Jamaica. 


Gatun Stage: Water Cay. 
Coll. ga, East Grape Point Creek. 
fiill ra, Banana River. 


Ostrea costaricensis, n. sp. Plate 21, figure 2 


Ostrea sculpturata Bose, 1906, Boll. de Inst. Geol. de Mexico, numero 
22, p. 76, pl. 9, ig. 4. Not of Conrad, 184o. 

Shell of medium size, subcircular in form but somewhat ob- 
lique; the shell is somewhat flexed or slightly saddle-shaped, the 
concave side of which corresponds to the left valve; both valves 
are strongly sculptured, the right with narrow, rib-like costz 
and wider interspaces, the left valve with the reverse of the 
sculpture of the right; the ribs are more or less nodose or knob- 
bed and are crossed by irregular concentric lines, which in some 
cases may become raised or lamellar-like; the hinge margin is ir- 
regularly crenulated: 

Height 50, length 47, diameter I9 mm. 

The type is aspecimen from the Gatun of the Rio Blanco 
with both valve tightly closed so that the interior cannot be 
seen. What appears to be the same species, is represented by 
several sessile and free valves from East Grape Point Creek, and 
which are associated with O. megodon. 


It differs from the Ostrea gatunensts - haitensis group by its 
smaller size; less heavy shell and in that the ribs of the valves 
are narrow and coste-like and not long, strong plicee. Bose 
has recorded and figured this species as Ostrea sculpturata Con- 


369 Costa Rica MIocENE—-OLSSON 197 


rad from the Miocene (as Pliocene) of the State of Oaxaca, 
Mexico. Itis very distinct from the East Coast, Chesapeake 
Miocene Os/rvea sculpturata both by its form and sculpture. 


Gatun Stage: Rio Blanco. 

Coll. ga, East Grape Point Creek. 

The collection also contains fragments or much decayed 
specimens of two or more large oysters, one of which is probably 
the gatunensis Brown and Pilsbry of the Canal Zone, and a 
large, heavy selleformis-like species, possibly the Galbertharrist 
Maury of the Dominican Miocene. The specimens are however 
too fragmentary for positive determination. 


Genus PECTEN, Muller 
Pecten gatunensis Toula Plate 16, figures 3, 4 


Pecten (Flabellipectcn) gatunensis Toula, 1908, Jahrb. der K-K Geol. 
Reichsanstalt Wien, vol. 58, p. 711, pl. 29, fig. 2. 


Pecten gatunensis Brown and Pilsbry, 1911, Proe. Acad, Nat. Sci. 
Phila., vol. 63, p. 365. 

Shell rather large, subcircular; the left valve is nearly flat 
or slightly vaulted transversely across the middle of the shell; 
the right valve is moderately convex; the ears are small and 
nearly equal,those of the right valve slightly flaring at the sides; 
the right valve has 23 or 24 fairly strong ribs separated by inter- 
spaces a little more than % of their width; in large right valves, 
some of the central ribs become dichotomously or trichotomously 
divided but this feature is not always developed to the same ex- 
tent on all shells; the left valve has 18 or 19 ribs which fade out 
on the wide dorsal submargins; the ribs of the left valve are 
narrow with the interspaces nearly twice as wide; the surface is 
covered with fine, even, raised concentric threads best preserved 
on the left valves; interior fluted. 


Length 62, height 60? diameter 12.5 mm. 
62 60? 4 mm (left valve) 


198 BULLETIN 39 37° 


We have redescribed this species as Toula had but a speci- 
men of the right valve. Itisa true Pecten, distantly related to 
the Pecten hemicyclicus Ravenel from the Chesapeake Miocene of 
Florida and the Carolinas, but is smaller and less broad. In both 
species, the ribs on the right valves of large specimens become 
divided in the same manner. 

Gain Stage Gatun GC. Z: 

Water Cay, 


Pecten MacDonaldi, n. sp. Plate 16, figures 1, 2 


Shell large, subcircular; ears of medium size and equal; left 
valve is slightly convex due tothe middle of the shell being 
transversely humped or vaulted and a depressed zone follows on 
the inner slope of each of the raised dorsal submargins; the 
right valve is slightly but evenly convex; sculpture of the right 
valve consists of about 26, low ribs which widen out as they 
approach the ventral margins; their interspaces are at first nearly 
as wide as the ribs themselves but become only % or 4% as wide 
ventrally; the left valve has 21 or 22 narrow ribs and wider inter- 
spaces; on the dorsal submargins, the ribs are small and fade 
away; surface with fine, even, raised lines best seen on the left 
valve. 


Length 106, height 97, semi-diameter 12 mm. 
83 80 mim. 

The fragmentary Toro limestone at Gatunand Toro Point 
contains very few good fossils,although the rock itself is compos- 
ed almost entirely of broken and ground fragments of shells and 
barnacles. Dall has described an Afztonzum toroensis from the 
Toro Point limestone where it is fairly common. ‘The Pecten 
MacDonald is found in the Toro limestone which caps the hills 
just west of the locks at Gatun. 

This species has much the same contour as the recent Pecten 
maximus Linné, from Europe. It isa larger species than the 
Pecten gatunensts Toula. 


Toro Limestone. Gatun, west of the locks. 


Sur Costa Rrcsa MrockNE—OLSSON 199 


Pecten coralliphila, n. sp, Plate 17; figures 5, 6 


Shell small, very thin and generally translucent; broader 
than high; right valve moderately convex, the left concave with 
raised submargins; ears small and subequal; sculpture of the 
right valve of about 23 low, flat ribs, with flat interspaces a 
little more than ¥%4 the width of the ribs; some of the ribs may 
become divided medially; the ribs of the left valve, about 20 in 
number, are narrow, usually somewhat irregularly spaced so 
that their interspaces are of variable width and new ribs may 
occasionally appear inthe wider interspaces near the ventral 
margin; surface in addition finely sculptured with even, raised 
concentric threads; interior of the shell fluted well into the cav- 
ity of the shell, in harmony with the external ribs. 


Length 34, height 29, semi-diameter 4 mm (right valve) 


This Pecten is fairly common in the coralline phases of the 
Miocene and its associated marls and sands. It is extremely thin 
and fragile in texture. Its sculpture resembles that of /Pecten 
gatunensts Toula but is much more irregular. The interspaces 
of the left valve are of very variable width. 


Gatun Stage: Port Limon. 


Subgenus CHLAMYS, Bolton 
Section PLAGIOCTENIUM Dall 


Pecten levicostatus Toula Plate 16, figures 5-7 


Fecten tevicostatus Toula, 1908, Jahrb. der K-K Geol. Reichsanstalt, 
Wien, vol. 58, p. 713, pl. 26, figs. 4, 5, 6. 

FPecten (Chlamys) santarosanus Bose, 1906, Boll. de Inst. Geol. de 
Mexico, numero 22, p. 73, pl. 6, figs. 3,4, ? FPecten (Chlamys) 
santarosanus Bose, op. cit. p. 23, pl. 1, figs. 1 and 4, 2 and 5. 

A scallop of the general type of Pecten ventricosus Sowerby; 
both valves equally and strongly convex; ribs about 21, which 
are square or rectangular in section on the umbos and with deep 
interspaces; the ribs generally flatten or become more rounded in 


206 BULLETIN 39 372 


large shell near the ventral margins; the interspaces are covered 
with fine, raised, concentric lines, best seen on the umbos or on 
young shells; they become more crowded towards the ventral 
margins and may pass over the tops of the ribs as well; the dor- 
sal and posterior submargins are strongly flattened and sculp- 
tureless; the anterior ear is a little larger, projects forward and 
carries, in the right valve, a byssal notch; it is sculptured with 
several radial riblets; the posterior ear has its upper half nearly 
smooth or but faintly sculptured, but its lower half carries a 
band of 5 or 6 strong, radial cords. 
Height 29, breadth 34, thickness 10.50 mm (right valve) 
34 38 23 (both valves) 
39 43 T4 (left valve) 


This is the common /Pecten in Costa Rica and frequently 
forms zones in the lower part of the Gatun formation. All our 
specimens have 21 ribs, which vary from strongly rectangular 
in section with deep interspaces to more flattened and rounded 
on the larger individuals. 


Bose has described as Fecten santarosanus, a small scallop 
shell from the Miocene of the State of Oaxaca, Mexico. The 
ribs number 23 to 25. Later in the same volume, he figured a 
larger Pecten from the Miocene of La Barranca de Santa Marie 
Tatella, as Pecten santarosanus. ‘This shell has but 21 ribs and 
is equivalent to Toula’s Jevicostata. If this larger specimen of 
Bose’s is equivalent to his typical saztarosanus, this name must 
of course take priority over Toula’s levicostatus. As Bose’s typi- 
cal santarosanus has 23 to 25 ribs, whilein /evicostatus the ribs 
appear to beconstantly but 21 in number, they probably repre- 
sent two different species. 


In the Dominican Miocene J/evzcostafius is represented by the 
Pecten excentricus Gabb, which appears to differ constantly by its 
lower and more rounded ribs and smaller ears. The recent 
Pacific Pecten ventricosus Sowerby is closely related and probably 
the direct descendent of /evicostatus. In that species, the ribs 
are low and rounded as in excenfricus and as occasionally seen in 


373 Costa Rica MiocENE—OLSSON 201 


large examples of /evicostatus. ‘The sculpture of the posterior 
ear is uniform. Inthe Chasapeake Miocene fauna the /ecten 
eboreus Conrad, variety comparilis Tuomey and Holmes is dis- 
tantly related, but is larger, less convex and generally with 
more numerous ribs. 
Gatun Stage: Gatun, C. Z. Quitana Creek. 

Water Cay. 

Flone Creek. 

Banana River. 

ftstrella River. 


Pecten costaricensis, n. sp. Plate 18, figures 8-11 


Shell small, inequivalve, oblique; the right valve is moder- 
ately convex, the left but slightly; right valve with 17 flat-top- 
ped ribs, square in section and with deep U-shaped interspaces 
of a width equal to the ribs; these interspaces are sculptured 
with fine, concentric, raised lines which are quite far apart on 
the umbos, but become crowded ventrally or disappear entirely; 
the left valve has 17 or 18 ribs which are lower and more round- 
ed in section; the fine concentric lines on the left valve are con- 
fined to the umbos, the interspaces over the greater part of the 
shell being smooth; the anterior and posterior dorsal slopes are 
flattened and smooth, but sculptured below in each valve by 
strong radial threads on the lower side of the ears; ears subequal; 
the right valve with a deep, byssal notch in its anterior ear; the 
left valve has the lower half of each ear with 3 strong radial 
threads, while on the upper half, the radial threads are largely 
obsolete; valves internally grooved in harmony with the exter- 
nal ribbing to about the center of the shell. 


Height 26.50, breadth 28, thickness 7.50 mm (right valve) 
24 24 4.00 mm (left valve) 
2g 28 5.50 mm (left valve) 


Differs from the Pecten levicostatus Toula, by its unequal 
valves, less convexity and fewer ribs. It is somewhat like the 
Pecten ineqgualis Sowerby, from which it is distinguished by its 


202 BULLETIN 39 374 


more flattened left valve and lower, smaller ribs of its right. In 
inequalis, the fine, lamellose, concentric lines cover the entire 
interspaces between the ribs on both valves, while in the present 
species the interspaces of the left valve are largely smooth with 
the concentric lines confined to the extreme umbonal area. 
Gatun Stage: Coll. 4, Red Cliff Creek. 
ZI mile south of the beach, along Old Man Sam 
Creek. 
Coll. 4, East Grape Point Creck. 
Rio Blanco. 


Section AEQUIPECTEN, Fischer 
Pecten (Aequipecten) preglyptus, n. sp. Plate 17, figures 2, 7 


Shell of moderate size, subcircular in form and convex; ears 
of usual size, subequal and with a straight hinge-line; sculpture 
of about 16 or 17 wide, little elevated ribs and narrower, scarce- 
ly distinguishable interspaces; the surface is otherwise smooth 
or with fine, concentric lines; interior of the shell with 29 or 30 
Amustum-like internal liree which are evenly spaced and not in 
pairs; these liree are very faint in the middle of the shell cavity 
but strong distally. 


Length 50, height 49, semi-diameter 6.5 mm. 


An Amustum-like species found in the lower sandstones and 
conglomerates of the Gatun formation in upper Cocles Creek. 
It is closely related to the recent deep-water Pecten glyptus Ver- 
rill from the cost of Hatteras and Marthas Vineyard (see Dall 
Brocy US Nat-iMiusiivol Mas pe 2asiuple) Ss. . fgs) 22) bot 
species having about the same number of low, wide ribs and in- 
ternal liree. ‘The fossil shell seem to have been nearly circular 

_ while glypfus is decidely oblique in outline. 

Gatun Stage: Upper Cocles Creek. 


375 Costa R1cA MIOCENE 


OLSSON 203 


Subgenus PSEUDAMUSIUM, H. and A. Adams 
Section PSEUDAMUSIUM, s. s. 


Pecten almirantensis, n. sp. Plate 18, figure 16 


Shell small, thin, compressed or but slightly convex; the 
posterior and anterior dorsal slopes are straight, meeting the 
beaks at an acute angle; the right valve is finely sculptured with 
close, regular, concentric lamella and wider interspaces; the 
posterior and anterior sides show a few, irregular, radial threads 
which cross the concentric lamellze; the ears are unfortunately 
broken; the interior of the shell cavity is filled with a thin de- 
posit of callus; ventral margin plain; hinge line minutely groov- 
ed with vertical lines. 


Length 4.75, height 4.75, diameter of right valve 60 mm. 
This small, elegant species is represented solely by an im- 
perfect right valve and represent the first true Pseudamusium to 
be discovered in the Miocene beds of Panama. Its sculpure is 
like the recent deep water Pseudamustum strigillatum Dall from 
the West Indies, but differs in having its anterior and posterior 
dorsal submargins meeting at the beaks at a much more acute 
angle. 


Gatun Stage: Bocas del Toro. 


Section CYCLOPECTEN, Verrill 
Pecien oligolepis Brown and Pilsbry 


Fecten aff. subhyalinus Smith, Toula, 1911, Jahrb. der K-K Geol. 
Reichsanstalt, Wien, vol. 61, p. 492, pl. 31, figs. Ia, b, c. 


Fecten (Cyclopecten) oligolepis Brown and Pilsbry, 1912, Proc. Acad. 
Nat. Sci. Phila., vol. 64, p. 512, text-figure 5. 

A very small species with broad, nearly smooth valves and 
large ears. The right anterior ear has a deep byssal notch and 
its surface is sculptured with fine, raised, concentric, lamellose 
lines. The ears of the left valve are similiarly sculptured but fin- 


204 BULLETIN 39 376 


er. The surface of the valves appear nearly smooth but in well- 
preserved specimens very faint concentric lines and fine radial 
strisee may be seen. The left valve is more coarsely sculptured. 
The hinge is finely grooved with vertical lines. The measure- 
ments given by Pilsbry for his Gatun specimen, length 2.8, 
height 2.7, diameter of right valve .8 mm, are the usual dimen- 
sions for the Costa Rican specimens. 


Gatun Stage: Gatun, C. Z. 
Coll. 4, East Grape Point Creek. 
Middle Creek. 


Pecten aotus, n. sp. Plate 18, figures 17, 18 


Shell small, thin, depressed or but slightly convex and 
Amusium-like; both valves except on magnification appearing 
smooth; ears of the left valve large and subequal; the anterior 
ear of the right valve about twice that of the posterior and car- 
rying a deep, byssal notch; the surface of the left valve on mag- 
nification is seen to be sculptured with fine, radial lines which 
often occur in irregular streaks or blotches, imparting a peculiar 
appearance to the shell; the right valve is perfectly smooth and 
featureless, except for a band of rough scales on its anterior ear 
above the byssal notch; interior of shell cavity shallow, a simple 
ventral margin and the hinge-line vertically grooved with fine 
lines. 


Length 4.5, height 4.5, diameter .75 mm. left valve. 


Differs from Pecten oligolepis Brown and Pilsbry in its larg- 
er size, thinner and more Amuszum-like shell, and strongly sculp- 
tured left valve. This sculpture consists of fine lines or camp- 
tonectes striations, which may be uniformly distributed but more 
usually occur in irregular streaks or blotches, imparting a dis- 
tinctive appearance to the surface. This sculpture is seen only 
on a slight magnification, the shell otherwise appearing smooth 
and featureless. It is quite common in the coralline limestones 
of Port Limon. 


377 Costa RicA MrocENE—OLSSON 205 


Gatun Stage: Port Limon. 
Coll. 5, Red Cliff Creek 


Genus AMUSIUM, Bolton 


Amusium luna Brown and Pilsbry Plate 17, figure I 


Amusium Mortont Bose, 1906, Boll. de Inst. Geol. de Mexico, Numero 
22, p. 24, pl. 1, figs. 3,6, 7,9. Not Amusium Mortoni Ravenal, 


1844, 
Amusium Mortoni Bose, op. cit. p. 74, pl. 8, figs. 1, 2; pl. 3, fig. 3. 
Pecten (Amusium) cf. Mortoni Toula, 1908, Jahrb. der K-K Geol. 
Reichsanstalt, Wien, vol. 58, p. 714, pl. 26, figs. 8, 9. 


Pecten (Amustum) luna Brown and Pilsbry, 1912, Proc. Acad. Nat. 
Sci. Phila., vol. 64, p. 514, pl. 23. fig. 1. 


This is the common Gatun Amusium in Panama and Costa 
Rica. Its valves are nearly equal and but slightly convex; ears 
equal and small, and defined from the rest of the shell by a 
sharp ledge; the umbos, as well as the rest of the shell disk are 
smooth, except for the faint lines of growth; the internal ribs 
are in pairs, numbering about 23 and spaced at intervals nearly 
twice the width of the pairs of ribs themselves. 


Bose and Toula both referred this species to the Upper 
Chesapeake Miocene and recent Amusium Mortont Ravenel, 
from which it is very distinct. Amuszum Mortoni is not only 
much larger (height 170 mm), but is porportionately much 
broader, and with larger ears which are defined simply by a line 
from the rest of the shell. 


The Dominican A. papyraceum Gabb has larger ears, the in- 
ternal ribs are more crowded and the left valve is generally 
strongly flexed along the anterior and posterior submargins. 
Amustum Toxule Brown and Pilsbry, is found in the Gatun of 
the Canal Zone. Its surface is marked with dark-colored rays 
and is said to contain no internal ribs. 


206 BULLETIN 39 378 


Gatun Stage: Gatun, C. Z. 
Flotel Creek. 
Zone 6, Old Man Sam Creek. 
Coll. 4, East Grape Point Creek. 


Amusium bocasensis, n. sp. Plate 17, figures 3, 4 


Shell rather small, thin and subcircular in outline; moderate- 
ly convex; ears proportionately larger than in 4. /una, and with 
their dorsal margins slightly inclined upwards from the beaks to 
their outer edges; the surface is smooth and polished, finely 
covered with concentric growth-lines and faintly showing the 
position of the internal lire; interior of shell with about 34 
lire, which are usually nearly evenly spaced and not in pairs; 
the internal lire commence well in the interior of the shell, but 
are strongest distally. 


Height 4o, breadth 43. 
25 38, thickness 6 mm. (right valve) 


A much smaller and more delicate species than the Amuszum 
luna Brown and Pilsbry. The internal liree are more or less 
equally spaced and not in pairs. It differs from Amuszum Lyonit 
_Gabb from the lower Miocene of Sapota, Costa Rica by its much 
smaller ears. 


Gatun Stage: Bocas del Toro. 


Genus SPONDYLUS, Linnzeus 
Spondylus chiriquiensis, n. sp. Plate 20, figures 1, 2, 5, 6 


Shell large, ponderous, equivalve and strongly convex; both 
valves of the same convexity, but with the beak of the right, 
high and projecting above the strongly incurved beak of the left; 
in large shells both valves are provided with a cardinal area, 
that of the right being both high and wide and for the greater 
part straight but the beak curving over it toa small extent at 
its upper end; the cardinal area of the left vale is much smaller, 


379 Costa RicA MIocENE—OLSSON 207 


narrow and appears only with maturity and is more or less hid- 
den by the strongly incurved beak; the sculpture consists of 
strong radial cords and small ribs divisible into 3 sets; a primary 
set of 7, which in perfect specimens and in the young carry large, 
strong spines; a secondary thread in each of the wide interspaces 
and when perfect bearing smaller spines, and tertiary threads of 
a variable number between the secondaries and primaries; the 
whole surface is in addition, finely, longitudinally starited with 
minute, scale-bearing threads; young shells have a small foliace- 
ous area on the umbo of the sessile or right valve and strong 
spines on both valves. 
Height 151, length 120, diameter 115 mm. 

A large, ponderous species with nearly equal and strongly 
convex valves which show no distortion due to attachment | 
Young shells associated with the type have a small, somewhat 
foliaceous attachment area on the right valve, and each valve 
bears large, strong spines on its primary set of radial cords and 
finer spines on its secondary and even on some of the tertiary 
threads. 

Of described species, the chzrigutensis seems nearest related 
to the dostrychttes Guppy from the Miocene of Santo Domingo 
and Jamaica but differs by its much larger size, proportionately 
more convex whorls and by its cardinal area which is more up- 
right and with a more incurved beak at itsupper end. The 

sculpture of chtviqutenszs is more spiny,the spines being borne by 
the primary radials and to a less degree by the secondaries, and 
even by the tertiaries. In dostrychites only the primaries bear 
spines, the others appearing as large, nearly smooth, intermedi- 
ate threads and cords. We have specimens ofa large, closely 
allied Spondylus from the Chesapeake Miocene of Florida, dif- 
fering mainly in its cardinal area and in details of sculpture. 


Gatun Stage: Water Cay. 


Spondylus gumanomocon Brown and Pilsbry Plate 21 figure I 


Spondylus americanus Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, 
p. 257. Not of Lamarck. 


208 BULLETIN 39 380 


Spondylns gumanomocon Brown and Pilsbry, 1912, Proc. Acad. Nat. 
Sci. Phila., vol. 14, p. 514, footnote. 


Spondylus gumanomocon Maury, 1917, Bull. Amer. Pal., vol. 5, p. 
355- 

This is a species with very unequal valves, that of the left 
or upper, being small, pectiniform and lacking ina cardinal 
area, while the right or lower valve is strongly convex, with a 
high produced beak and a high cardinal area. The sculpture of 
the two valves is similiar, except that the umbo of the right is 
strongly foliaceous. It occurs in the Miocene of Santo Domin- 
go, where it was referred to the recent S. americanus by Gabb. 


Gatun Stage: Coll. 2, Hone Walk Creek. 
Port Limon. 


Genus PLICATULA, Lamarck 


Plicatula marginata Say Plate 28, figures 6, 7 


Flicatula marginata Say, 1824, Journ. Acad. Nat. Sci. Phila., vol. 4, 
pp. 136-137, pl. 9, fig. 4. 

Plicatula marginata Dall, 1898, Trans. Wagner Free Inst. Sci., vol. 3, 
pt. 4, p. 764. 

The Panama and Costa Rican examples are not typical but 
they approach more closely the Plicatula marginata Say than 
they do the densata which Dall has recorded from the Bowden 
beds of Jamica. 


The shells vary from subcircular to elongate and in some 
cases carry the dark, marginal band so frequently seen in typical 
marginata, ‘The ribs number five to seven; the shells with more 
rounded form and more numerous ribs suggest the dexsata, but 
the ribs are higher and more foliaceous, and can be exactly du- 
plicated by scores of true marginata in the Cornell collection. 
The above determination must however be considered as purely 
provisional at the present time. 

The typical P. densata was described by Conrad from the 
lower Chesapeake Miocene of New Jersey, but it also occurs in 


381 Costa Rica Mi0cENE 


OLSSON 209 


the synchronous Calvert formation of Maryland. It is also found 
as a lower Miocene fossil in Florida and Dall recorded it from 
the Bowden beds of Jamaica and from the Oligocene of Gual- 
lava, Costa Rica. The ?. marginata Conrad is an Upper Chea- 
peake Miocene and Pliocene fossil of the eastern United States. 
Gatun Stage: Water Cay, 
Old Man SamCreek, 1 mile south of the beach. 


Genus LIMAEA, Born 
Limaea solida Dall Plate 28, figure 10 


Limea solida Dall, 1898, Trans, Wagner Free Inst. Sci., vol. 3, pt. 4, 
p. 769, pl. 35, figs. 4, 5. 


A small, solid shell, externally sculptured like a small Gly- 
cymeris. It has 12 rounded ribs and a coarsely crenulated basal 
margin. Our single specimen from Costa Rica isa little smaller 
than Dall’s specimens from Bowden. 


Gatun Stage: Coll. 4, Red Cliff Creek. 


Genus ANOMIA, (Linné) Muller 
Anomia simplex d’Orbigny Plate 21, figure 6 


Anomia ephippium Conrad, 1845, Fossils of the Medial Tertiary, p. 
75, pl. 43, fig. 4. Not of Linné. 


Anomia simplex d’Orbigny, 1845, Hist. Pol. y Nat. Isla de Cuba, p. 


367. 

Anomia ephippium Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 
257: 

Anomia simplex Dall, 1898, Trans. Wagner Free Inst., Sci., vol. 3, 
pt. 4, p. 784. 


Anomia simplex Bose, 1906, Boll. de Inst. Geol. de Mexico, numero 
22, p. 25, pl. 2, figs. 18-23. 
Anomia simplex Maury, 1917, Bull. Amer. Pal., vol. 5, p. 355, pl. 26, 
fig. 15. 
The Anomia simplex is both common and characteristic in 
the Gatun of Costa Rica as well as in certain parts of the Prov- 


210 BULLETIN 39 382 


ince of Colton, Panama, near the Canal Zone. It frequently 
gives rise to important zones in the lower part of the Gatun for- 
mation, where it occurs in large numbers associated with Pecten 
levicostatus Toula. 

The specimens are generally large, thin-shelled, smooth and 
irregular in form, due to the object on which they were attach- 
ed. They average for the upper valves about 35 mm in 
height. 

Anomia simplex appears as a Miocene fossil in the eastern 
United States for the first time in the St. Mary’s formation of 
Maryland in the upper part of the Lower Chesapeake. It occurs 
throughout the Upper Chesapeake Miocene and Pliocene and 
passes into the recent fauna. Locally asa fossil it may become 
very abundant, as at the base of the Pliocene overlying Upper 
Chesapeake Miocene along the shores of Lake Waccamaw, 
North Carolina. It occurs rarely as a Miocene fossil in Santo 
Domingo, where it has been recorded by Gabb and Maury. In 
Mexico, it occurs in the Miocene of Santa Rosa, Vera Cruz as 
figured by Bose. 

Gatun Stage: Several localities, Prov. of Colon, Panama. 

Middle Creek. Comadre Creek. 
Fone and Hotel Creeks. 
Hill No. 2, Banana River, Soury Creek, ete. 


Genus PLACUNANOMIA, Broderip 
Placunanomia lithobleta Dall Plate 21, figures 3, 4, 5 


Placunanomtia lithobleta Dall, 1898,Trans. Wagner Free Inst. Sci.,vol. 
3; Ptr 4p. 778: 

Placunanomia lithobleta Maury, 1917, Bull. Amer. Pal., vol. 5, p. 356, 
Di s4A igs ine: 

The Costa Rican specimens are frequently widely plicate, 
nearly as much asin the Duplin Miocene P. plicata Tuomey and 
Holmes, but the surface is marked with fine, wavy, radial threads 
so that even very small fragments of this species are very easily 
recognized. It occurs in the Miocene of Jamaica, its type local- 
ity, and in the Miocene of Santo Domingo. 

Gatun Stage: Hill 3, Banana River. 


383 Costa RrcA MrockENE—OLSSON 211 


: ORDER ANOMALODESMACEA 


Genus THRACIA, Blainville 
Subgenus CYATHODONTA, Conrad 
Thracia Tristani, n. sp. Plate 20, fig. 3 


Shell thin, subquadrate, the left valve slightly convex; the 
beaks situated at the posterior 1-3 of the shell, are low and in- 
conspicuous; anterior end, wide and broadly rounded, the poster- 
ior contracted, depressed and truncated at its extremity; the sur- 
face is sculptured with regular wave-like folds, which on the 
middle of the shell are concentric, but are oblique on the anter- 
ior portion; they are lacking from the posterior area; in addi- 
tion the whole surface is very finely pustulose and crossed by 
very indistinct growth-lines. 


Length 42, height 30, diameter of the left valve 7.00 mm- 


The Cyathodonta gatunensis Toula of the Canal Zone isa 
similiar but larger species, reaching a length of 52 mm. It dif- 
fers from the Costa Rican species in its greater proportional 
length, the more central beaks and in its sculpture, which ac- 
cording to Toula’s figure is strictly concentric. 

The C. undulata Conrad, a recent species from the Gulf of 
California, is sculptured like 77zsfanz with oblique riblets but 
the left valve is depressed and strongly flattened. 

This species is named for Professor J. Fidel Tristan, Director 
of the National Museum of Costa Rica. 


Gatun Stage: Rio Betey. 


212 BULLETIN 39 384 


ORDER TELEODESMACEA 


Genus CRASSATELLITES, Kruger 


Crassateilites Reevei Gabb Plate 20, figure 4 


Crassatella antillarum Reeve? C. Reevet Gabb 1873, Trans. Amer. 
Phil. Soc. vol. 15, p. 252. 


Crassatellites reevei Brown and Pilsbry, 1913, Proc. Acad. Nat. Sci. 
Phila., vol. 64, p. 515, pl. 23, fig. 5. 


The Crassatellites Reevei Gabb is not an uncommon fossil 
in the Gatun beds of western Panama and Costa Rica. Our 
specimens agree quite closely with Brown and Pilsbry’s figure of 
the single right valve collected by Gabb in the Miocene of Santo 
Domingo. Young and full-grown specimens show a strong 
posterior angulation which extends from the beak to the poster- 
ior ventral extremity. “The posterior end is obliquely truncated 
but less so in young shells. ‘The surface sculpture consists of 
even, concentric costze which on the slightly flattened umbos are 
a little wider apart asin the Cvassatedlites of the section Scam- 
bula Conrad. In this later character, our shells disagree with 
Brown and Pilsbry’s redescription of Gabb’s type specimen, in 
which the sculpture is said to extend without any irregularity 
upon the rather flattened umbos. 


The C. mediamericanus Brown and Pilsbry from Sapote, 
Costa Rica is an earlier shell and is higher, less obliquely trun- 
cated behind, more coarsely sculptured and lacks the posterior 
angulation. In these characters except the lack of the posterior- 
umbonal angulation it is like the young shells of Reevez. 


Length 55, height 36.5, semi-diameter 10.5 mm. 
50 35 diameter Pe iGaabonl, 
58 42 seni-diameter 14 mm. 


385 Costa RicA MI0ockENE—OLSSON 202 


Gatun Stage: Water Cay. 
Old Man Sam Creek, Coll. No. 6: 


Banana River. 


Subgenus CRASSINELLA, Guppy 


Crassatellites bowdenensis Dall, variety costaricensis, n. var. 
Plate 29, figure 12 


Cf. Crassatellites (Crassinella) bowdenensis Dall, 1903, Trans. Wagner 
Hreevinsty sci Vol) 3 pt. GN p.T4 76s" pla) 50st eSy Qua. 


Shell small, triangular, slightly convex, subequilateral, the 
anterior end generally a little longer and more pointed; beaks 
triangular, slightly pointing forwards; lunule large, narrowly 
lanceolate, smooth; escutcheon small; surface sculptured with 
small, even, concentric riblets which on some shells may be 
strong and persistent over the greater part of the shell, or on 
some specimen sub-obsolete; interior of shell cavity shallow, with 
a moderately heavy hinge. 

Wength!)2o0, height 2.75, diameter }75 mm: 
Ba5O) 250) 1715) Mal 

The typical dowdenenszs Dall from the Bowden Miocene of 
Jamaica is a more solid, more convex and triangular shell. The 
variety costaricensis is very abundant at several localities in Costa 
Rica in certain horizons. The sculpture varies from strong, 
even, concentric riblets covering the greater part of the shell, to 
forms which are nearly smooth. 

Gatun Stage: Hiill ra, Banana River. 

Middle Creek. 
Zone 3, Quttana Creek. 


Crassatellites midiensis, n. sp. Plate 20, figure 11 


Shell small, trigonal, slightly convex and equilateral; the 
beaks are central and erect; the dorsal margins straight and 
meeting at the beaks nearly at right angles; the surface is sculp- 
tured with subregular, distant, concentric lamelle-like riblets, 


214 BULLETIN 39 386 


numbering about 10; the interspaces carry irregular, finer, con- 
centric lines; lunule narrowly lanceolate, with the escutcheon 
narrow and smaller; interior of shell cavity shallow, with a mod- 
erately heavy hinge. 


Length 3.25, height 3.25, diameter of one valve 1.00 mm. 


A small, rare species, somewhat like the C. Guppyz Dall of 
the Miocene of Jamaica, but more trigonal, with more central 
beaks and somewhat different sculpture. The sculpture and 
form are very similar to specimens of C. galvestonensis Harris in 
the Cornell Collection from the Chesapeake Miocene of York- 
town, Va. ‘This latter species is generally much worn but per- 
fect specimens have a coarse concentric sculpture like that of 
lunulatus Conrad. ‘The Costa Rican shells are however very 
much smaller. 


Gatun Stage: Middle Creek, C. R. 
Coll. 4, East Grape Point Creek. 
Water Cay, Panama. 


Genus CARDITA, (Bruguiére) Lamarck 
Cardita caribbeanensis, n. sp. Plate 26, figure 22 


Shell small, moderately convex, elongate, its length a little 
less than twice its height; beaks very near the anterior end are 
small but distinct; an umbonal ridge extends from the beaks to 
the posterior-ventral extremity and along which and bordering 
the ribs are largest; the right valve with about 20, hight, narrow 
and sharply nodulose ribs, separated by wide, triangular, smooth 
interspaces; 6 of the ribs are situated on the posterior-dorsal 
submargins, and of which the 4th and 5th from the dorsal mar- 
gin are much smaller than the others; of the remaining ribs, the 
largest are found on the umbonal ridge, and become progressive- 
ly smaller anteriorly; lunule small but distinct; hinge of the 
right valve that of typical Cardita, with 3 cardinal teeth of which 
the middle one is much the largest, no laterals; ventral margin, 
internally fluted in harmony with the external ribs. 


387 Costa RicA MIocENE—OLSSON 215 


Length 18, height 12.50, diameter of the right valve 
5.50 mim. 


This small species represented by a single right valve in our 
collection appears to be a true Cavdzfa, its hinge being essential- 
ly that of the recent West Coast Cardzta laticostata Sowerby. 
With the exception of a few doubtful species in the Eocene of 
United States, true Cardita has heretofore not been found as a 
fossil in the Americas and its recent American species are con- 
fined to the Pacific coast. 


Compared with the recent Cardzta laticostata Sowerby, which 
is abundant along the Pacific Coast of Panama, the fossil shell 
differs in its smaller size, smaller umbos and in form. Its lunule 
is like that of /aticostata. 


Gatun Stage: Banana River. 


Section CARDITAMERA, Conrad 
Cardita matima, n. sp. Plate 32, figure 8 


Shell robust, solid, strongly inequilateral with the beaks 
situated at the anterior extremity of the shell; umbos 
large and full with strongly curved beaks above the small, 
sunken lunule; outline of the shell from within is subrectangu- 
lar with straight ventral and posterior sides which meet at 
nearly right angles; sculpture of about 18 strong, heavy cord- 
like ribs, separated by interspaces 1-3 of their width; the ribs 
are strongly nodulated on the disk and umbonal portion of the 
shell, but below they are simply irregularly wrinkled; a deep 
furrow extends from the beaks to the posterior extremity, just 
above the umbonal angle; the rib forming the summit of the 
ridge between the umbonal furrow and the hinge-line is larger 
than those on each side; shell cavity deep with a crudely den- 
ticulated ventral and posterior margin, hinge large (broken in 
our specimen), with a well-developed anterior lateral tooth in its 
left valve. 

Length 30, height 25, semi-diameter 12 mm. 


216 BULLETIN 39 288 


But a single specimen of this very distinet spceies was col- 
lected by Dr. A. C. Veatch from the Upper Gatun of Old Man 
Sam Creek in eastern Costa Rica. It differs from the other 
known American Carditameras by its convex shell and strongly 
curved beaks. The ribs are large and strongly nodulated at 
first, becoming irregularly wrinkled near the ventral margin. 
The hinge is unfortunately broken on our specimen, a left valve, 
but the small anterior lateral still remains which shows this shell 
to belong to the section Carditamera 

Gatun Stage: Old Man Sam Creek, 1 mile south from 

shore. 


Genus VENERICARDIA, Lamarck 
Venericardia scabricostata Guppy Plate 32, figures 14, 17 


Cardita scabricostata Guppy, 1866, Quart. Journ. Geol. Soc. London, 
Voll22h pi 2oB4 ple ce yon nos 

Cardita scabricostata Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 
252. 

Venericardia scabricostata Maury, 1917, Bull. Amer. Pal., vol. 5, p. 
ZBO2s plus Milo: 

This species originally described by Guppy from the Bowden 
beds of Jamaica is common in the Miocene of Costa Rica and 
frequently forms small zones in the Gatun beds. It occurs also 
in the Miocene of Santo Domingo. 

The ribs numbering about 18 are high, narrow and strongly 
granulated. The interspaces are wide and generally smooth. 

Gatun Stage: Zone E, G, Saury creek. 

Zone l, 3, East Grape Point creek. 
Coll. 5, Red Cliff creek. 
fleadwaters of Middle creek. 

Coll. C, Comadre creek. 


Venericardia Terryi, n. sp. Plate 32, figures 12, 13 


Shell of medium size, solid, moderately convex and sub-cir- 


G3 
e=2) 
Ns) 


Costa Rica M1ockNE—OLSSON Zi 


cular in form; beaks situated well forward and but slightly 
back of the extreme anterior end of the shell; the beak itself is 
small and loosely coiled over the small sunken lunule; sculpture 
consists of about 13, low, wide, smooth ribs which fit so closely 
together that there are no interspaces between them; the ribs are 
rounded in section on the umbos, wide and low ventally; the ribs 
are largest about the middle, small and fading away on the poster- 
ior and anterior dorsal submargins; interior concealed in a hard 
sandstone matrix. 
Length 22, height 21, semi-diameter 6 mm. 


The first specimens of this interesting species was collected 
by Mr. R. Terry from some Gatun sandstones in Quitana Creek, 
a small tributary streams of the Rio Betey. In that locality, it 
is an abundant species and occurs in small zones. 


It is related to the recent Pacific, V. crassicostata Sowerby 
but is a much smaller species. 
Gatun Stage: Outtana Creek. 
fiili No. 3, Banana River. 


Subgenus PLEUROMERIS, Conrad 
Venericardia Conradiana Gabb, var. limonensis, n. var. Plate 26, figs. 19, 20 


Cardita Conradiana Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 
2nd series, p. 377, pl. 47, fig. 79. 

Shell small, subcircular, moderately convex; umbos high 
and prominent, with small, inconspicuous beaks at the anterior 
1-3 of the shell; the sculpture consists of about 18 heavy, gran- 
ulated ribs, separated by somewhat narrower interspaces; interi- 
or of the shell deep, the hinge with 2 cardinal teeth and a small 
posterior and anterior lateral. 

Length 7.00, height 7.00, diameter of the left valve 
3.00 mm. 

Gabb specimen described from the Pliocene beds of the 
Limon Peninsula, measures 1% mm in length. His enlarged 
figure is very similiar to the present shell, in its subcircular fori 


218 BULLETIN 39 390 


and in number of ribs. The figure of the hinge shows the two 
cardinal teeth but no laterals. 


The common ¢ridentata Say of the Upper Chesapeake Mio- 
cene and recent fauna of the West Indies differs mainly in its 
more triangular form. 


Gatun Stage: Port Limon. 


Genus CHAMA, (Linné) Bruguiére 


Chama congregata Conrad Plate 28, figure 11 


Chama congregata Conrad, 1833, American Journal of Science, vol. 23, 
Pp. 341. 

Chama congregata Conrad, 1838, Fossils of the Medial Tertiary, p. 32, 
plen7 A tiga: 

Chama congregata Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, 
pt. 6, p. 1400. 

Chama congregatoides Maury, 1917, Bull. Amer. Pal., vol. 5, p. 200, 
pl. 33, fig. 8. 

Our specimens from Costa Rica are all small and belong to 
the upper or free valve, which in this species is the right. These 
specimens cannot be distinguished from true congregaza of the 
same size from the eastern United States, where it is an abund- 
ant fossil throughout most of the Chesapeake Miocene. The 
upper or right valve is finely, radiately frilled or fluted and these 
frills are cut by the concentric lamelle. The attached or left 
valve is more coarsely sculptured,both radially and concentrical- 
ly. 

Gatun Stage: Water Cay. 

Coll. Va Red i Glo Cxeek: 


Genus ECHINOCHAMA, Fischer 


Echinochama antiquata Dall Plate 28, figure 8 


Chama antiquata Guppy, 1866, Quart. Journ. Geol. Soc. London, vol. 
22, p. 294. Not of Linné. 

Echinochama antiquata Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 
3, pt. 6, p. 1404, pl. 54, fig. 9. 


391 Costa RicA MIocCENE—OLSSON 219 


Not “chinochana antiquata Maury, 1917, Buil. Amer. Pal., vol. 5, p. 
365, pl. 33, fig. 10. F. vaguensis Maury. 

This species described by Dall from the Bowden beds of 
Jamaica differs from the common, recent avcinella Linné of the 
West Indies, by its more numerous, irregular and lower ribs 
which bear only small, short spines. In arcenella the middle 
ribs are heavy and bear large, stump-like spines and the wide 
interspaces are reticulated with a coarse, mesh-like sculpture. 


Gatun Stage: Middle Creek, 
Coll. 6, Old Man Sam Creek. 
Flill No. 1, Banana River. 
Rio Blanco. 


Echinocama yaquensis Maury Plate 28, figure 5 


Chama arcinella Moore, 1853, Quart. Journ. Geol. Soc. London, vol. 
9, p. 130. Not of Linné. 

Chama arcinella Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 
251: 

Chama arcinella Guppy, 1874, Quart. Journ. Geol. Soc. London, vol. 
Behn Ou eee 

E-chinochama antiquata Maury, 1917, Bull. Amer. Pal., vol. 5, p. 365, 
pliiz3, fg. ro!) Not of Dall. 


Echinochama antiquata var. yaquensis Maury, 1917, op. cit. p. 365, pl. 
Bos SIT L2. 

E-chinochama trachyderma Pilsbry and Johnson, 1917, Proc. Acad. 
Nat. Sci. Phila., p. 197. 

The £Echinochama yaguensis Maury from the Miocene of 
Santo Domingo, where it is the prevaling species, differs from 
the antiguata by the reduced number of its ribs, which in some 
cases are nearly obsolete and in having its surface covered with 
a coarse, mesh-like sculpture of raised pustules. 


In Costa Rica, the azzzqguata is the common species, but we 
have also a single species of yagzenszs collected from the lower 
Gatun near Old Harbor. 


220 BULLETIN 39 392 


Gatun Stage: Old Harbor. 
(Gong. (Cs be 


Genus DIPLODONTA, Brown 


Section DIPLODONTA, s. s. 
Diplodonta collina, n. sp. Plate 32, figure 15 


Shell small, subcircular, convex; umbos wide, with small 
inconspicuous beaks, slightly posterior of the middle; anterior 
and posterior extremities well rounded into the base forming a 
part of the same circle; surface sculptured with fine, even, con- 
centric threads which are nearly lacking from the smooth umbos; 
hinge normal, and a plain ventral margin. 

Length 6.75, height 6.75, diameter of the right valve 
1.60 min. 

A small shell resembling the Dzplodonta nucletformis Wag- 
ner of the Upper Chesapeake Miocene and the recent West In- 
dian fauna, but less convex and more finely sculptured. 

Gatun Stage: Middle Creek. 


Section FELANIELLA, Dall 
Diplodonta insula, n. sp. Plate 32, figure 21 


Shell small, slightly convex, subcircular; beaks slightly 
posterior of the middle so that the posterior is a little longer 
than the anterior; beaks low and inconspicuous; both anterior 
and posterior extremities widely rounded, the posterior and the 
ventral margin forming a part of the same circle; surface with 
fine, concentric threads heaviest on the extremities, but nearly 
lacking from the umbonal area; interior of shell shallow, the 
right valve with a strongly bifid posterior cardinal; ventral mar- 
gin plain. 

Length 6.60, height 6.50, diameter of the right valve 
1.25 mm. 


393 Costa RicA M1ockNE—OLSSON 221 


A small, subcircular shell, neatly sculptured with fine, con- 
centric threads. DY. mznor Dall from the Bowden beds is strong- 
ly oblique. 

Gatun Stage: Water Cay. 


Genus MYRTAEA, Turton 


Myrtaea limoniana Dall Plate 32, figures II, 23 


Myrtea limoniana Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, 
pt. 6, p. 1358, pl. 52, fig. ro. 

A small, thin and slightly convex shell, its surface sculptur- 
ed with fine, thin and sharp, elevated, concentric lamelle. We 
have several specimens from the coral limestones of Port Limon 
as well asalarger specimen from Zone 4, of Red Cliff Creek. 
This later shell measures: heigth 9.50, length 10.00, diameter of 
the right valve 2.50 mm. It occurs also in the Bowden beds of 
Jamaica. 

Gatun Stage; Port Limon. 

Coll. 4, Red Cliff creek. 


Genus PHACOIDES, Blainville 


Subgenus CALLUCINA, Dall 
Phacoides radians Conrad, var. medioamericanus, n. var. Plate 32 fig. 16 


The Phacotdes radians Conrad is a fossil of the Upper Ches- 
apeake Miocene of the Carolinas but has continued through the 
Pliocene to the recent fauna. The Costa Rican fossils differ from 
typical radians in having their umbos sculptured with about 
6 ribs much stronger than the concentric, while in typical 
radians, the ribs are more numerous and about equal to the con- 
centric lines. The medioamericanus is also smaller as may be 
seen in the measurements given below, while vadzans is large 
(20 mm) and often quite convex. 


Length 9.5, height 9, semi-diametes 2.5 mm. 


222 BULLETIN 39 394 


Gatun Stage: Hleadwater of Middle creek. 
Old Man Sam creek, 1 mile south of the beach. 


Subgenus HERE, Gabb 


Section CAVILUCINA, Fischer 


Phacoides trisuleatus Conrad 


Lucina trisulcata Conrad, 1841, Trans. Amer. Assoc. Nat. and Geol., 
vol. I. p. IIo. 


Lucina trisulcata Conrad, 1845, Fossils of the Medial Tertairy, p. 71, 
pl. 40, fig. 5. ? 

Phacoides (Cavilucina) trisulcatus Dall, 1903, Trans. Wagner Free 
Inst. Sci., vol. 3, pt. 6, p. 1369. 

Few specimens from the Gatun of Middle Creek. They 
are identical with typical specimens from the Chesapeake Mio- 
cene of the eastern United States. The species also occurs in the 
recent fauna. 


Gatun Stage: Middle creek. 
Old Man Sam creek. 


Subgenus LUCINISCA, Dall 


Phacoides costaricensis, n. sp. Plate 32, figure 5 


Shell subcircular, moderately thick and but slightly convex: 
beaks small and inconspicuous, a little in front of the middle; 
dorsal area impressed and well defined; sculpture consists of 
strong, regular, radial threads, widely separated by interspaces 
2 or 3 times the width of the threads themselves; they are cross- 
ed by distant but regular concentric ridges, the intersection form- 
ing small beads or granulations, strongest on the anterior and 
posterior extremities; the lunule is narrow, lanceolate and with 
the concentric sculpture predominanting; interior of shell cavity 
rather shallow, the ventral margin coarsely serrated by the ex- 
ternal sculpture. 


295 Costa Rica MrocENE—OLSSON 22 


WwW 


Length 11.50, height 11.00, diameter of the left valve 
2.25 mm. 

This species is recognized by its depressed or but slightly 
convex shell and coarse sculpture, of which the radial is a little 
stronger. Phacotdes cribrarius Say and nassula Conrad are both 
more convex, with the concentric lamellee more distantly spaced 
on the umbos and dominating over the radial. PP. murtcatus 
Spengler of the West Indies is much more finely sculptured. 

We have also a young shell from Water Cay which is pos- 
sibly P. zassula, var. caloosana Dall of the Florida Pliocene. It 
is very close to specimens of that species in the Cornell collec- 
tion. 

Gatun Stage: Middle creek. 


Phacoides hocasensis, n. sp. Plate 32, figure 6 


Shell thin, subcircular, depressed; dorsal area well defined 
by a marked change in sculpture; beak small, but distinct, placed 
a little in front of the middle; the sculpture on the disk of the 
shell is fine, composed of subequal, radial threads, separated by 
narrow interspaces; the ribs and interspaces are crossed by fine, 
concentric threads or ridges which produce a finely scabrous 
surface; the concentric threads are a little more widely spaced 
on the umbos; the dorsal area is defined by a stronger radial, be- 
yond which follows a wide radial band, strongly sculptured with 
the concentric threads or lamelle but lacking in radials; the up- 
per half of the dorsal band bears in addition to the concentric 
lamelle, 2 or 3 weak but scabrous radial threads; the lunule is 
very narrow, lanceolate and with a few strong, concentric lame- 
lle; interior of the shell cavity shallow, with the ventral margin 
finely crenulated. 

Length 10.00, height 9.25, diameter of the left valve 
2.00 mim. 

A depressed and very finely sculptured species whose gener- 
al form is that of P. muricatus Spengler of the recent West In- 
dian fauna. It is more regularly sculptured than the P. hAzspar- 


224 BULLETIN 39 396 


zolana Maury of the Cercado formation of Santo Domingo but 
the main difference lies in their dorsal areas, that of hzsbantolana 
Maury being sculptured more or less like the rest of the shell; 
in bocasensts, as described above, the sculpture of the dorsal area 
is dominantly concentric, only the upper half bearing 2 or 3, low 
but scabrous radials. 


Gatun Stage: Bocas del Toro. 
Subgenus PARVILUCINA, Dall 


Section BELLUCINA, Dall 


Phacoides actinus Dall Plate 32, figure 24 


Phacoides ( Bellucina) actinus Dall, 1903, Trans. Wagner Free Inst. 
Sci., vol. 3, pt. 6, p. 1385, pl. 52, fig. 3. 


Dall gives for his typical specimens from the Bowden beds 
of Jamaica, a height of 4.50 mm, while the Panama shells, all 
from Bocas Island, do not exceed 3 mm in height but agree in 
other respects. It is also recorded by Maury from the upper 
part of the Cercado formation of Santo Domingo. 


Gatun Stage: Bocas del Toro. 
Genus DIVARICELLA, von Martens 


Divaricella quadrisulcata d’Orbigny Plate 32, figure 20 


Lucina divaricata Conrad, 1840, Fossils of the Medical Tertiary, p. 39, 
pl. 20, fig. 3, but not of Linné. 

Lucina quadrisulcata d’Orbigny, 1846, Voy. Am. Mer., p. 584. 

Divaricella quadrisulcata Dall, 1903, Trans. Wagner Free Inst. Sci., 

vol. 3, pt. 6, p. 1389. 
Lucina ( Divaricella) quadrisulcata Bose, 1906, Boll. de Inst. Geol. de 
Mexico, numero 22, p. 77, pl. II, figs. 2, 3. 

This species is very rare in Costa Rica. The shells are rath- 
er thin and generally more finely sculptured and approach in 
this respect the D. compsa Dall of the Florida Pliocene. Our 
larges specimen measures: height 14 m, length 1o mm, diameter 


397 Costa R1icA MrocENE—OLSSON 225 


of left valve 4.50 mm. The D. prevaricata Guppy of the Mio- 
cene of Santo Domingo and Jamaica measures only 8 mm in 
height. 

The D. guadrisulcata is a recent species of the West Indies 
appearing in the east coast Miocene of the United States for the 
first time in the Upper Chesapeake. 

Gatun Stage:Coll. 4,5, Red Cliff creek. 

Middle creek. 


Genus CARDIUM, Linné 
Subgenus TRACHYCARDIUM, Moerch 


Cardium stiriatum Brown and Pilsbry Plate 27, figures 4, 5 


Cardium (Trachycardinm) stiriatum Brown and Pilsbry, 1911, Proc. 
Acad. Nat. Sci. Phila., vol. 63, p. 366, pl. 28, fig. 11. 


This is most elegant of the several species of Cardium in 
the Miocene of Panama and Costa Rica. The shell is high and 
inflated with about 30 radial ribs which are peculiarly sculptur- 
ed with twisted or obliquely projecting tuberclers placed on the 
posterior side of the ribs and overhanging the adjacent inter- 
space. Large specimens may reach a height 50 mm. It is re- 
lated to the recent West Coast Cardium belcheri Broderip. 


Gatun Stage: Gatun, C. Z. 
Water Cay, Panama. 
Hill No. 1, Banaxe River. 


Cardium dominicense Gabb Plate 27, figure 2 


Cardium )Trachycardium) dominicense Gabb, 1873, Trans. Amer. 
Phil. Soc., vol. 15, p. 25. 

Cardium ( Trachycardium) dominicense Gabb, 1881, Journ. Acad. Nat. 
Sci. Phila., 2nd series, vol. 8, p. 344. 

Cardium (Trachycardium) gatunensts Toula, 1908, Jahrb. der K-K 
Geol. Reichsanstalt Wien, vol. 58, p. 720, pl. 27, fig. 4. Not Cara- 
tum (Fragum) gatunensis Dall, 1900. 


226 BULLETIN 39 308 


Cardiuin ( Trachycardium ) dominicense Brown and Pilsbry, 1911, Proc. 
Acad. Nat. Sci. Phila., vol. 63, p. 367. 

A large species abundant in the Canal Zone but usually only 
as internal casts. It appears to be rare in Costa Rica, our col- 
ection containing only a few fragments of a large individual 
from Hast Grape Point Creek. It is distinguished not only by 
its large size, but very numerous (60+) fine ribs. 

Gatun Stage: Gatun, C. Z. 

East Grape Point Creek, C. R. 
APOE, 3 Gara G-aoo)) 


Cardium costaricanum, n. sp. Plate 27. neurens 


Shell ovate, solid, inflated; umbos wide and full with high 
beaks; sculpture of about 24 strong ribs, rounded or subtrian- 
gular in section with their apex leaning posteriorly except those 
on the posterior slope which lean anteriorly; interspaces narraw, 
formed by the sides of the ribs; the ribs are mostly smooth ex- 
cept those on the anterior half of the shell which have their pos- 
terior side more or less granulated; this granulation is found al- 
so on a few of the central ribs near their ventral margin; hinge 
narrow; internal margin fluted in harmony with the external 
ribs. 

Length 24.50, height 34, semi-diameter 15 mm. 

This species is not uncommon in Costa Rica and may easily 
be confused with the C. dominicanum Dall. It differs in its 
larger side, narrow umbos, more pointed beaks and more flatten- 
ed ribs. 

Gatun Stage: Coll. 4, East Grape Point Creek. 


Subgenus FRAGUM, Bolton 


Cardium medium Linnzeus Plate 27, figure 6 


Cardinm medium T,innzeus, 1758, Sys. Nat. ed. 10, p. 678. 

Cardium medium Reeve, 1844, Conch. Icon., Cardium, pl. 6. fig. 
30. 

Cardium (Fragum) medium Dall, 1900, Trans. Wagner Free Inst. 
SCiivolwa ap tsii pw lLOLe 


3909 CosTa R1icA MiocENE—OLSSON 227 


A recent West Indian species but also common as an Up- 
per Miocene and Pliocene fossil along the Atlantic coastal plain 
of eastern United States. The ribs number 24 to 27 in front of 
the umbonal angle. Dall has described a Cardium (Fragum) 
gatunensis from Gatun, on which the ribs number only 16 in 
front of the truncation and 10 behind. 


Gatun Stage: Port Limon. 


Section TRIGONIOCARDIA Dall 
Cardium heredium, n. sp. Plate 27, figure to 


Shell small, solid, oblique, convex and subquadrate; strong- 
ly carinate on the posterior-umbonal slope and truncated; umbos 
high and prominent; shell sculptured anterior to the truncation 
with 12 or 13 strong ribs, separated by interspaces about % to 
Y% as wide; there are 7 or 8 smaller ribs on the posterior trun- 
cation; the interspaces are finely sculptured with small, elevated 
cross-threads; the anterior set of ribs on the type specimens are 
wide, subequal and smooth; the posterior are unequal, with the 
three, bordering the umboal angulation much the smaller and 
bearing small and scattered pustules; on the posterior ribs where 
the pustules are lacking, are small dot-like depressions. 


Height 11.50, length 11.50, diameter 6.00 mm. 


This species approaches the Cardium apateticum Dall from 
the Oak Grove sands of Lower Miocene age in Florida. It dif- 
fers by its more oblique umbos and proportionately higher shell. 
The pustules are variously developed and on some shells are 
more strongly developed and may also appear on some of the 
ribs anterior to the truncation. These more strongly scalptured 
shells may prove with more material to belong to another 
species. 

Gatun Stage: Rio Betey. 

Banana River. 
Fistrella River. 


228 BULLETIN 39 460 


Subgenus PAPYRIDEA, Swainson 


Cardium spinosum Meuschen, var. Turtoni Dall Plate 27, figure I 


Cardium (Papyridea) spinosum var. Turtoni Dall, tg00, Trans. Wag- 
mer Free Inst., Sci., vol. 3, pt. 5; p. 1108. 

The fossil shells from Costa Rica, are referrible to this var- 
iety of spimosum, distinguished by having the interspaces be- 
tween the ribs marked with a flat thread, between sharp grooves. 
This variety is also found fossil in the Chesapeake Miocene at 
Jackson Bluff Florida and in the Pliocene Caloosahatchie beds of 
the same state. It the recent fauna it is confined, according to 
Dall, to the eastern Atlantic, with the typical spzzosum in the 
West Indies and a third variety aspersum Sowerby along the 
Pacific side. 


Gatun Stage: Port Limon. 


Subgenus LAEVICARDIUM, Swainson 
Cardium serratum Linnzeus Plate 27, figures II, 12 


Cardium serratum Linneeus, 1758, Syst. Nat., ed. 10, p. 680. 


Cardium (Levicardium) serratum Dall, tg00, Trans. Wagner Free 
Insts Sciesivolai2s ptrs4) p.ULtos 


Cardium (Levicardium) serratum Brown and Pilsbry, 1911, Proc 
Acad. Nat. Sci. Phila., vol. 63, p. 367. 


Cardium (Levicardium) serratum Maury, 1917, Bull. Amer. Pal., vol. 
iy 1 27s) OLA) Ios) a8lreZ4 Xe) 

Most of the fossil shells are indistinguishable from recent 
examples of servatum from the West Indies. Others shells are 
decidely oblique and approach in form the C. sublineatum Con- 
rad of the Upper Chesapeake Miocene of eastern United States. 
The C. venustum Gabb from the Miocene of Santo Domingo is 
very distinct from serratum and characterized by its faintly flex- 
uous outline, strong radial striz, andin having the lower cardin- 
al tooth large and strong. 


401 ' Costa Rica MriockENE—OLSSON 229 


Gatun Stage: Gatun, C. Z. 
Muddley Greek Gj 0h. 
BO WA, (Go I 


Genus PROTOCARDIA, Beyrich 


Protocardia costaricensis, n. sp. Plate 27, figure 13 


Shell small, rather thin, convex; beaks a little anterior to 
the middle, with high, inflated umbos; the anterior end is wide- 
ly rounded, the posterior somewhat truncated; the posterior area 
is well differentiated and occupies the posterior one-third of the 
shell and is sculptured with numerous small radial threads sep- 
arated by wider interspaces; these interspaces are finely sculp- 
tured with cross-threads; anterior of the posterior area the shell 
surface is sculptured with small radial threads which are finely 
beaded; on the anterior end of the shell, the beading is more ir- 
regular and is seen to be made by the concentric sculpture. 


Height 18, length 21, diameter of the left valve 7 mm. 


This species differs from both P. jamazcensis Dall from the 
Bowden beds of Jamaica and P. zslahispaniole Maury from the 
Miocene of Santo Domingo by its smaller posterior area. It ap- 
proaches more closely the Dominican shell but is shorter and 
more coarsely sculptured. 

The P. Newberryana Gabb from Gatun is based on a large 
internal cast which is entirely unrecognizable. 


Gatun Stage: Comadre creek. 


Subgenus LOPHOCARDIUM, Fischer 


Protocardia gurabica Maury Plate 27, figures 7-9 


Protocardia gurabica Maury, 1917, Bull. Amer, Pal., vol. 5, p. 377, pl. 

36, fig. Io, 
Shell thin, inflated, a little longer than high and with the 
greatest convexity about the center of the shell; the umbos are 
high, prominent and wide, and project well above the hinge 


230 BULLETIN 39 A402 


margin; the anterior end is well rounded into the base; the pos- 
terior end is more contracted and gaping slightly at its extremi- 
ty; surface sculpture on the body of the shell, anteriorly con- 
sists of narrow, concentric wrinkles which are best developed on 
the anterior end but become obsolete on the middle of the shell 
and posteriorly; in addition the whole surface carries fine, 
sub-obsolete radial threads; on the posterior area, the sculpture 
is of heavier and coarser radial threads, lacking the concentric, 
except t he irregular growth lines. 


Height 56, length 32, diameter 20 mm. 


The subgenus Lophocardium, of which there are two recent 
species along the West Coast of Mexico, differs from the true 
Protocardia, in its gaping posterior end and lack of lateral teeth 
as well-as in important difference in its anatomical structure. 
The shells are strongly convex, with high, inflated umbos and 
are extremily thin and fragile. 


The P. gurabica was described by Dr. Maury from the Gur- 
abo formation of Santo Domingo, and compared by her with the 
Vickburgian P. dzversa Conrad, but it appears to me to be a typ- 
ical Lophocardium and representing the first member of this in- 
teresting subgenus to be found in the fossil state. As her speci- 
men is very fragmentary, I have redescribed the species from 
better preserved Panama and Costa Rican material. The poster- 
ior end is gaping and the internal mold shows no impression of 
lateral teeth along the hinge margin. It approaches closely the 
L. Annette Yall, dredged from 8 to 27 fathoms in the Gulf of 
California, except that it is proportionately longer and lacks the 
concentric wrinkles on its posterior area. The L. Cumingz Brod- 
erip, the type of the subgenus, carries between its posterior area 
and the general surface of the shell, a thin radial lamina or crest, 
extending from the umbos to the ventral margin. 


The fossil shell is extremily thin. It generally occurs simp- 
ly as internal molds which may still retains a part of the shell 
or impression of its sculpture on its surface. 


403 Costa Rica M1ocENE—OLSSON 231 
Gatun Stage: Gatun, and Mt. Hope, C. Z. 
Water Cay. 
Margarita Trail, C. R. 
Genus DOSINIA, Scopoli 
Section DOSINIDEA, Dall 
Dosinia acetabulum Conrad Plate 31, figure I 


Artemis acetabulum Conrad, 1833, Fossils of the Tertiary Formations, 
PZOvepl won fie. Te 

Dosinia acetabulum Bose, 1906, Boll. de Inst. Geol. de Mexico, num- 
ELO}2 2 py Oly iplo Tl, Ss. 17), Wl 

Dosinia (Artemis) cf. acetabulum Toula, 1908, Jahrb. der K-K, Geol. 
Reichsanstalt, Wien, vol. 58, p. 727, pl. 27, figs. 8, 8a. 

The Dosinia acetabulum has been recorded by Bose from 
Mexico and by Toula from Panama, but their determinations 
were based on imperfect material or casts. The Costa Rican 
collection however contains a large series of finely preserved Do- 
sinias which are unquestionably the Dosinza acetabulum Conrad 
of the Chesapeake Miocene. 

The surface of the Costa Rican specimens are generally 
neatly sculptured with even, concentric bands or ribbons which 
on the middle of the shell disk average 2 mm in width. The 
form of the shell is subcircular and like the typical D. acetabu- 
Jum of the Yorktown formation of Virginia and North Carolina. 
The Costa Rican shells are equally as large as the Chesapeake 
shell measuring in height 70 mm or more. 

The Dosinia acetabulum in the eastern United States, is a 
most characteristic fossil of the Chesapeake Miocene, where its 
range extends throughout most of the lower and upper Chesa- 
peake formations. It is lacking from the Lower Miocene or the 
Alum Bluff formation of Florida, as well as from the succeeding 
Pliocene. Inits range through the Chesakeake Miocene, it 
gives rise to several well-marked varieties of which the broad 


222; BULLETIN 39 404 


subcircular shell of the Upper Chesapeake is the typical form. 
The Costa Rican shells are most like the Upper Chesapeake 
typical acetabulum. 

Brown and Pilsbry have described a smaller and more finely 
sculptured species from the Gatun Spillway as delicatisstma. 
This spceies is represented in our collection from the same place 
and appears to be distinct. Toula’s specimen from Gatun, how- 
ever, is a true acetabulum. 

The Dosinia acetabulum is a common fossil in the Gatun 
beds of Costa Rica, where it occurs most abundautly in the 
lower part of the formation. 

Gatun Stage: Gatun, C. Z. 

Flotel and Hone Creek. 
Pumbri Creek. 

fiill No. 37, Banana River. 
Coll. 6, Estrella River. 


Genus CLEMENTIA, Gray 
Clementia dariena Conrad Plate 31, figure 4 


Meretrix dariena Conrad, 1856, Pacific R. R. Reports, vol. 5, p. 328, 


pl. 6, fig. 55. 

Clementia dariena Gabb, 1881, Journ. Nat. Sci. Phila., vol. 8, p, 344, 
DLV AAN fet 6s 

Clementia dariena Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, 
DEVON AI225. 


Clementia daviena Toula, 1908, Jahrb. der K-K. Geol. Reichsanstalt, 
vol. 58, p. 7:25, ple 27,, fges.o,) 10: 

Clementia dariena Hubbard, 1920, Scientific Survey of P. R. and the 
Virgin Islands, vol. 3, pt. 2, p. 118, pl. 19, figs 10-12. 

This species is very common in the lower Gatun of the 
Canal Zone. It is very rare in western Panama and in Costa 
Rica, where it occurs in the lower part of the Gatun formation 
and in the Uscari shales. Romanes* has recorded it from Bar- 


* Quart. Journ. Geol. Soc. London, 1912, vol. 68, p. 125. 


405 Costa Rica M1rocENE—OLSSON 233 


ranca near Punta Arenas onthe Pacific coast of Costa Rica, 
from beds probably equivalent to the Lower Gatun or to the 
Uscari. 

The shell is very thin and fragile, and hence the specimens 
are very frequently much crushed and distorted in shape. Typi- 
cally the shell is broad and high, the beaks anterior in position 
and the surface sculptured with narrow, wave-like undulations 
which are best developed on the umbos. 

Length 70, height 60. diameter 34 mm (Sousi Creek). 

Uscari Stage: Sapote, C. Z. (Gabb) 

Gatun Stage: Gatun, C. Z. 

Nancy's Cay, Prov. of Bocas del Toro. 
Upper Hone Creek, and tts tributary Soust 
Creek. 


Genus GAFRARIUM, Bolton 


Section GOULDIA, C..B. Adams 
Gafrarium limonensis, n. sp. Plate 32, figure 18 


Shell small, subcircular, moderately convex, umbos inflated, 
beaks anterior to the middle: surface finely sculptured with even, 
concentric riblets, more or less cancellated by fine, radial threads 
which are strongest on the anterior and posterior extremities, 
while the concentric riblets predominate on the middle of the 
shell; lunule distinct, narrowly lanceolate; interior of the shell 
deep, the hinge of both valves with 3 cardinal teeth and an an- 
terior lateral; ventral margin concentrically grooved but not 
tangentially asin 7vansenella. 

Wengthia.75, height 4:25, diameter) ot) the lett: valve 
1.50 mm. 

This small Gouldia, is fairly abundant in the Miocene coral 
limestones of Port Limon, and is characterized by its small, con- — 
vex shell and subcircular form. Gabb has described a small 
Gouldia from the Pliocene beds of the Limon Peninsula, but that 


234 BULLETIN 39 406 


species is more depressed, less circular and more trigonal in out- 
line. Gabb’s species may prove to be but a small form of the 
recent West Indien G. cerzza C. B. Adams, 
Gatun Stage; Port Limon. 
Old Man Sam creek, 1 mile south of the beach. 


Gafrarinm altum Dall, va. costaricensis, n. sp. Plate 32, figures I9, 22 


ef. Gafrartum altum Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, 
pt. 6, p. 1249, pl. 57, fig. 5, 

Shell small, high, rounded trigonal, moderately convex; 
beaks small and pointed slightly forward, in young shell nearly 
central but becoming with maturity slightly anterior; anterior 
and posterior extremities nearly similiar and evenly rounded; 
surface with fine, concentric lines, most distinct and even to- 
wards the ventral margin, and with faint, radial strize on the 
anterior and posterior slopes; lunule large, lanceolate, defined by 
an impressed line; interior of shell deep, with a concentrically 
grooved margin. 


Length 4.85, height 4.75, diameter of right valve 1.50 
min. 

The Costa Rican shells differ from the typical altum, de- 
scribed by Dall from the Oak Grove sands of Florida, in being 
more trigonal in form and with higher beaks. The sculpture of 
its surface is mainly concentric, the radials showing only as 
very faint striae on the posterior and anterior extremities. 

Gatun Stage: Middle creek. 

Coll. 6, Estrella River. 


Genus MACROCALLISTA, Meek 


Section CHIONELLA, Cossmann 
Macrocallista maculata Linnzeus Plate 31, figures 6, 7 
Venus maculata Vinnzeus, 1758, Syst. Nat., ed. 10, p. 686, ed. 12, p. 
rri2% 
Cytherea dariena Conrad, 1857, Pacific R. R. Report, vol. 6, p. 72, pl. 
Ip, 1 0egg yA 


407 Costa R1icA MIocKNE—OLSSON 235 


Callista maculata Gabb, 1881, Journ, Acad. Nat. Sci. Phila:, 2nd series, 
VOLS) p41 372: 


Macrocallista (Chionella) maculata Dall, 1903, Trans. Wagner Free 
IS tNSCi..) voles. pt. (Gn pos) 256: 


Macrocallista maculata Brown and Pilsbry, 1911, Proc, Acad. Nat. Sci. 
Plila., vol. 63, p. 370. 

The Macrocallista maculata is a common and widely dis- 
tributed fossil in the Costa Rican Miocene and is readily recog- 
nized by its depressed, porcellaneous and highly polished shell. 
In the usual sandy phases of the Gatun, the species is quite typi- 
eal, but becomes high and broad in the coralline or transitional 
phase. 


A variety cuneata has been proposed by Gabb to include 
shells in which this posterior-dorsal slope is straight and not 
humped as usual. We have seen no examples of this type. 


The Macrocallista maculata isa common recent species of 
the West Indies, appearing for the first time, in the lower Mio- 
cene beds of the Chipola River, Florida. 

Gatun Stage: Headwaters of Middle creek. 

Rio Blanco. 


Genus CALLOCARDIA, A. Adams 


Subgenus AGRIOPOMA, Dall 
Callocardia gatunensis Dall Plate 32, figure 1 


Callocardia (Agriopoma) gatunensis Dall, 1903, Trans. Wagner Free 
Inst. Sci., vol. 3, pt. 6, p. 1260, pl. 54, fig.1. 


Callocardia (Agriopoma) gatunensis Brown and Pilsbry, 1911, Proc. 
Acad. Nat. Sci. Phila., vol. 63, p. 370. 

A convex, cordate shell very abundant in the Gatun beds 
of the Canal Zone, especially in the quarries at Mt. Hope. It 
is a rare species in Costa Rica, but generally larger, more trian- 
gular in outline and somewhat coarsely sculptured. These shells 
are nearest to Dall’s variety mudltzfilosa. 


236 BULLETIN 39 408 


Dall also records the species from the Miocene of Santo Do- 
mingo and Jamaica. 


Gabb’s C. sapotensts, from Sapote, Costa Rica, we have not 
seen. It isa higher and more Dosinoid shell with nearly cen- 
tral beaks. It is possibly but a variety of this species. 


The following measurements are of Costa Rican examples: 


Length 46, height 28, thickness 35 mm. 
533 44, 16 mm (right valve) 


Gatun Stage: Gatun and Mt. Hope, C. Z. 
fiill No. 1, Banana River. 
Soust Creek (Upper Fone creek). 


Genus PITARIA, Roemer 


Section LAMELLICONCHA, Dall 


Pitaria circinata Born Plate 31 figures 3, 9 


Venus civcinata Born, 1780, Test. Mus. Vind. p. 61, pl. 4, fig. 8. 

Cytherea alternata Broderip, 1835, Proc. Zool. Soc., p. 45. 

Cytherea Juncea Guppy, 1866, Quart. Journ. Geol. Soc. London, vol. 
22, p. 682, pl. 26. fig. 13. 

Chione circinata Gabb, 1873, Tarns. Amer. Phil. Soc., vol. 15, p. 250. 

Cytherea juncea Guppy, 1876, Quart. Journ. Geol. Soc. London, vol, 
32, p. 531. 

Pitaria (Lamelliconcha) circinata Dall, 1903, Trans, Wagner Free 
Inst. Sci., vol. 3, pt. 6, p. 1269. 


Pitaria circinata Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila. 
vol. 63, p. 370. 

Pitarvia (Lamelliconcha) ciycinata Maury, 1917, Bull. Amer. Pal., vol. 
5, P- 379, Pl. 37, fig. I. 

The Pitaria circinata Bornis a common, recent Veneroid 
along the Caribbean coast of Panama and Costa Rica. It occurs 
also on the west or Pacific side but ina slightly different form, 
as the variety a/ternata Broderip. This variety differs from the 
typical civcinata by its larger, more convex shell and more dis- 
tant concentric ribs. 


409 Costa RicA MIocENE—OLSSON 237 


The fossil shells from Costa Rica are generally more like the 
West Coast alternata, but are smaller and with somewhat more 
pointed beaks. The species is also found as fossil, in Santo Do- 
mingo, Trinidad and Venezuela. 

Gatun Stage: Gatun, C. Z. 

Lowe EN Saray Green Canine 


Pitaria Hillii Dall, var. musanica, n. var, Plate 31, figure 5 


Shell elongate, moderately convex and rather solid; beaks 
situated at the anterior %, with the anterior extremity widely 
rounded but obtusely pointed posteriorly; lunule small, ridged 
in the middle and defined by a faint line; posterior-dorsal area 
broad and flat; surface of the shell sculptured with low, irregu- 
lar, rounded, concentric ribs, which are persistent along the 
posterior-dorsal slope but lacking elsewhere near,the ventral mar- 
gin; interior of shell unknown. 

Length 42, height 27, thickness (left valve) 10.50 mm. 

This species has a solid shell like that of Wacrocallista. It 
differs from the true /7z//7zz described by Dall from the Gatun of 
the Canal Zone, by its more rude and irregular concentric sculp- 
Blane: 


The P. planivieta Guppy, from the Miocene of Jamaica and 
Santo Domingo is a much shorter and more regularly sculptur- 
ed shell. 


Gatun Stage: Hill No. 2, Banana River, C. R. 


Pitaria Guppyana Gabb Plate 31, figure I1 


Caryatis Guppyana Gabb, 1881, Journ. Acad. Nat. Sci. Phila., 2nd 
series, vol. 8, p. 373, pl. 47, fig. 73. 

Shell small, ovate, moderately convex and _ porcellaneous; 
dorsal side straight; ventral side gently convex; posterior ex- 
tremity sub-truncate but wide, meeting the dorsal side nearly at 
right angles; anterior extremity much narrower, produced and 
rounded; lunule narrowly lanceolate, sculptured with the con- 


238 BULLETIN 39 410 


tinuation of the concentric ribs and separated from the shell disk 
by an impressed line; surface of disk sculptured with rather wide, 
low or appressed ribs formed between incised lines, but the 
shell as a whole is porcellaneous and smooth; as the ribs pass 
over the dorsal-posterior area towards the dorsal margin they 
decrease in number through fusion; interior concealed in a hard 
matrix. 


Length 32, height 18, diameter (right valve) 5 mm. 


A small species of the coralline limestones and marls of the 
Limon Peninsula, from which it was described by Gabb. It 
approaches the P. planivieta Guppy of the Miocene of Jamaica and 
Santo Domingo but is smaller and more rectangular in outlines. 
It is rather convex with a smooth and polished surface and with 
regular, even, concentric ribs. 


Gatun Stage: Port Limon. 


Pitaria VanWinklez, n. sp. Plate 32, figure 2, 3 


Shell elongate, depressed; anterior extremity broadly round- 
ed, posterior more acute; lunule small, lanceolate and defined by 
an impressed line; escutcheon long and very narrow; surface 
sculptured with narrow, closely spaced and reflected ribs; these 
ribs which number about 100 on the shell disk are even on the 
center of the shell, but generally uneven on the anterior extrem- 
ity; the ribs are continuous from the edge of the escutcheon to 
the anterior dorsal margin; the rounded posterior dorsal slope 
carries two small, faint rows along which the concentric ribs are 
slightly higher and occasionally raised into small incipient 
spines. 

Length 41.5, height 32, diameter (left valve) 10 mm. 


This species is quite similiar to the P. vosea Broderip and 
Sowerby of the West Coast in its depressed shell and general 
form. ‘The concentric ribs are very regular, even and continu- 
ous across the entire surface of the shell. Where these ribs cross 
the posterior-dorsal slope, they occasionally become raised into 


4AXY Costa RicA M1ocENE—OLSSON 239 


two rows of incipient spines. In P. rosea there is only one row 
of incipient spines and the dorsal-posterior area is smooth. 

This species is named for Miss Katherine Van Winkle of the 
Paleontological Department of Cornell University and who is at 
present engaged on a Monograph of the American species of 
Veneride. 

Gatun Stage: Hill No. 3. Banana River, C. R. 


Pitaria boucaryensis, n. sp. Plate 31, figure 10 


Shell rounded quadrate, thin and moderately convex; pos- 
terior and anterior extremities rounded, umbos rather large; 
lunule small and feebly defined by a faint, impressed line; sur- 
face of disk sculptured with about 4o even, distant, concentric 
lamellae; interspaces generally sinooth or irregularly sculptured 
with growth lines; interior of shell concealed. 


Length 31, height 22, diameter (right valve) 10 mm. 


As the single type specimen is fragmentary and imbedded 
in a hard sandstone, the above measurements are but approxi- 
mate. It isa very distinctive species, differing from the other 
fossil Costa Rican Pitarias by its more delicate and convex shell 
and more distant, even, concentric lamellee. 


Gatun Stage: Boucary creek, C. R. 
Genus ANTIGONA, Schumacher 


Antigona multicostata Sowerby Plate 30, figure I 


Venus multicostata Sowerby, 1835, Proc. Zool. Soc., p. 22. 

Venus multicostata Reeve, 1863, Conch. Icon., vol. 11, pl. 3, fig. 9. 

Cytherea ( Cytherea) mutlticostata Dall, 1902, Proc. U.S. Nat. Mus., 
vol. 26, p. 390. 

This is a large species living on the south or Pacific side of 
the Isthmus. A perfect specimen was collected from the Gatun 
beds of Water Cay, identical in every respect with recent ex- 
amples from the Bay of Panama. It is another instance among 
the many which we have had occasion to note in the course of 


240 BULLETIN 39 412 


the present study of the occurrence of West Coast species in the 
Caribbean Miocene, proving conclusively from a purely paleon- 
tological standpoint the rather late separation of the Atlantic 
and Pacific through the final uplift of the isthmian lands. 


The recent A. Lister? Gray is a smaller, more elongate 
species and with finer and sharper concentric lamelle. 


Length 107 mm, height 102 mm, diameter 69 mm. 
Gatun Stage: Water Cay. 


Antigona tarquina Dall Plate 30, figure Io 


Venus magnifica Gabb, 1673, Trans. Amer, Phil. Soc., vol. 15, p. 249. 
Not of Sowerby, 1853. 


Cytherea tarquina Dall, 1900, Trans. Wagner Free Inst., vol. 3, pt. 5, 
pl. 38, figs. 2, 2a; vol. 3, pt. 6, p. 1274. 

Antigona tarquina Dall, 1915, Bull. go, U. S. Nat. Mus., p. 147. 

Antigona tarquina Maury, 1917, Bull. Amer. Pal., vol. 5, p. 381. pl. 
37, fig. 4. 

A small shell identical or closely related to A. tarquina 
Dall was collected from Zone 3 of Saury Creek. It is similiar 
to Dr. Maury’s specimen of 4. farguina from Santo Domingo. 

Typical A. targuina Dall is a common fossil of the Orthau- 
lax pugnax fauna of the Tampa Silex beds of Florida. It was 
also collected by Professor Gabb and Dr. Maury from the Mio- 
cene of Santo Domingo. 


Gatun Stage: Zone E, Saury creek, near Cahutta, C. R. 


Section VENTRICOLA, Roemer 


Antigona rugosa Gmeiln Plate 30, figure 4 


Venus rugosa Gmelin, 1792, Syst. Nat., p. 3276. 
Venus rugosa Reeve, 1863, Conch. Icon., vol. 11, pl. 7, fig. 23. 


Several specimens from the coralline limestones of Port 
Limon. ‘They are practically identical with recent examples of 
A. rugosa from the West Indies. This species is distributed 
throughout the West Indies but is also found living in the Gulf 


413 Costa Rica MrocENE—OLSSON 241 


of California. This distribution on both sides of the Isthmus is 
in harmony with its occurrence as a Miocene fossil. 


Gatun Stage: Port Limon. 
Antigona Blandiana Guppy Plate 30, figures 5, 6 


Venus Blandiana Guppy, 1874, Geol. Mag., London, Decade ii, vol. 1, 
P4366, ply 17.) fs. 6: 

Cytherea (Ventricola\ Blandiana Dall, 1903, Trans. Wagner Free Inst. 
Sci Voli 35 pt. 6) p. 1277. 

Antigona (Ventricola) Blandiana Maury, 1917, Bull. Amer. Pal., vol. 
5, P- 381, pl. 37, fig. 5. 

This species was described by Guppy from the Bowden beds 
of Jamaica but also collected by the Maury expedition in the 
Miocene of Santo Domingo. Dali also records it from the lower 
Miocene of the Chipola beds and from Curacoa, Dutch West 
Indies. It is asmall species of nearly circular outlines and 
sculptured with distant primary lamelle and finer secondary 
threads in the interspaces. 

Our Costa Rican shells consist of two specimens, one from 
the coral limestones near Port Limon, the other from the lower 
Gatun of Comadre Creek near Cahuita. They differ from Dr. 
Maury’s specimens from Santo Domingo in having the primary 
set of lamella more distantly spaced and their interspaces more 
closely striated with the secondary threads. 

Length 30, height 24, diameter 9g mm (left valves) Com- 
adre. 
Length 24, height 23, diameter 8.5 mm (left valve) Port 
Limon. 
Gatun Stage: Comadre creek, near Cahuita. 
Port Limon. 


Antigona Harrisiana, n. sp. Plate 30, figure 3 


Shell small, convex and nearly circular in outline; umbos 
full with the beaks situated a short distance in front of the mid- 
dle; lunule small but broadly cordate and sculptured with the 


242 BULLETIN 39 | 414 


continuation of the concentric ribs and lamella; escutcheon lan- 
ceolate, bordered by an angled ridge from the shell disk; disk 
sculptured with about 28 even, concentric ribs which appear 
as close undulations of the shell surface; the ribs and inter- 
spaces are in addition sculptured with fine lamelle-like threads, 
each finely crenulated or radially striated; these lamellee vary in 
size, those of the interspaces or troughs between the ribs are fine 
and 5 in number, with 2 large ones forming the tops of the ribs 
themselves, so that the ribs appear as if mesially divided; the 
radially striated character of the ribs and lamellz is much strong - 
er on the anterior and posterior submargins; interior of the shell 
deep but concealed by matrix in the type specimens. 

Length 21.5, height 19.5, diameter 6.25 mm _ (right 

valve). 

This elegant species occurs in the coralline phase of the 
Gatun formation in Costa Rica. Its relations are with the 4. 
Blandiana Guppy and the recent A. strigillina Dall, but differs 
from both in its more elegant sculpture. This sculpture consists 
of close, regular, wave-like undulations or ribs, the crests of 
which carry 2, low lamine, the troughs or interspaces, 5 or 6 
finer threads. ‘These fine thread-like laminz are delicately etch- 
ed with fine radial striations. 


It is named for Professor G. D. Harris of the Paleontologi- 
cal Department of Cornell University. 
Gatun Stage: Port Limon. 


Genus CYCLINELLA, Dail 


Cyclinella beteyensis, n. sp. Plate 31, figure 2 


Shell thin, Dosinoid or sub-circular in form, the margins of 
the shell nearly a perfect circle; slightly convex; beaks small 
but distinct, situated at the anterior % of the shell ; sculpture 
with very fine, concentric threads, which are a little elevated and 
strongest on the anterior end, nearly smooth on the middle; lu- 
nule small, lanceolate and defined by a small, incised line; hinge 


45 Costa R1icA MiockNE—OLSSON 243 


normal, the left valve with 3, narrow; cardinal teeth and a wide 
ligament. 
Height 39, length 38? diameter 5 mm left valve. 

Represented by a single, imperfect specimen from the Gatun 
of Rio Betey. It is but slightly convex, and in form approach- 
es very closely a species in the Newcomb collection from the 
Bay of Fonseca, which is probably the C. Avoyeri Philippi. The 
fossil shell is more perfectly circular in form. 


Gatun Stage: Rio Betey. 


Cyclineila subquadrata Hanley, var. quitana, n. sp. Plate 31, fig. 8 


Shell small, thin, convex and subcircular in form; beaks 
small but distinct, with inflated umbo and small beak situated 
at the anterior %; the anterior end is narrow and slightly pro- 
duced, while the posterior end is wide and with the hinge-margin 
is subquadrate in form; the surface is sculptured with fine, ir- 
regular, concentric growth-lines, which are strongest and more 
regular on the posterior dorsal submargin; the interior 1s con- 
cealed. 

Height 28, length 31, diameter of the right valve 7 mm. 


This is a smaller and more convex species than the beteyen- 
sts and like that species, it seems to find its recent analogue on 
the Pacific side, in this case the C. swbqguadrata Hanley. On the 
Pearl Islands in the Bay of Panama I collected a few small 
valves which are probably the C. subguadrata Hanley, but they 
are not so produced anteriorly as is shown in Reeve’s figure. 
These Pearl Island shells are very similiar to the Costa Rican 
fossil, differing mainly in their more inflated umbos and in 
slight difference in form. 

From the Gatun beds of the Canal Zone, Dall has described 
C. gatunensis, but that species is very distinct from the two 
Costa Rican Cyclinellas. 


Gatun Stage; Outtana creek. 


244 BULLETIN 39 416 


Genus CHIONE, Megerle von Miihlfeld 
Chione Rowleei, n. sp. Plate 30, figure 2 


Shell rather large, convex, ovate; lunule broadly cordate 
and concentrically sculptured; escutcheon long, narrow, smooth 
and defined by a small cord-like ridge; surface of the disk with 
about 30 slightly elevated concentric ribs which on the umbos 
are thin and lamellar but lower on the shell disk are wide and 
trianglar at the base, but with a thin zigzag or fluted lamellar 
ridge on top; the interspaces, much wider on the umbos is sculp- 
tured with even rib-like cords which also flute in harmony the 
dorsal face of the concentric ribs; the ventral face of the ribs are 
finely striated by radial incised lines which form cord-like ridges 
along % as wide as the radial cord of the interspaces; interior of 
the shell deep, with a small pallial sinus and subequal adductor 
scars; basal margin finely crenulated as well as the lunular mar- 
gin and the extreme posterior extremity. 

Length 42, height 37.5, thickness (right valve) 17 mm. 

This large, elegant Chione is quite common in the Gatun 
beds of Costa Rica. It also occurs in the lower Gatun at Gatun, 
usually in the form of casts. It recalls in a general way, the re- 
cent West Coast C. amathusia Philippi, but differs in its heavier 
shell, higher and fuller umbos and difference in details of its 
spiral and radial sculpture. In the Costa Rican shell, the radial 
cords are simple, relating it more closely with the small C. 
Woodwardi Guppy of the Miocene of Jamica and Santo Domin- 
go, while in C. amathusia, they are double and alternating. 


The C. tegulum Brown and Pilsbry we have not seen. It 
is a smaller shell, related to the C. Woodwardi. In C. tegulum 
the concentric lamellee are much closer and the wide cordate 
lunule is sculptured only with radial lamellee. 


It is named for Professor W. W. Rowlee, who collected a 
few specimens from the Banana River in the course of botanical 
investigation of the Balsa tree (Genus Ochroma), a very light 
wood used in the construction of life-buoys, etc. 


417 Costa RicA Mi1ocENE—OLSSON 245 


Gatun Stage: Gatun, C. Z. 
Banana River. 
Coll. 6, Estrella River. 


Chione costaricensis, n. sp. Plate 32, figure 4 


Shell of moderate size, convex, ovate-trigonal; the dorsal 
portion of the type specimen is lacking; the surface is sculptured 
with fine, subregular, concentric lamella spaced on the center of 
the shell disk about .75 mm apart; the interspaces and the ven- 
tral surfaces of the lamellee are marked with regular, incised lines 
forming radial bands of an average width of .40 mm; a faint rad- 
ial sinus extends from the beaks to the posterior; ventral margin 
of the shell is slightly arcuate at its posterior end; interior of 
ventral margin finely crenulated. 

Length 34, height 29, diameter of the right valve 10 
mm. 

This shell will be distinguished from the other Costa Rican 
Chiones by its more crowded, concentric, lamelle and regular, 
radial stria. The striz occur on the ventral faces of the lamel- 
lze and on their interspaces. 


Gatun Stage: Hill No. 3, Banana River. 


Section LIROPHORA, Conrad 


Chione mactropsis Conrad Plate 30, figures 7, 8 


Gratelupia? mactropsis Conrad, 1856, Pacific R. R. Reports, vol. 5, p. 
328, pl. 6, fig. 54. 

Chione (Lirophora) mactropsis Dall, 1903, Trans. Wagner Free Inst. 
Sci., vol. 3, pt. 6, p. 1294. 


Chione (Lirophora) mactropsis Brown and Pilsbry, Ig911, Proc. Acad. 
Nat. Sci. Phila., vol. 63, p, 369. 


Chione (Lirophora) ulocyma Brown and Pilsbry, to11, /dem, p. 369. 
Not of Dell. 1903. 

Until the construction work on the Panama Canal had com- 

menced in earnest, the abundant and finely preserved fossils of 


246 BULLETIN 39 4i8 


the Canal Zone, so easily obtained at the present time, were not 
available, and the only fossil remains collected by the early trav- 
vellers were generally in the form of poorly preserved casts. In 
1856 there appeared in the Pacific Rail Road Reports, the de- 
scription of an internal cast as Gratelupia? mactropsts, together 

with scarsely recognizable Turritella gatunensis and altilira. 
These specimens obtained by Mr. W. P. Blake from the 
Isthmus are the first noticed from the fossiliferous rocks in 
Panama and Central America. 


The Livophora mactropsis described by Conrad from an in- 
ternal cast as Gratelupia ?, is one of the most abundant fossils in 
the Gatun beds of the Canal Zone, associated with the small- 
er and more finely sculptured holocyma Brown and Pilsbry. JL. 
mactropsis is a species of variable size, but moderately convex, 
longer posteriorly and usually with the basal margin more or 
less arcuated about the posterior extremity. The sculpture con- 
sists of irregular, more or less confluent concentric lamelle, 
which are overrun by fine radial lines. These radial lines may 
remain strong or become obsolete as usual with specimens from 
the Canal Zone. A narrow foliaceous band is developed on each 
side of the escutcheon and another about the anterior extremity 
below the lunule. The Costa Rican examples are often very 
much larger than those of the Canal Zone, the concentric lamel- 
le are finer and less confluent and with more persistent radial 
lines. The following measurements will illustrate the range in 
size: 

Length 30, heigth 23, thickness 15 mm, Central Zone. 


ce ce 


37) 28, 19 

41.5, 22) 9.5 mm (right valve) 
Old Man Sam Creek, C. R. 

Bilas) Rhigyen 21.5 mm, Sousi, C. R. 


Gatun Stage: Gatun, and Mt. Hope, C. Z. 
Old Man Sam creek, C. R. 
Soust creek (Upper Fone creck) 


Banana River. 


419 Costa Rica MrocENE—OLSSON 247 


Chione holocyma Brown and Pilsbry Plate 30, figure 9 


Chione ulocyma Dall, variety holocyma Brown and Pilsbry, 1911, Proc. 
Acad. Nat. Sci. Phila., vol. 63, p. 369. 

Associated with mactropsts, in the Central Zone is a more 
convex and finely sculptured shell, allied to the wlocyma Dall of 
of the Floridian Miocene. It is the /olocyma of Brown and Pils- 
bry. We have seen no typical wlocyma from Panama. 

From mactropsis, the holocyma will be distinguished by its 
generally smaller, more convex and more circular shell. Its 
concentric lamellae are more numerous and less confluent and 
with fine but persistent radial strice. It is generally best distin- 
guished by its smaller and less clearly differentated foliaceous 
area on its anterior extremity, which in mactropsis is as wide 
an area as that along its posterior dorsal margin. 


Heigth 29, length 36.5, thickness 19 mm. 
Gan Stage Mt. Flope, C- Z. 


Chione hotelensis, n. sp. Plate 32, figure 7 


Shell small, subtrigonal, heavy, but slightly convex; sculp- 
ture of about 15, thick, rounded and evenly spaced concentric 
lamellee; on the posterior-dorsal slope, the ends of the lamellce 
become attenuated, thin and somewhat appressed as in glyptocy- 
ma Dall of the Flordian Miocene; the lamellze themselves are 
smooth and rounded but with their interspaces finely radially in- 
cised by small lines; escutcheon rather large, smooth; lunule 
small, cordate and smooth except for the growth lines, 

Length 23, height 19, thickness 12.5 mm. 

This elegant little species is distantly related to the C. gly- 
ptocyma Dall from the Oak Grove sands of Florida, but isa 
smaller and more compact shell. The radial striations commence 
on the ventral surface of the lamellze and are continued across 
the interspace to the base of the next rib. 

All of our specimens were collected in a hard sandstone in 
the lower Gatun of Hotel Creek near Old Harbor, Costa Rica. 


248 BULLETIN 39 420 


Uscart Stage: Hotel creek, C. R. 


Chione chiriquiensis, u. sp. Plate 32, figure 9, Ic 


Shell small, solid, depressed; lunule rather large, cordate, 
smooth; escutcheon large, smooth and defined by an angled 
ridge; surface of the disk sculptured with about 8, large, irregu- 
lar ribs, like those of Chione latilirata; these ribs do not extend 
to the edge of the escutcheon, but commence a short distance in 
front leaving a narrow band which is smooth and sculptureless; 
the ribs are of variable size, flat-topped and smooth; they are 
rounded on their ventral side, but shelving or recurved back- 
wards on the dorsal face; interspaces smooth or only sculptured 
with fine growth-lines; interior of shell of moderate depth, with 
a small pallial sinus and with a slightly larger posterior adduct- 
or scar; basal margin finely crenulated. 

Length 13.75, height 12.50, diameter 9.50 mm. 

The few specimens which we have from Water Cay, are 
probably immature, but they possess such distinctive characters 
that the species cannot be mistaken. 

The heavy, irregular concentric ribs recall certain varieties 
of Chione latilirata Conrad. ‘They differ in being abruptly cut 
off or flattened a short distance behind the dorsal-posterior angle 
leaving a band asin C. paphia Linnzus, but smooth and sculp- 
tureless. The interspaces are smooth, 

Gatun Stage: Water Cay. 


Section TIMOCLEA, Brown 
Chione grus Holmes, var. Plate 26, figure 21 


Tapes grus Holmes, 1858, Post-Pliocene Fossils of S. Car., p. 37, pl. 7 
fig. 5. 
Chione ( Timoclea) grus Dall, 1903, Trans. Wagner Free Inst. Sci., 
vol. 3, pt. 6, p. 1299. 
We have but a single and not entirely perfect specimen of a 
Timoclea closely related to C. grus Holmes. It is somewhat 


42t Costa Rica MI1ocENE—OLSSON 249 


more convex than typical C. grvus from the Carolinas and more 
finely sculptured. More material is needed to show whether or 
not these charaters are constant. The C. gvus appears asa 
fossil in the Upper Chesapeake Miocene of the eastern United 
States and is recent from Cape Hatteras to Yucatan. 


Gatun Stage: Coll. 4, Red Cliff creek. 


Genus TELLINA, (Linné) Lamarck 
Teilina rovala, n. sp. Plate 26, figures 7, 8 


Shell elongate, slender, subequilateral with the beaks situ- 
ated near the center of the shell; anterior side long, with a 
straight dorsal margin and narrowly rounded at its extremity; 
posterior side much narrowed and rostrate, the left valve with 2 
radial bands forming the rostrum, the dorsal one a fold, the oth- 
era wide sinus; ventral margin rounded anteriorly, nearly 
straight about the middle and a wide sinus posteriorly; the sur- 
face is sculptured with even, concentric lamelle; this sculpture 
on the rostrum is heavier and on crossing the fold and sinus be- 
come reversed in position. 

Length 29, height 12, semi-diameter 2.5 mm. 
36? T4 3 mm. 

We have two specimens of this typical 7e//ima from Water 
Cay. The larger specimen is fragmentary but probably measur- 
ed when complete about 36 mmin length. The species recalls 
the recent West Indian 7. zxzterrupta Wood (7. antonzz Philippi) 
in its form and sculpture but lacks the finer radiating strie. 


Gatun Stage: Water Cay. 
Subgenus ARCOPAGIA, (Teach) 
Section MERISCA, Dall 


Tellina crystallina Wood Plate 26, figure 10 


Tellina crystallina Wood, 1815, General Conchology. 


250 BULLETIN 39 422 


Tellina (Merisca) crystallina Dall, 1901, Proc. U. S. Nat. Mus., vol. 
Pew oye ener ols 7s) never.) 3(6) 

Tellina (Merisca) crystallina Maury, 1917, Bull. Amer. Pal., vol. 5, p. 
387, pl. 38, fig. 4. 


This Zel/ina, the type of the section JZerisca, is recognized 
by its thin, depressed shell, strongly sculptured with distant, but 
regular, raised, concentric lamellee. It is very rare as a fossil, 
our collection containing but a single specimen from the Gatun 
beds of the Banana River. It has previously been recorded by 
Maury as a Miocene fossil in Santo Domingo. 

In the recent fauna, Tellina crystallina is distributed along 
both the Atlantic and Pacific coasts of Central America. I have 
collected a few specimens from the beaches of Bocas Island and 
more abundantly at Bucaru, Province of Los Santos, on the 
Pacific side of Panama. 


Gatun Stage: Hill No. 2. Banana River. 
Tellina nasua, n. sp. Plate 26, figure 17 


Shell sub-rhomboid, slightly convex, thin; beaks small, in- 
conspicuous, posterior to the middle; the anterior end longer, 
the dorsal-anterior dorsal margin descending, the extremity 
rounded into the gentle curve of the base; (the posterior side 
broken in the type specimen); the greatest convexity of the shell 
lies along the broad, umbonal slope anterior to the beaks; the 
surface is finely sculptured with regular, sharp, concentric 
ridges, separated by narrow, band-like interspaces; interior of 
the right valve with a large pallial sinus which extends to the 
anterior 1% of the shell; hinge appearing normal for the section, 
but partly broken away, but showing a large anterior lateral 
tooth. 

Length 20? Height 15.25, diameter of the right valve 
3.00 mm. 

Our specimen if fragmentary, the posterior portion of the 
shell being broken away. There is however little doubt that 
the species is a true JZerisca, allied to 7. egutstriata Say and 


423 Costa RicA M1ocENE—OLSSON 251 


TZ. lintea Conrad. It differs from eguzstriata in its greater dor- 
sal anterior slope and finer sculpture, and from 7. /zztea of the 
recent fauna, in its somewhat greater proportional height and 
greater convexity. 


Gatun Stage: Coll. 4, Red Cliff creek. 


Section PHYLLODINA, Dall 
Tellina lepidota Dall Plate 26, figures 4, 5 


Tellina (Phyllodina) lepidota Dall, 1900, Trans. Wagner Free Inst. 
El mVOla 35 Pts 5, .p: LO22. pliv AO whoa les 

Dall’s type of this species is based on a fragmentary right 
valve from Gatun. We have several complete and large speci- 
mens from Water Cay, where the species is fairly common. The 
right valve is flat and depressed, the left more convex. Both 
valves are nearly similarly sculptured with 2 or 3 wide undul- 
ations of the beaks, which are soon replaced by even, sharp, con- 
centric lamellee, of which 3 or 4 are higher and more lamellose 
on the dorsal margin. 


Length 24, height 12.5, diameter of the left valve 2.75 


mim. 
Length 21.5, height 11.00, diameter of the right valve 
eA sonben, 


Gatun Stage: Water Cay. 


Section EURYTELLINA, Fischer 


Tellina costaricana, n. sp. Plate 26, figures 6, 9 


Shell oblong, depressed; beaks small and inconspicuous 
about the middle of the shell so that the anterior and posterior 
ends are of nearly the same length; posterior end widely round- 
ed the anterior but slightly narrowed and less _ round- 
ed at its extremity; anterior and posterior dorsal margins nearly 
straight; surface of the disk finely sculptured with even, con- 


252 BULLETIN 39 424 


centric lines which are slightly lamellose on the posterior-dorsal 
margin; interior concealed. 


Length 32, height 19, semi-diameter 2 mm. 


This species is related to the recent West Indian 7. angulosa 
Gmelin and its Pacific analogue, the 7. rubescens Hanley. The 
fossil shell differs by its more central beaks and sharper concen- 
tric sculpture, which is continued across the entire surface of 
the disk. 


Gatun Stage: Hill No. 3, Banana River. 


Tellina dariena Conrad Plate 26, figure 3 


Tellina dariena Conrad, 1857, Pacific R. R. Reports, vol. 5, p. 328, 
pl. 6, fig. 53. 

Tellina semilevis Gabb, 1861, Proc. Acad. Nat. Sci. Phila., vol. 12, p. 
567. 

Tellina Dariena Gabb, 1881, Journ. Acad. Nat. Sci. Pnila., vol. 8, p. 
343, pl. 44 fig. 13. 

Tellina Rowlandi Toula, 1908, Jahrb. der K-K. Geol. Reichsanstalt 

vol. 58, p. 728, pl. 28, fig. rr. 

Tellina Rowlandi Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. 

Phila., vol. 63, p. 368. 

Conrad’s figure of 7e/lina dariena is very poor but probably 
represents this species, the most abundant 7Ze//zma at Gatun. 
Conrad’s figure shows a subtrigonal shell, with a produced and 
truncated posterior extremity. In 1861, Gabb described Jellina 
semtlevts together with Arca chiriguiensts from the Gatun beds 
of the Chiriqui Lagoon. This species Gabb however later con- 
sidered synonymous with the earlier 7e//ina dariena of Conrad. 
Toula’s TZellina Rowlandi is certainly identical with Gabb’s 
Tellina semilevis as figured by Gabb. 

Shell but slightly convex, subequilateral with the beaks sit- 
uated a short distance posterior to the middle of the shell; anter- 
ior and posterior dorsal slopes straight and evenly descending; 
posterior end rounded and subtruncate, the anterior rounded ; 
beaks low; surface striated with very fine concentric lines and 


425 Costa R1icA MIocENE—OLSSON 253 


the shell may sometimes appear nearly smooth and polished 
especially on the umbonal areas; hinge of the left valve with a 
bifid anterior cardinal, a small posterior and 2 laterals; the an- 
terior adductor is bordered on the posterior side by a thickened 
internal ray; pallial sinus long, ascending above in the middle 
of the shell and touching the anterior adductor scar. 
Length 34, height I9.5, semi-diameter 3.50 mm. 
Gatun Stage: Gatun, C. Z. 
Water Cay. 
Chiriqui Lagoon and Oronli creck (Gabo) 


Subgenus ANGULUS, Mergerle 


Section ANGULUS, s. s. 
Tellina umbra Dall Plate 26 figures 12, 15 


Tellina (Angulus) umbra Dall, 1900, Trans. Wagner Free Inst. Seine 
vol. 3, pt. 5, p. 1033, pl. 46, fig. 13. 


The Costa Rican specimen consists of a small individual 
with both valves. It is somewhat smaller than typical wmora, 
from the Upper Chesapeake Miocene of the Carolinas, but agrees 
in form and in the fine surfrce sculpturing. 

The Tellina umbra is a common fossil in the Upper Chesa- 
peake Miocene of the eastern United States and in the Pliocene 
of Florida, It is closely allied tothe recent Tellima sybaritica 
Dall of the West Indies, differing most importantly in its less an- 
gular posterior extremity. 

The Costa Rica specimens measures: 

Length 8.00, height 4.15, diameter 2.25 mm. 


Gatun Stage: Coll. 4, Fast Grape Point creek. 
Section SCISSULA, Dall 


Tellina cercadica Maury 


Tellina (Scissula) cercadica Maury, 1917, Bull. Amer. Pal., vol. 5, p. 
388, pl. 38, fig. 9. 


254 BULLETIN 39 426 


The Costa Rican specimens are fragmentary but there can be 
little doubt of their identity with the TZel/ina cercadica Maury 
from the Miocene of Santo Domingo. 

The shell is thin and strongly depressed. ‘The surface is 
polished and obliquely striated with fine lines. It is related to 
the recent Pacific 7ellina delicatula Deshayes. 


Gatun Stage: Coll. G, Saury creek. 
Estrella River. 


Genus TELLIDORA, Moerch 
Tellidora cristata Recluz Plate 26, figurce 1, 2 


Lucina cristata Recluz, 1742, Revue Cuvier., p. 270. 
Tellidova cristata Dall, 1900, Trans. Wagner Free Inst. Sci., vol. 3, 
pt-.4) \p) 1037. 

A recent species of the West Indies and previously knowa 
as a Pliocene and Pleistocene fossil from Carolinas. The fossil 
Costa Rican shells are frequently larger than the recent form 
and with the dorsal margins more nearly in line. It is not rare 
in the form of impressions but the perfect shell is seldom found. 


Gatun Stage: Hills ra, 2 and 3, Banana River. 


Genus STRIGILLA, Turton 
Strigilla pisiformis Linné 


Tellina pisiformis Linné, 1758, Syst. Nat., ed. 10, p. 677. 

Strigilla pisiformis Dall, 1900, Trans. Wagner Free Inst. Sci., vol, 3, 
pt. 5, p. 1038. 

Strigilla pisiformis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 389, pl. 
39, fig. 6. 

This species is rare asa fossil in Costa Rica. It is very 
common asa recent shell on the sandy beaches along the en- 
tire north coast of both Panama and Costa Rica, where it occurs 
with Donax denticulata Linné and afew other species. It is 


427 CosTA RicA MiocENE—OLSSON 255 


also found as a Miocene fossil in the Bowden beds of Jamaica 
and in Santo Domingo. 


Gatun Stage: Middle creek. 


Strigilla musanica, n. sp. Plate 26, figure 18 


Shell subtrigonal, moderately convex, subequilateral; the 
umbos are broad and wide, situated about the middle of the shell, 
with the greatest convexity of the shell lying along the anterior- 
umbonal slope; the anterior end is broad and rounded, the pos- 
terior slightly narrow, but not produced; the sculpture is divid- 
ed between three areas, the posterior, central and anterior, each 
of which is set off from the adjacent one, by a marked change in 
sculpture along a well-defined line; on the anterior area, it con- 
sists of a concentric series of wave-like incised lines, the crest of 
the wave lying near the posterior side and directed as usual dor- 
sally; on the central area, the lines are vertically directed, com- 
mencing on the lines bordering the posterior side; on the poster- 
ior area, the lines commence on the anterior side and pass verti- 
cally, forming therefore with the middle lines very acute an- 
gles; the interior is concealed in our specimen. 

Length 13.00, height 11.50, diameter of the right valve 
3.00 mm. 

Differs from Strigzlla carnaria Linné of the recent West In- 
dian fauna, by being more inequilateral, more convex and in de- 
tails of its sculpture which is sharply divided between three areas 
as described. ‘The interior of the type specimen is concealed, so 
that the features of the pallial line and sinus cannot be deter- 
mined. 


Gatun Stage: Hill No. 3, Banana River. 
Genus METIS, H. and A. Adams 


Metis barba, n. sp. Plate 29, figures 3, 4 


Shell subquadrate, inequilateral, with the beaks situated at 
the anterior two-thirds of the total length of the shell; anterior 


256 BULLETIN 39 428 


end longer with a sloping dorsal margin and rounded anterior 
extremity; posterior side shorter, with a dorsal margin sloping 
more rapiding to the wide and flexed posterior end; the left valve 
is convex but depressed along the narrow posterior submargin or 
wing; the right valve is less convex with a broad depressed area 
extending from the beaks to the ventral margin; this depressed 
zone is sharply separated from the posterior-dorsal slope or wing 
by a strong angle or flexure which corresponds to the sinus in 
the middle of the wing on the left valve; surface finely striated 
with concentric growth-lines. 


Length 48, height 37, diameter 19 mm. 


Pilsbry and Johnson have described without figuring, 
two species of Metis from the Miocene of Santo Domingo (the 
M. efferta and postrema). ‘These species, they compare with the 
M. chipolana Dall, from the Chipola beds of Florida and men- 
tion in their description the nearly central location of the beaks. 
In the Costa Rican shell the beaks are at the posterior one-third 
of the shell. It is very distinct from the (7. chipolana Dall. JZ. 
trinttaria Dall from ‘Trinidad and Porto Rico differs in its more 
central beaks, fuller umbos and has the right valve much less 
flexed posteriorly. 


Gatun Stage: Betey creek. 


Genus MACOMA, Leach 
Macoma costaricana, n. sp. Plate 26, figures 14, 16 


Shell broadly ovate, thin and slightly inflated; dorsal and 
ventral margin straight and parallel; the posterior dorsal side 
sloping to the rounded ventral extremity; anterior extremity 
rounded; the posterior side is very slightly flexed; surface 
smooth except for very fine, incremental growth lines; interior 
of shell concealed. 

Length 29, height 19, semi-diameter 4.5 mm. 
The reference of this species is Macoma is doubtful as the 


429 Costa Rica M1ocENE—OLSSON 257 


hinge is completely concealed in all our specimens. It has the 
form of the JZ. Conradi Dall, from the Chesapeake Miocene of 
United States but is higher and differs in many other respects. 
It is shorter and higher than 4. gatunensis Toula with which it 
miay occur. 
Gatun Stage; Hill No. 3, Banana River. 
kio Betey. 


Subgenus PSAMMACOMA, Dall 
Macoma gatunensis Toula Plate 26, figure 13 


Macoma ( Tellina) gatunensis Toula, 1908, Jahrb. der K-K Geol. Reich- 
sanstalt, vol. 58, p. 729, textfigure Io. 


Tellina gatunensis Brown and Pilsbry, IgtI, Proc. Acad. Nat. Sci. 
Phila., vol. 63, p. 368. 


This shell is extremely similiar to the recent M. elongata 
Hanley of the Pacific coast of Panama and I have failed to find 
any character which will distinguish the fossil shells from the 
recent ones. Our shells were all collected loose near Hill 3, of 
the Banana River and were probably derived from beds low in 
the Gatun formation. 


Gatun Stage: loose, near Hill 3, Banana River (Terry). 


Macoma panamensis Dall, var. canatis, n. var, Plate 26, figure II 


Cf. Macoma (Psammacoma) panamensis Dall, 1900, Proc. U. S. Nat, 
Mus., vol. 23, p. 324, pl. 4, fig. 3. 

The Vacoma panamensis Dall was described from a left 
valve dredged from 33 fathoms of water in Panama Bay. ‘The 
fossil shell, also a left valve from the Gatun of Mt. Hope, differs 
most importantly from Dall’s figure in being somewhat higher. 
The shell is thin and extremely fragile, with the surface simply 
scultured with the fine growth lines. The interior is concealed 
so that the characters of the pallial sinus cannot be seen. The 
specimens measures as follows: 


258 BULLETIN 39 430 


Length 30.50, height 15.00, diameter of the left valve 
3.00 mm. 


Gatun, Stage Mi Fope. (Gi Zh 


Genus SEMELE, Schumacher 
Semele levis Sowerby, var. costaricensis, u. var. Plate 29 figure 1 


Cf. Amphidesma levis Sowerby, in Reeves, 1853, Conch. Icon., pl. 7, 
fig. 50. 


Shell large, subelliptical, but slightly convex; beaks very 
nearly central, the posterior side wide and well rounded, the an- 
terior somewhat narrowed, contracted and. slightly the longer; 
the surface is nearly smooth and porcellaneous, faintly marked 
on the anterior and posterior extremities by fine, concentric 
lines. 


Length 72, height 65.50, semi-diameter 8 mm. 

This large Semele is quite common in the Lower Gatun beds 
of the Banana River. It is closely related to the recent Pacific 
S. levis Sowerby, but differs in being slightly longer and more 
evenly rounded at its posterior extremity. 


Gatun Stage: Hill No. 3, Banana River. 
fiiull ra, Banana River. 


Semele Sayi Toula Plate 29, figure 8 


Semele Sayt Toula, 1908, Jahrb. der K-K Geol. Reichsanstalt, Wien, 
vol. 58, p. 730, pl. 28, fig. 17. 

Semele Sayt Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila. 
vol. 63, p. 368. ; 

Shell convex, rather thin, with the umbos nearly central. 
The anterior side is somewhat narrow or contracted, and round- 
ed at its extremity. The surface is sculptured with fine, hair- 
like, concentric threads. The species was described by Toula 
from the Canal Zone. Our record from Costa Rica is based ona 
single, imperfect specimen from Sousi creek, where it occurs as- 
sociated with Clementia, Chione mactropsts and Turritella altilira 


431 Costa R1rcA MIocENKE—OLSSON 259 


variety costaricenszs in the lower part of the Gatun formation. 


Gatun Stage: Sousi creek. 


Semele Claytoni Maury Plate 29, figure 2 


Semele Claytont, Maury, 1917, Bull. Amer. Pal., vol. 5, p. 227, pl. 35, 
fig. 9. 


The S. Claytont was described by Dr. Maury from the Cer- 
cado formation, Santo Domingo of Lower Miocene age. Itisa 
large species, reaching a length of 66 mm or more and charact- 
eristically sculptured with even, concentric lamellz like the re- 
lated species from the Tertiaries of eastern United States as chipo- 
lana Dall, Leana Dall and perlamellosa Heilprin. From these, 
the S. Clayfonz is distinguished in having its radial strize nearly 
obsolete. Our specimen from Costa Rica is fragmentary but 
probably measured when perfect 38 by 30 mm. 


Uscart Stage: Hotel creek. 


Genus ABRA, (Leach) Lamarck 


Abra triangulata Dall Plate 29, figure 10 


Abra triangulata Dall, 1900, Trans. Wagner Free Inst., Sci., vol. 3, 
pt. 5, p. 997, pl. 49, fig. 4. 


A small, rotund or trigonal species abundant in the shale 
beds of Bocas Island. ‘The valves are nearly equilateral, the an- 
terior end slightly the larger and inflated. Surface smooth and 
shining. The type was described by Dall from the Bowden beds 
of Jamaica. Our specimens are a little larger, measuring: 


Wength 7.25) height |\6450, diameter of) the) left) valve 
1.75 mim. 


Gatun Stage: Bocas del Toro. 


260 BULLETIN 39 A432 


Genus PSAMMOSOLEN, Risso 


Psammosolen gatunensis Toula, var. costaricensis, n. var. 


Plate 29, figure 7 
Solecurtus gatunensis Toula, 1908, Jahrb. der K-K. Geol. Reichsan- 
stalt, vol: 58, p. 732, pl. 28, fig. 12. 


Solecurtus gatunensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. 
Phila., vol. 58, p. 372. 


Shell rather large, elongate, convex, the dorsal and ventral 
margins straight and parallel; the low, inconspicuous beaks sit- 
uated at the anterior 4, the anterior end rounded, the posterior 
subtruncate; two low, radial depressed rays extend from the 
beaks obliquely to the ventral margin; the central part of the 
shell and the anterior end is sculptured roughly with irregular 
growth lines; the posterior dorsal slope has strong, vertical or 
slightly oblique lines which cross the finer concentric growth 
lines; in addition the whole surface of the shell is roughened by 
small, short threads as if the surface had been painted with a 
stiff brush. 


Length 63, height 25.50, diameter of the right valve 
14.50 mm. 

Toula’s figure of an internal mold shows a smaller shell, a 
strong anterior-umbonal angle and a somewhat shorter anterior 
end. It is possible that the two forms are distinct but we lack 
a good comparative series of the Gatun shell. A smaller and 
more delicate species of Psammosolen occurs at several localities 
in the Costa Rican Gatun. It is not the P. strigzl/atus (L) of 
Toula, differing in its pointed anterior end. 


Gatun Stage: Rio Betey. 
Genus SANGUINOLARIA, Lamarck 
Section SAMMOTELLA, Blainville 


Sanguinolaria alouatta, n. sp. Plate 29, figures 5, 6 


Shell rather large, elongate; and rostrated behind; left 


433 Costa RicA M1ocENE—OLSSON 261 


valve nearly flat, the right valve much more convex; the right 
valve carries a strong umbonal keel from the tip of the beaks to 
the posterior extremity; this keel is but poorly developed on the 
left valve; anterior end broadly rounded with the dorsal and 
ventral margins nearly parallel; the posterior side is much nar- 
rower and with its extremity pointed; the surface is smooth; the 
dorsal-posterior slope of the left valve is less smooth and carries 
coarse, irregular growth-lines; the interior is not known. 
Length 54, height 30, semi-diameter 4 mm (a left valve) 
Length 60, height 27.5 semi-diameter 7 mm (a right 
valve) 

This species stands intermediate between the S. Smzthwood- 
wardt Maury from the Miocene of Santo Domingo and the re- 
cent West Indian S. oferculata Gmelin. It differs from the S7zz¢h- 
woodward, in being much larger with a less depressed left valve: 
From oferculata the Costa Rican shell has higher and more 
pointed beaks as well as being longer posteriorly. In full-grown 
specimens of operculata, the surface generally shows radial lines 
and striz which may become quite strong near the posterior 
ventral margin. The Costa Rican shells are smooth except for 
the growth lines. 


Gatun Stage: Hill No. 3, Banana River. 


Genus TAGELUS, Gray 
Tagelus cebus, n. sp. Plate 29, figure 9 


Shell elongate, moderately convex and solid; beaks central 
with the anterior and posterior ends nearly equal, and with the 
dorsal and ventral margins parallel and nearly straight; anterior 
and posterior extremities subtruncate; a broad depressed zone 
extends from the umbos to the ventral margin of the valves; 
surface marked with irregular concentric lines of growth; inter- 
ior concealed. 


Length 36, height 13, diameter of the left valve 3.00 
mim. 


262 BULLETIN 39 434 


The Szligua subegualis Gabb, from the Miocene of Santo 
Domingo is doubtless a 7agelus, belonging to the Section J/es- 
opleura Conrad. It differs from the present shell in being thin- 
ner, more depressed and more pointed posteriorly. Gabb’s species 
approaches very closely the recent 7. divisus Spengler of the 
West Indian fauna, judging from the specimens in the Maury 
collection at Cornell, differing mainly in its larger size. 


The interior of the Costa Rican shell is concealed so that the 
presence or absence of an internal rib cannot be determined. 
Its heavier shell and general form, is more that of typical 7age- 
lus differing from the recent gzbdus Spengler in its more central 
beaks. 


Gatun Stage: Rio Betey. 
Genus MACTRA (L.) Lamarck 


Subgenus MACTRELLA, Gray 


Kactra exoleta Gray Plate 28, figure 2 


Mactra exoleta Gray, 1837, Mag. Nat. Hist., p. 372. 
Mactra exoleta Reeve, 1854, Conch. Icon., Mactra, fig. 16. 


Comparision of our fragmentary shell with recent examples 
from the Bay of Panama show no inportant differences. The 
species is notable for its high, narrow, inflated umbos, strongly 
depressed posterior-dorsal margin and alation of its dorsal-pos- 
terior angle. 


At the present time, this species is restricted to the Paciliic- 
side, but represented on the Caribbean coast by the next, the 
Mactra alata Spengler, common along sandy beaches. It is 
therefore of more than usual interest to find these two species, 
together in the Miocene beds of the Caribbean slope of Costa 
Rica. 


Gatun Stage: Boucary creek. 


435 Costa RicA M1iocENE—OLSSON 263 


Mactra alata Spengler Plate 28, figure 3 


Mactra alata Spengler, 1802, Skriv. Naturch. Selsk., vol. 5, part 2, 
Pp. 99. 

Mactra alata Reeve, 1854, Conch. Icon., Mactra pl. 8, fig. 29. 

Mactra (Mactrella) cf. alata Maury, 1917, Bull. Amer. Pal., vol. 5, 
P- 395- 

A thin, fragile species distinguished by its sharp dorsal-pos- 
terior angle and hatchet-shaped shell. It is related to the exo- 
feta, already referred to, but is a very much broader shell. In 
exoleta, the dorsal-posterior area is flat or concave, with a slight 
raised line which is bounded on each side by a markly depressed 
or sunken band. In a@/afa this area is convex. 


It is living today on the Caribbean side of the Isthmus, be- 
ing quite common on sandy beaches. 


Gatun Stage: Boucary Creek. 
fill No. 3, Banana River. 


Section HARVELLA, Gray 


Macira estreilana, n. sp. Plate 28, figure 1 


Shell rather large, ovate, thin, slightly convex; surface reg- 
ularly, concentrically plicated; the fragmentary type specimen 
about 38 mm in height has about 32 plicee which on the center 
of the shell disk are spaced about 2 mm apart; these pliceze com- 
mence as in typical arvella at the dorsal-posterior carina which 
bounds the escutcheon and extends across the surface to the an- 
terior extremity; the surface is, in addition, finely, concentrically 
striated; lunule large elliptical, smooth or finely striated; escut- 
cheon smooth; interior not known. 


Length 50, height 38, semi-diameter 8 mm, 
Although this is not a very rare species in the Costa Rican 


Miocene, its thin, delicate shell renders the collecting of good 
specimens very difficult and it is usually secured only as distort- 


2604 BULLETIN 39 436 


ed internals molds. It is related to the recent Pacific 7. elegans 
Sowerby, but differs in being less convex, wider and with more 
numerous and closer concentric plicee. 
Gatun Stage; Coll. 6. Estrella River. 
Coll. 2, Quitana creek. 


Macira sincola, n. sp. Plate 28, figure 4 


Shell thin, broadly subtrigonal, depressed, nearly equilater- 
al; beaks small and pointed anteriorly; lunule absent; escutche- 
on linear-lanceolate and defined from the shell disk by an elevat- 
ed carina; the surface is sculptured with strong, wave-like, con- 
centric plicee which are continuous from the anterior-dorsal mar- 
gin to the edge of the escutcheon; on the type specimen there 
are about 30 pliceze on the center of the shell disk, spac ed about 
I.75 mm apart; the intervals between the concentric plicz 
are deep and trough-like and in width equal to the plice them- 
selves; in addition the whole surface of the shell is finely sculp- 
tured with small, radial, vermiculate threads, best seen on the 
summits of the plicze; the escutcheon is ridged on the middle 
and smooth; the hinge, largely concealed in the sandy matrix, is 
mactroid. 

Length 36, height 27, diameter of the right valve 5.5 
mm. 

The systematic position of this interesting shell is very 
doubtful. Its general surface sculpture of strong, concentric 
plicze and well-defined escutcheon resembles that of Havrvella. 
The surface is covered with fine, vermiculate threads as seen on 
Labiosa. 


Gatun Stage: Saury creck. 
Genus LABIOSA, (Schmidt) Moller 


Labiosa lineata Say Plate 21, figure II 


Lutraria lineata Say, 1821, Journ. Acad. Nat. Sci. Phila., vol. 2, p. 
310. 


437 Costa Rica MIocENE—OLSSON 265 


Mactra Nuttallii Reeve, 1854, Conch., Icon., Mactra, fig: 125. 
Labiosa lineata Dall, 1898, Trans. Wagner Free Inst. Sci., vol. 3, pt. 
4, p- go6. 

This species is represented in our collection by fragments of 
the hinge and a perfect mold of the interior which is figured. 
This mold compares very closely with recent specimens of Ladz- 
osa lineata Say. 

Dall records this species as fossil in the Pliocene Caloosa- 
hatchie beds of Florida and we have several fragments in the 
Cornell collection from the Upper Chesapeake Miocene of Nat- 
ural Well, North Carolina. 


Gatun Stage: Saury creek. 
Coll. 6, Estrella River. 


Genus MULINIA, Gray 
Muiinia lateralis Say, var. sauryensis, n. var. Plate 28, figures 23. 24 


The species of Mulinta are extremlly variable and we have 
therefore associated this form with the common /atevalis of Say, 
although it may later prove to be a distinct species. 

The general form of the shell is like corbulotdes Deshayes, 
the common Gulf variety of lateralis of Say. It differs most im- 
portantly from /aferalis in having the beaks situated a short dist- 
ance above the hinge-line as is seen in the Chesapeake Miocene 
M. congesta Conrad. In other characters, the Costa Rican shells 
are more closely related to the J/atervalis, than to the congesta. 
The shell is fairly convex and thin. 

Length 20, height 16, semi-diameter 6 mm. 


Gatun Stage: Zone E, Saury creek. 
Genus MESODESMA, Deshayes 
Mesodesma costaricensis, n. sp. Plate 28, figure 18 


Shell ovate, moderately convex, inequilateral, thin; anterior 


266 BULLETIN 39 438 


end about % again as longas the posterior, rounded; beaks 
high, triangular; posterior and anterior dorsal margins straight; 
ventral margin evenly rounded; surface nearly smooth; but with 
the dorsal submargins finely concentrically grooved; interior 
concealed. 


Length 22, height 16, semi-diameter 4 mm. 
A rather thin, Dozax-like shell from the Lower Gatun beds 


of the Banana River. Its surface is smooth and polished with 
concentric grooves on its posterior and anterior dorsal area. 


Gatun Stage: Hill No. 3, Banana River. 
Genus GASTROCHAENA, (Spengler) Cuvier 


Gastrochena rotunda Dall 


Gastrocheena ovata Sowerby, variety rotunda Dall, 1898, Trans. Wag- 
ner Free Inst. Sci., vol. 3, pt. 4, p. 825. 


Gastrochena rotunda Dall, 1915, Bull. N. S. Nat. Mus., No. 90, p. 157, 
pl. 19, fig. 2. 

A single, small specimen of a Gastrochena was collected in 
the Gatun of Red Cliff Creek. It is widely expanded about the 
posterior portion and probably represents a young shell of G. ro- 
tunda which Dall records from the Bowden beds of Jamaica. 
This species is alsofound in the Tampa Silex beds and in the 
Lower Miocene of Florida. 


Gatun Stage: Coll. 4, Red Cliff Creek. 
Genus CORBULA, Lamarck 


Section ALOIDIS, Megerle 


? Corbula vieta Guppy Plate 28, figures 15, 16 


Corbula meta Guppy, Quart. Journ. Geol. Soc. London, vol. 22, p. 
580, pl. 26. fig: 8. 

Corbula disparilis Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 
247. 

Corbula (Aloidis) vieta Dall, 1898, Trans. Wagner Free Inst. Sci., 
vol. 3, pt. 4, p. 859. 


439 - Costa RicA MiocENE—OLSSON 267 


Corbula (Aloidis) vieta Maury, 1917, Bull. Amer. Pal., vol. 5, p. 395, 
pl 39, fig. 13. 


The members of this group of Corbule are difficult to sepa- 
rate satisfactorily as all are quite similiar in their characters. The 
right valve is broad, high and coarsely sculptured with regular 
concentric coste, while the left valve is much smaller, longer 
than high, with its surface smooth or faintly lined with un- 
equally distributed radial threads. 


The C. viefa was described by Guppy from the Manzanilla 
beds of Trinidad and most authors have regarded the A7ycina 
tensa described at the same time as its left valve, an indentifica- 
tion which is open to question. Guppy’s figure of fezsa, shows 
a Veneroid shell strongly sculptured with even radials and 
fine concentric lines, and quite unlike the left valves of Corbule 
of the A/ozdzs section. 


? Corbula vieta from Costa Rica is identical with Maury’s 
specimens from the Miocene of Santo Domingo. It is a smaller 
and more circular shell than the Coréula heterogenea Guppy 
which we have from the Canal Zone. 


Length 6.25, height 6.50, semi-diameter 3 mm. (right 
valve) 


Gatun Stage: Water Cay. 
Hull 1a, 3. Bananavketver 


Section CUNEOCORBULA. Cossman 
Corbula oropendula, n. sp. Plate 28, figures 12, 13, 14, 26, 27 


Shell small, solid, elongate, both valves of nearly equal con- 
vexity but the the right valve somewhat larger and overlapping 
on the left; beaks nearly central, the anterior end well rounded, 
the posterior somewhat contracted, slightly rostrated and with 
a well-marked keel; ventral margin more or less rounded, ter- 
minating atis posterior end ina small sinus, just anterior to 


268 BULLETIN 39 440 


the keel; the basal margins of both valves are folded in and 
flattened as seen in C. contracta Say but toa less extent and 
least in the right valve; surface sculpture of rather heavy con- 
centric costee, most widely spaced on the umbos; the surface 
may also show fine radial lines of small pustules or puncte 
which are best developed on the rostrum; no lunular pit. 
Length 6.5, height 4.25, diameter 4 mm (Banana River ). 
9 6 semidiameter 2 mm (Saury). 
We have chosen as the type of this spceies, the smaller and 
more abundant form as found in the Gatun bed of the Banana 
River, rather than the larger shells from the more inaccessible 
Saury. It is related to the C. contracta Say, but differs in hav- 
ing the ventral margin more curved, and the posterior extremi- 
ty more emarginate. Its surface is sculptured with rather 
strong, concentric costee which are more widely spaced on the 
umbos, and submicroscopic radial lines of fine punctze or small 
pustules, best seen on the rostral areas. 
Gatun Stage: Hills ra, 3, Banana river (abundant). 


Zone G, Saury creek. 
Gol On Rea Oli enecr: 


Coraula Dietziana C. B. Adams Plate 21, figure 8 
Corbula Dietziana C. B. Adams, Contr. Conch., vol. 12, p. 235. 


Corbula (Cuneocorbula) Dietziana Dall, 1898, Trans. Wagner Free 
Imsty Seis) voli2. pt Ayip isso: 
A recent species quite common in the coralline limestones 
of the Limon Peninsula, from which it has previously been re- 
ported by Dall from specimens collected by Hill. 


In its earlier stages of growth, this species resembles the 
C. eguivalvis Phil._(C. cubantana d’Orb) in its rectangular from 
and strong dorsal keel. Later in life, the shells become strongly 
distorted, the ventral margin is abruptly bent downwards nearly 
at right angles to the former plane of the shell as is seen to a 
much less degree in’the recent C. contyvacta Say. Its surface is 


441 Costa RicA MIocENE—OLSSON 269 


coarsely concentrically grooved, the two valves being sculptured 
alike. 


Gatun Stage: Port Limon. 


Corbula hexacyma Brown and Pilsbry Plate 21, figures 12, 13 


Corbula hexacyma Brown and Pilsbry, 1912, Proc. Acad. Nat. Sci. 
Phila., vol. 64, p. 518, pl. 26, fig. 4. 


A species of the Gatun beds of the Central Zone. The shell 
is rather large, solid and with its valves similiary sculptured 
with a few, deep, wave-like, concentric ribs. Its right valve is 
slightly more convex than its left. Our specimens measure: 


Length 20, height 14, diameter of the left valve 6.00 
mm. 
Length 20, height 13.50, diameter of the rigth valve 
5.75 mm. 
Gatun Stage: Gatun, C. Z. 


Corbula orosi, n. sp. Plate 21, figures 9, 10 


Shell elongate, convex with the low inconspicuous beaks 
about the middle of the shell, so that the posterior and anterior 
ends of the shell are about equal in length; the right valve is 
slightly larger and its basal margin folds over slightly that of 
the left valve; anterior end rounded, the posterior narrow and 
more pointed but with the keel only very poorly developed; the 
surface is finely and evenly concentrically ribbed and both valves 
are similiarly sculptured; no lunule. 


Length 20, height 11.5, diameter of both valves 9.5 mm. 


A rather large species with nearly equally convex valves; 
surface sculptured with fine, regular, concentric ribs. It 
resembles the C. sarda Dall, from the Chipola Miocene but is 
much larger. 


Gatun Stage; Hill No. 3, Banana River. 


270 BULLETIN 39 442 


Section BOTHROCORBULA, Gabb 


Corbula viminea Guppy Plate 28, figure 25 


Corbula viminea Guppy, 1866, Quart. Journ. Geol. Soc. London, vol. 
22D A2Oa nla Zon iong Tere 

Lothrocorbula viminea Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15 
p. 247. 

Corbula ( Bothrocorbula) viminea Dall, 1898, Trans. Wagner Free Inst. 
Sci., vol. 3, pt. 4, p. 850. 

Corbula (Bothrocorbula) viminea Maury, 1917, Bull. Amer. Pal., vol. 
5, Pe 39754 Dl- 39) es. 20.) 2) 


This is a very characteristic species, distinguished by its 
large, deep lunule and its surface strongly sculptured with 
coarse, concentric, wave-like coste. It is known from the Mio- 
cene of Jamaica and Santo Domingo. 

The Corbula hexacyma Brown and Pilsbry, still known on- 
ly from the Gatun of the Canal Zone is similiar in form and 
sculpture but lacks the deep lunular pit. 

Length 15, height 10, semi-diameter 4.85 mm (right 


valve). 
Gatun Stage: Middle creck. 


443 


Fig. 


Costa RicA MiocENE—OLSSON 271 
EXPLANATION OF) RE AduES 
PLATE 1 (4) 
Page 
I. Terebra bipartita Sowerby, Saury Creek, height 77 mm........ 35 
2. Terebra dipartita Sowerby, Middle Creek, height 53 mm. 
3. Terebra cf. haitensis Dall, Sousi Creek, height 37 mm......... 35 
4. Terebra gatunensis Toula var., Gatun, height 58 mm............ 36 
5. Terebra gatunensis Toula, Gatun, height 52 mm. 
6. Terebra gatunensis Toula, Gatun, height 42 mm. 
7. Terebra spirifera Dall, var. midiensis, n. var., Middle Creek 39 
8. Terebra gausapata Brown and Pilsbry, Mt. Hope, height 
RUUWbe cba NARA MAAN RAO Hoa ra cab Woan Suan Kae AUABBO be 38 
9. Terebra estrellana, n. sp. Estrella River, height 7.5 mm...... 40 
Io. Terebra gausapata Brown and Pilsbry, Mt. Hope, height 
PXS) NOW oO peKeedeasccqdsooRaccQEHSeS SGouuacboaseBseupoasoogde 38 
11. Terebra Wolfgangi Toula, Mt. Hope, height 38 mm.............. 37 
12. Terebra Wolfgangi Toula, Mt. Hope, height 26 mm. 
iewlerebraspintiera Dall iBananapRaivetpecsecnee se seeeea eae a tees antes 37 
14. Conus gracilissimus Guppy, Bocas del Toro...... .........2.--0+55 52 
I5. Conus tortuosostriatus Toula, Bocas del Toro..................-...- 49 
ROwMMverebra estrellanay nm.Wsp', Estrellay Rivet nes. yal meee ain aaans ate 4o 
17-18. Terebra Berlinerze Maury, Boucary Creek.. ........ PE Asse A 39 
19. Terebra costaricensis, n. sp., Banana River, height 13 mm... 40 
20.) (Conus marginatus Sowerby, Red Clift Creel nee eee seo eee ee 51 
Zig COnuUS mW tS triatusVvBOSe Water Calycsauneeseeceesensyeescee cece 54 
2249) COMMS MUScensis. ns Spy, Bamana Ravietesece amy cenchecseeeaeeeestece as 47 
22a Conus muUltistriatus BOSE Water) Caine teases seecceee ee eeeee 54 
DARN COMMS HMUSZENSIS) My Spy. bata WRU CT enseetaeeneseeceacer ete cee 47 


25- 26. Terebra benthalis Dall, var. bocasensis, n. var., Bocas 


Daf 


Caled Wid oy oy MAU ian NE EA a aR URE 4I 


B72 


Fig. 


Fig. 


Fig. 


BULLETIN 39 444 
PLATE 2 (5) 

Page 

I-2. Conus molis Brown and Pilsbry, Banana River............ ..... 42 
3-40) /Conus proteus) /Euwassi) NViaitern © aiyesie. eats arene ane eeeane 43 
5. ConusiViea teh ty yaknspy MMW eit ergs Cay euch aera Ue eae ie Unc nts a manuae 44 
6. Conus imitator Brown and Pilsbry, Mt. Hope....... .....-.....00- 45 
Yo \ Orbis) WdosroreraIsS,) sole Gio; | IROIKO (Cals ccssopicnodiccscddcesudcoo eeoBobeuoade 48 
8), Conus Veatch misma a NVave rth @aly aula suse ease menene 8) 0 icsaee 44 
93) Conus! recognitus) Guppy. Banana Rivenroar eee ue .cee eee 46 

PACD) 2400 6)) 

jig) (CONS Je levewisi, tol, Siow. IBkewatebnel IRIE. shocc wssosoneeaes! \aune ;cbosodosooce 53 
2. Conus|stenostomus) Sowerbys) Bone Mu im Oneeeee senna. oe see ene 42 
3. Conus floridanus Gabb, var. costaricensis, n. var Banana R... 45 
AE: Cloyohntsy JeyouRel'd akahqobbl JBxoRS. | NWE\dese (OAy7 155 sooosodcosucsoocseascocab0SazC 52 
6))) Conus tortuosopunctatus Moulaw Mit. elope. ee a ease ene eee 54 
7freys) (Clovonbis}, IhhooVoyoVeraisyisy 014 /G}o},4) Jetorae ILpbooVha aqcoopocadécoosdocbndebocaods AT 
g. Conus floridanus Gabb, var. costaricensis, n. var., Banana R. 45 
Lov Conus planilinatus) Sowerby MeOnts lei O mts tren seaee eae tee ene 50 
Tt.) \Conustortuosopunctatus)Moulav Mth Hiopersice-csssseencceeeeeees 54 
12. Conus cf. interstinctus Guppy, Banana Rivetr..................0.00+ 48 
13.) Conus planiliratus) Sowerby, Port, Wimonm sc: se4 ee sseeee cece 50 
14. Conus planiliratus, var. bocasensis, n. var., Bocas del Toro.. 57 
152), Conus pranozonatus) Guppy, Rio) Blanco ws wesse.sseeneeceee ese eeae 50 


16. Conus planiliratus var., bocasensis, n. var., Bocas del Toro.. 51 


PLATE 4. (7) 


1-2, Turris albida Perry, slightly reduced, Toro Cays............ 58 
3. Scobinella Morierei (Laville) Cossmann, length 48 mm, Toro 
CaS eae Ee awa aaa sistent eet tees 79 
4. Scobinella Morierei (Laville) Cossmann, length 52 mm., 
Moro) Cay sys oseaucnweaeecece sete 79 
5. Drillia papaya, n.sp., length 32 mm., Rio Cocles............... 63 
6. Turricula lavinoides, n. sp., length 54 mm., Banana River.... 55 


Halia americana, n.sp., length 66 mm., Mt. Hope............... 79 


445 


Fig. 


CORSON CME ee Oo UE 


22. 


23. 


Costa RicA Mr1ocENE—OLSSON 273 

Page 

Drillia consors Sowerby, Grape Point Creek, length 60mm... 61 
Drillia venusta Sowerby, Banana River, length 30 mm........ 61 
Drillia consors Sowerby, Mt. Hope, length 33 mm......... ..... 64 


Drillia carruca, n.sp., East Grape Point Creek, length 18mm. 71! 
Turricula lavinoides, var. limonensis, n. var., length 97 mm., 


MOrOy Cary Sey CMe aati castle 56 
Drillia consors Sowerby, length 4o mm, Banana River......... 61 
Bullaria paupercula Sowerby, Middle Creek, length22 mm... 34c 
Drillia limonica, n. sp., Port Limon, length 19 mm.............. 66 
Ancistrosyrinx Dalli, n. sp., Toro Cays, length 14 mm......... 60 


Ancistrosyrinx elegans Dall, var., Banana R.,length 15 mm _ 60 


PLATE 5. (8) 


Drillia Cristobali, n. sp., Bocas del Toro, heigth 20 mm........ 72 
Drillia chirquiensis, n. sp., Bocas del Toro, height 25 mm.. 64 
Turricula taurina, n. sp., Bocas del Toro, height 25 mm........ 57 
Turricula Terryi, n.sp., Bocas del Toro, height 23.5 mm..... 57 
Drillia bocasensis, n.sp., Bocas del Toro, height 22 mm..... 73 
Drillia bocatoroensis, n. sp., Bocas del Toro, height 23 mm.. 67 
Driliiarcoccos,s 1) Spis)\Cocovelum\sheigiae 2mm) sees ecee se. 64 
Drillia limonensis, n. sp., Port Limon, height 20 mm......... 71 
Drillia limonensis, n. sp., height 15 mm. 

Drillia limonetta, n. sp., Port Limon, height 12.25 mm......... 70 
Drillia theobroma, n. sp., Bocas del Tcro, height 21 mm...... 68 
Drilliacitriat nusp..\/Lort Wimontohieleipyien santa sense erste 65 
Drillia citria, n. sp., Port Limon, height 11 mm. 

Drillia cocosina, n. sp., Bocas del Toro, height 12 mm........ 75 
Drillia aurantia, n. sp., Port Limon, height 11 mm..... ...... 68 
Drillia aquanica, n. sp., Water Cay, height mm.................. 65 


Drillia aquanica var. 
Drillia lithocolletoides, n. sp., height 10.75 mm., 


IBXOY CRIS GIS NoidO\an nds neu ausecosendhesoqsseads 74 
Leucosyrinx chloris, n. sp., height 14.50 mm., 
BocasrdelgMorou tee mean wasn uae! 69 
Cytharella limata, n. sp., height 9 mm, Port Limon.............. Ta 
Cythara terminula Dall, var. costaricensis n. var., height 13 
mm., Hill 1a, Banana River...... 77 
Cythara terminula Dall, var. costaricensis n. var., height 13 
mm., Hill ra, Banana River......... 77 
Borsonia cocoensis, n. sp., Coco Plum, height 16 mm............ 78 


2 Woe G74 BULLETIN 39 446 


Page 

Fig. 24. Borsonia cocoensis, n.sp., Coco Plum, 14 mMl.........-.....2006 78 

25. Drillia aquzensis, n. sp., Water Cay, height 15 mm.............. 73 

26. Drillia musa, n.sp., Banana River, height 16 mm............... 74 

27-28. Drillia musacina, n.sp., Banana River, height 9 mm ..... 69 
29-30. Glyphostoma moinica, n. sp., Port Limon, height, 

Eni}: 0dbaDY, ond Gon dooasopaseeaoneedoRonooDseagce 76 


Fig. 


PANE) 361199) 


I. Mitra Swainsoni Broderip, var. limonensis, n.sp., Port 


IWibesvoyay,  oKeRKelole * /Al: TaBWEO,. «Ass nogaaeaqocdodse 16606 100 
2-3. Cancellaria Plummeri, n.sp., Toro Cays, height 34 mm... 85 
4. Cancellaria toroeusis, n.sp., Toro Cays, height 29 mm ....... 84 

5. Cancellaria Maury, n. sp., Cercado de Mao, Santo Domingo, 
JaKesGegalty /GX6), SOUTH ag aenoonvosadoogaosodccoogs oooaECdosS 82 
6. Cancellaria Barretti Guppy, slightly enlarged, Banana River. 81 
7. Cancellaria Rowelli Dall, East Grape Point Cr., height 30mm. 84 
8. Cancellaria dariena Toula, about natural size, Toro Cays........ 80 
g. Cancellaria Cossmanni, n. sp., somewhat enlarged, Banana R. 81 
Lo.) Matra longa Cabby iGatun) heiglit) oy mimeo se pasccecnacssacseees IOI 
11. Cancellaria Cossmanni, n. sp., somewhat enlarged, Banana R_ 81 
12. Cancellaria islacolonis Maury, enlarged, Red Cliff-Creek........ 86 
13. Mitra cf. rudis Gabb, Hone Walk Creek, height 33 mm.......... 102 
14-15. Marginella MacDonaldi Dall, Banana R., height 28. mm... 95 
16. Marginella mindiensis Cossmann, Gatun, height 9 mm.......... 96 
17. Marginella MacDonaldi Dall, Banana R., height 26 mm......... 95 

18. Mitra Almagrensis Toula, variety coralliophila, n. var., Port 
Limon, height 17 mm.......... 103 
19. Marginella MacDonaldi Dall, Banana River, height 25 mm.... 95 
20-21. Mitra poas, n. sp., about natural size, Port Limon.............0. 102 
22. Marginella leander Brown and Pilsbry, enlarged, Gatun........ 98 
23. Marginella mindiensis Crossmann, Gatun, height 17.5 mm...... 96 

24. Mitra Almagrensis Toula, variety coralliophila, n. var., Port 
Limon, height about 17 mm........ 103 

25. Mitra dariensis Brown and Pilsbry, Bocas del Toro, height 
2) G5 0008 ARB HAH ER aE i Sore OF ese asdabacée IOI 


I. Oliva cylindrica Sowerby, Old Man Sam Creek.......... Hocesclnae 88 
2-3. Oliva brevispira Gabb, Red Cliff Creek.......... awa aciaaas Bhansee 88 


447 


Fig. 


Fig. 


Costa RicA MIocENE---OLSSON 275 

Page 

4. Oliva brevispira Gabb, var. Giraudi Cossmann, Middle Creek. 88 

Sm Olivarcatinensis: Loulaw Gatun eesccecn ecco oe een ccretanseceeee 89 
6-7. Oliva Sayana Ravenel, var. immortua Pilsbry and Brown 

BananaiRiverncnsewudevossescieeeeccet 89 

8-9. Oliva mancinella, n. sp., East Grape Point Creek......... sesenT PO) 

LOmeeAnelllariajaqiuzensis, ni/Spi Water Cay.iueeesesacesseneseerese sce ce 94 

11 Olivella muticoides Gabb, East Grape Point Creek................ 92 
12-13. Oliva testacea Lamarck, var. costaricensis, n. var., Banana 

RAVE TA eee ANU ARLE) SU aL 90 

12. Olivella muticoides Gabb, Red Cliff Creek... .............. ceceeeee ee, 92 

15-16. Olivella Boussaci Cossmann, var., Banana River............... 93 

17-18. Olivella muticoides Gabb. Red Cliff Creek. ..................5. 4 92 

19-20. Olivella limonensis, n. sp,, Port Limon.................60eeseeeee 93 

21. Marginella avena Valenciennis, Banana River ..................0+- 97 

pom Olivelia Goliath, im. Sp Red Ciith\ Greeks usec sesseseseeencee- 92 


23. Olivella Goliath, n. sp., Banana River 
24-25. Olivella limonensis, var., bocasensis n. var., Bocas del 


TOTO Te Se Mane MN e Ua EU OMaLEE UIs Lue IU 94 
26-27. Marginella collina, n. sp., Banana River................0000eeeeees 97 
28. Marginella avena Valenciennes, Port Limon.....................-:. 97 


PLATE 8. (If) 


1. Fasciolaria MacDonaldi, n. sp., Banana River, height 70 mm. 106 
2. Voluta Alfaroi Dall, Banana River, height 63 mm............... 99 
3. Solenosteira chiriquiensis, n. sp., Water Cay, reduced........... 114 
4. lLatirus taurus, n. sp., Toro Cay, slightly reduced................. 108 
5. Fusus miocosmius, n. sp., Bocas del Toro, height 100 mm..... 103 
6. Fusus honensis, n. sp., Hone Walk Creek, height 63 mm ..... 104 
7. Solenosteira Vaughani Dall, variety medioamericana, n. var. 
Toro Cary Sis aera neal iar un) iain 2 
8. Solensoteira Dalli Brown and Pilsbry, Gatun, slightly en- 
langed seni i Men at wataae cr 113 
g. Fasciolaria Gorgasiana Brown and Pilsbry, Rio Betey, height 
Ko Vb cab oe aM anUON NAW enue eto tata UML Ae nag 105 
10. Latirus infundibulum Gmelin, Pumbri Creek, slightly re- 
GUC ed eset ee ued GOS en Ns 107 
11. Peristernia insula, n. sp., Water Cay, height 23 mm............ IIO 
12. lLatirus irazu, n. sp., Port Limon, height slightly reduced..... 107 


13. Peristernia tortugera, n. sp., Port Limon, height 29 mm....... 110 


BULLETIN 39 448 


PLATE 9. (12) 


Page 


Melongena consors Sowerby, Old Man Sam Creek, height 
TEINS) (SBODBHIS Gaba0G54 Hansa pup oabodoonbEedodcdec5 112 

Phos Veatchi, n. sp., Hill 1a, Banana River, height 43 mm.. 121 

Phos Veatchi, n. sp., Hill 1a, Banana River, height 41 mm.. 121 


. Phos gatunensis Toula, Gatun, height 30 mm................-..5. Mie) 
Phos beteyensis, n. sp., Rio Betey, height 26 mmu................. 122 
Northia northiz Gray, variety miocenica, n.var., tip, Banana 

ISOMEIEY Sretondessancos sooBdOuSeRABRposacHodona 124 

. Phos Moorei Guppy, variety costaricensis, n. var., Banana 

RVs Jolemelate, 2X6) weoknalsesoaqeac GudonoodC 118 
Phos mexicanus Bose, Water Cay, height 29 mm...... ........5. 117 


Phos mexicanus Bose, Water Cay, height 27.5 mm. 

Phos elegans Guppy, variety limonensis, n. var., Port Limon 
leraerel nl 237/, TaabIal se ccosddondooaboooosds eo00e 119 

Phos elegans Guppy, variety limonensis, n. var., Port Limon 
length 22.50 mm 

Northia northie Gray, variety muocenica, n. var., Banana 


VET Sh el pac Gu ASEM ROn Rae HaRAA MR nen aie aio 124 
Phos snbsemicostata Brown and Pilsbry, Saury Creek, height 

AD) MGSO oa seoeadce BBen Aan nee sone aenboe se 121 
Phos metuloides Dall, Mt. Hope, height 19.5 mm............... 122 
Phos estrellensis, n. sp., Estrella River, height I9 mm......... 120 


Phos estrellensis, n. sp., Estrella River, height 16 mm. 
Alectrion merenda, n. sp., Old Man Sam Creek, height 7 mm 124 


Alectrion ranuncula, n. sp., Banana River, height 14 mm..... 123 
Alectrion losquemadica Maury, Port Limon, height 11.50mm 123 
Alectrion ranuncula, n. sp., Banana River, height 14 mm...... 123 


PLATE 10 (13) 


Marginella latissima, variety Pilsbryi, n. var., Banana R...... 96 

Marginella latissima, variety Pilisbryi, n. var., Gatun 

. Clava costaricana, n. sp., Banana River, height 40 mm........ 144 

Metula Harrisi, variety limonensis, n. var.. Port Limon, 
InlSsheloly) 7/ Swaheolyonane? Shouse soqsobacdosedco 116 

Metula Harrisi, variety limonensis, n. var., Port Limon, 
eto hey 25 TAT esescs pennant 116 

Strombina matima, n. sp., Banana River, height 12 mm........ 127 


449 Costa RicA M1ocENE—OLSSON 277 


8. Strombina costaricensis, n. sp., Middle Creek, height 15 mm 128 


g. Strombina ambigua Guppy, Banana River, height 26 mm...... 126 
to. Metula Harrisi, n. sp., Coco Plum, height 27 mm................. 116 
11. Metula Gabbi Brown and Pilsbry, Mt. Hope, height 28.5 mim 115 
12. Metula cancellata Gabb, Bocas del Toro, height 18.5 mm...... 115 
13. Strombina Lessepiana Brown and Pilsbry, Gatun, height 

AS Sanvad Landay. aude jnppaauAaoooebadbaaonoca 126 

AMe SECO bia Chiniguiensisy maiSPilucamatnnene re acee ce ee eneeee oie eecesns 130 
15. Typhis alatus Sowerby, Toro Cays, height 23 mm................- 132 
16-17. Strombina costaricensis, n. sp., Middle Creek...............0. 128 
18. Strombina cricamola, n. sp., Water Cay, height 16 mm......... 127 
19. Strombina sincola, n. sp., Banana River, height 7 mm.......... 129 
20. Strombina Lessepiana Brown and Pilsbry, Gatun................. 126 
12. Strombina sineola, n. sp., Banana River, height 7 mm......... 129 
22. Typhis linguliferus, variety costaricensis, n. var., Bannan R. 132 
22y eoistrum) nodulosum ©. Bs Adams, Red Cliff) Creeks co .5.5-. 133 
PAM StLOm bia Chiriquiensiss das )Spi isan uen eel oe wasulatesleclsicele sels salle 130 
25. Marginella musacina, n. sp., Banana RiveT........ 0.0.0.0... se000 98 


26-27. Strombina costaricensis, variety musanica n. var. Banana R 129 
28. Cerithium costaricensis, n. sp., Red Cliff Creek, height 


PX oii DOOD AA ANUN GRRE AA BANE aaa SeRaeReToe see ae 143 
29. Typhis linguliferus, variety costaricensis, n. var., Banana R. 132 
30. Clava alajuela, n sp., Middle Creek, height 17 mm............. IA5 
31-32. Drillia estrellana, n. sp., Estrella River, height 7.50 mim... 69 
33-34. Columbella submercatoria, n. sp., Red Cliff Creek.............. 125 


PLATE 11 (14) 


Fig. 3. Xancus scopulus, n. sp., Banana River, height 265 mm......... TIt 


PLATE 12 ({5) 


Big: 1. Serpulorbis papulosa Guppy, Banana River........................-. 145 
2. Malea elliptica Pilsbry and Johnson, Banana River, height 
G3 STAY ee Ne SS et aera 139 
3. Sconsia levigata Sowerby, variety Gabbi, n. var., Toro Cays, 
cbvedondlyy, SrallahsegeC onda aouasdanopodad se: 136 
4. Siliquaria modesta Dall, variety limonensis, n. var., Port 
Limon, somewhat enlarged......... 147 


5. Siliquaria modesta Dall, variety limonensis, n. var., Port 
Limon, slightly enlarged............ 147 


278 


Fig, 


Fig. 


BULLETIN 39 450 
Page 
6. Siliquaria modesta Dall, variety limonensis, n. var., Port 
Limon, slightly enlarged............ 147 
7. Sconsia cocleana, n. sp., Rio Coles, height mm.................... 138 
8. Cypreea cinerea Gmelin, Rio Blanco.............0..cccccceesescereeese 140 
g. Cyprea almirantensis, n. sp., Water Cay, height 39 mm...... 140 
10, Cypreea parisimina, n. sp., Port Limon, height 39 mm......... 139 
11, Phalium moniliferum Guppy, Rio Betey, height 35 mm........ 135 
12. Sconsia bocasensis, n. sp., Bocas del Toro, height 35 mm...... 137 
13. Sconsia bocasensis, n. sp., Bocas del Toro. 
PLATE 13. (16) 
1-2-3. Strombus pugiloides Guppy, Banana River...................05- 142 
4. Fasciolaria tulipa Linnzeus, variety Port Limon................... 105 
5-6. Strombus gatunensis Toula, Water Cay, somewhat reduced.. 141 
7. Polinices Stanislas-Meunieri Maury, Banana River, slightly 
SRO NEKO los ngnocdoquosadesedddn sles 157 
8. Natica Youngi Maury, varietv cocleana, n. var., Cocles 
Creek, slightly enlarged............. 155 
ov Natica canrena Ieimmecisy Gatun eee see ee epee eeneee ee eee seeneee 155 
10-11-12. Architectonica granulata Lamarck, Banana River......... 154 
13-14-15. Natica Guppyana Toula, Banana River...............:06 see 156 
16-17. Polinices subclausa Sowerby, Banana River..................000+ 157 
PLATE 14. (17) 
i. Turritella oreodoxa, n. sp., Banana River, height 49 mm........ 152 
2. Turritella exoleta Linnzeus, variety limonensis, Port Limon, 
LOSS olle Ate Fools 4 Gescenugaopscasedoceoace 152 
3. Turritella exoleta Linnzeus, variety limonensis, n. var., Port 
Limon, height 36 mm 
4. Turritella altilira Conrad, variety chiriquiensis, n. var., 
Banana River, height 52 mm...... 150 
5. Turritella mimetes Brown and Pilsbry, Gatun, height 72 mm 149 
6. Turritella altilira Conrad, Mt. Hope, typical, height 50 mm 150 
7. Turritella altilira Conrad, Gatun, typical, height 36 mm...... 149 
8. Turritella altilira Conrad, variety chiriquiensis, n. var., Wat- 
er Cay, height 116 mm. ............. 150 
9. Turritella altilira Conrad, variety chiriquiensis, n. var. Ban- 


ana River, height 4o mm............ 150 


Fig. 


Costa Rica MioccENE—-OLSSON 279 


Page 


. Io. Petaloconchus sculpturatus H. C. Lea, Water Cay, height 


YAM oD OTB EBS RBA MACHO aa AN OR SAM SiN cane anr 
It. Turritella altilira Conrad, variety costaricensis, n. var., Sousi 
Greek Sheiolt QAi mms ease 
12, Turritella gatunensis Conrad, Water Cay, height 36.5 mm..... 
13. Turritella gatunensis Conrad, Water Cay, height 24 mm. 
14. Turritella altilira, variety chiriquiensis, n. var., Red Cliff 
Creeks iheiabtig aim malay eaeeeeseeeeees 
I5. Petaloconchus sculpturatus H. C. Lea, East Grape Point 
Creek Whleioliia a yirarreen ares see seen 


PVA TS len) 


1. Dentalium uscarianum, n. sp,, Rio Cocles, natural size......... 
2-3. Dentalium. bocasensis, n. sp., Bocas del Toro, somewhat 
enlarged. a eat NEO ANE I NE 
4. Astralium brevispinum Lamarck, variety basalis, n. var., Old 
Man Sam Creek, natural size...... 


5. Astralium brevispinum Lamarck, variety basalis, n. var., Old 
Man Sam Creek, natural size...... 

6-7. Calliostoma castilla, n. sp., Port Limon, slightly enlarged..... 

8. Calliostoma limonensis, n. sp., Port Limon, about natural 


STZ NaS MUU UT SOAK eae UE A ED 
g-10. Calliostoma mancinella, n. sp., Red Cliff Creek, about nat- 
WRAY SIZ SEN eR Ve uig acts N AUST Dba at 
tt. Calliostoma limonensis, n.sp., Port Limon, about natural 
SUZ OY UNM SD ANCA AT MEA PEED seat 


IMuboysaltuss n> sps, Red: Clit Creek height mmh ees) ee 
13. Scaphella costaricana, n. sp., Cocles Creek, height 29 mm..... 
14 Ptychosalpinx ? dentalis, n. sp., Cocles Creek, about natural 


15. Actzeon costaricensis, n. sp., Banana River, height 19.5 mm. 
16. Chlorostoma costaricensis, n. sp., Red Cliff Creek, diameter 


17. Crepitacella limonensis, n. sp., Port Limon, height 8 mm..... 
18. Ptychosalpinx ? dentalis, n. sp., Cocles Creek, about natural 


19. Bursa crassa Dillwyn, Banana River, about natural size........ 
20-21. Acteocina bullata Kiener, variety costaricana, n. var., Mid- 
die Créeks heiehtiropmmeses.scscr: 


150 


146 


166 


166 


Fig. 


Fig. 


is) 


BULLETIN 39 452° 


Page 
Neritina viridemaris Maury, Middle Creek, enlarged............ 159 
Alaba turrita Guppy, Estrella River, height 5 mm............... 153 


PLATE 16 (19) 


Pecten MacDonaldi, n. sp., type, Gatun (Toro limestone) 
right valve, height 96 mm.......... 198 
Pecten MacDonaldi, n. sp., cotype, same locality, left valve 
height 79 mm. 
Pecten gatunensis, Chagres Spillway, right valve, height 
COPA oTb Gols Aas nao ha MERE RE ARE Hone hada 197 
Pecten gatunensis, same locality, left valve, height 48 mm. 
Pecten levicostatus Toula, Hill No. 2. Banana River, slight- 


hf) sREYG NE CeL=YE Ae aon uaanoBooo sdonosdoed odor 199 
Pecten levicostatus Toula, same locality, left valve, slightly 
reduced 
Pecten levicostatus Toula, same locality, right valve slightly 
reduced 


PLATE 17 (20) 


Amusium luna Brown and Pilsbry, Chagres Spillway, some- 
Wiltatine duce disease esata en esc eee 205 

Pecten preglyptus, n. sp., Cocles Creek, natural size (type).. 202 

Amusium bocasensis, n.sp., Bocas del Toro, natural size, 


COLY POH ase dee sciseeeoneemeeessee eer 206 
Amusium bocasensis, n. sp., Bocas del Toro, interior of right 
valve. 
Pecten coralliphila, n.sp., type, Port Limon, slightly en- 
Pare nce ee CS ee Ue aa “t-99 
Pecten coralliphila, n. sp., Port Limon, left valve................ 199 


Pecten preglyptus, n.sp., interior view of type, natural size 202 
PLATE 18 (2i) 


Ostrea megodon Hanley, Banana River, natural size ........... 195 
Glycymeris canalis Brown and Pilsbry, Comadre Creek, about 
MatuTnalisize nesses wens sue 177 


453 Costa Rica MrockxE—OLSSON 281 


Page 

3. Glycymeris canalis Brown and Pilsbry, Banana River, about 
pacing INH Aer ANE AMR Ee At. Atta acts Neon 177 

4. Glycymeris canalis Brown and Pilsbry, variety, East Grape 
POM Cy Cree ke aN a eye aa Nofy 

5. Glycymeris canalis Brown and Pilsbry, variety, East Grape 
Pointy Greeley eee uu yy veakne enna tues Mola) 
6-7. Glycymeris canalis Brown and Pilsbry, Banana River......... 177 
Pecten costaricensis,n. sp., Red Cliff Creek, right valve........ 201 


g. Pecten costaricensis, n. sp., Red Cliff Creek, left valve. 
Io. Pecten costaricensis, n. sp., Red Cliff Creek, right valve. 
Ir. Pecten costaricensis, n. sp., Red Cliff Creek, left valve. 
12. Leda ensinoides, n. sp., Banana River, length 54 mm........... 174 
13. Glycymeris castaneus Lamarck, Banana River,length 30 mm. 180 
14. Glycymeris castaneus Lamarck, Banana River,length 23 mm. 

15. Leda Balboz Brown and Pilsbry, Banana River, enlarged..... 173 
16. Pecten almirantensis, n. sp., Bocas del Toro,heigth 4.75 mm. 203 


IPppecten aOLus, ni sp: Port Limon, height 4i5o,mam.-.34) +... 204 
18. Pecten aotus, n. sp., Port Limon, 4.75 mm. 

19, Leda quitanensis, n. sp., Quitana Creek, length 26 mm........ 174 
20uWeda Davila, m.\Sp.) Bocas deli Moros lenges 5-75) tate... sese-2 7a 
21-22-23-24. Nucula cahuitensis, n. sp., Saury, length 3.50 mm..... 171 


25. Nucula tenuisculpta Gabb, Red Cliff Creek,length 4.00 mm. 172 
PLAGE Loy (22) 


‘Fig. 1. Glycymeris carbasina Brown and Pilsbry, East Grape Point 


Creek,aboutimatunalllsizensseisce sce 179 
2. Glycymeris carbasina Brown and Pilsbry, same locality, 
AOUb Ta tually Siz Cleese enna ease 179 


3-4. Glycymeris jamaicensis, Red Cliff Creek, about natural size 178 
5. Glycymeris carbasina Brown and Pilsbry, East Grape Point 
Creek, small shell about natural 


SUZ Gs ameter eer eA near Me ERLE NAM IUD ERO Uy 179 
6-7. Glycymeris decussatus Lamarck, Old Man Sam Creek, nat- 
Trad SIZE WAN saiaranees UNA Aen Aen re 180 


PLATE 20 (23) 


Fig. 1. Spondylus chiriquiensis, n. sp., Water Cay, height 151 mm... 206 
2. Spondylus chiriquiensis, n. sp., young shell to show the small 
foliaceous attachment area. 
3. Thracia Tristani, n. sp., Rio Betey, length 42 mmi............... 211 


282 


Fig. 


Fig. 


nun 


Io. 


BULLETIN 39 454 


Page 


Crassatellites Reevei Gabb, Banana River, slightly reduced.. 212 
Spondylus chiriquiensis, n. sp., Water Cay, young right 


Spondylus chiriquiensis, n. sp., Water Cay, young left valve. 
PLATE 21 (24) 


Spondylus gumanomocon Brown and Pilsbry, Hone Walk 
Creek, shell greatly reduced....... 207 
Ostrea costaricensis, n. sp., Rio Blanco, about natural size... 196 
Placunanomia lithobleta Dall, Banana River, height 36 mm.. 210 
Placunanomia lithobleta Dall, Banana River, height 49 mm 
Placunanomia lithobleta Dall, interior of same specimen as 
figure 3. 
Anomia simplex d’Orbigny, Middle Creek, height 34 mm..... 209 
Corbula gatunensis Toula, Gatun, length 21 mm. 
Corbula Dietziana C. B. Adams,Port Limon, length 15.5 mm. 268 
Corbula orosi, n. sp., type, Banana River, length 19.5 mm... 269 
Corbula orosi, n. s9., type, Banana River, length tg mm. 
Labiosa lineata Say, Saury Creek, length 44 mm................ 264 
Corbula hexacyma Brown and Pilsbry, Gatun,length 26 mm. 269 
Corbula hexacyma Brown and Pilsbry, Gatun,length 18 mm. 


PLATE 22 (25) 


Arca occidentalis Phil., Port Limon, somewhat enlarged...... 181 
Arca umbonata Lamarck, East Grape Point Creek, slightly 
enlarged eee sy ree ee sae 182 


Arca auriculata Lamarck,,Old Man Sam Creek, young shell.. 190 
Arca Mauryze, n. sp., Port Limon, right valve of type, length 


Arca candida Gmelin, Port Limon, left valve, length 18 mm. 183 
Arca candida Gmelin, Port Limon, right valve length 


13 mm. 

Arca\Mauryce. (ns Spr ittenon Wlewy Onity Pea eene se eel-ieettanseee 182 

Arca honensis, n. sp., Hone Walk Creek, left valve type, 
slighitlys;nediiced wus teeesencee cee 187 

Arca honensis, n. sp., same locality, right valve of cotype, 
Sligdhtlyasne diced etn naieeheruecnescaee 187 


Arca dariensis Brown and Pilsbry, Water Cay, about natural 
SIZ ORME a Re A ICR NR tat 184 


Fig. 


Fig. 


Fig. 


1a & 


Pao 
3. 


Costa RicA MrockENE—OLSSON 283 


Page 


Arca dariensis Brown and Pilsbry, Water Cay, about natural 


Arca dariensis Brown and Pilsbry, Gatun, about natural size. 
Arca dariensis Brown and Pilsbry, Gatun, about natural size. 


PLATE 23 (26) 


Arca Veatchi, n.sp., Water Cay, left valve of type, length 

GMAT Gata ecine sesame cionainieigs eel neige 189 
Arca Veatchi, n. sp., Water Cay, interior of left valve of type. 
Arca Veatchi, n. sp., Water Cay, right valve of type. 


4-5. Arca golfoyaquensis Maury, variety medioamericana,n. var., 


6. 


Hone Walk Creek, valve of type, 
Ikesaverdoy’ Gey Semboouogousaqnesodaqomabadeosde 188 

Arca golfoyaquensis Maury, variety medioamericana, n. var., 
Hone Walk Creek, left valve of 
type, same locality. 


7-8. Arca actinophora Dall, Red Cliff Creek, nearly natural size.. 185 


I 
De 
Be 
6. 
7 


Xe 


14. 


aes 


PLATE 24. (27) 


Arca cacica, n. sp., type, Comadre Creek, length 36 mm...... 190 
Arca Pittiern Dall Sauny) Creek mabunalisizessnuniecsssess secre ces 192 


4-5. Arca-Pittieri Dall, Hill 1a, Banana River, natural size. 


Arca Pittieri Dall, Hill 1, Banana River, natural size. 

Arca Hindsi, n. sp., type, East Grape Point Creek, natural 
RIYAS ei socsanerdodasnimacasods sdscoosoosesdeE 193 

Arca Hindsi, n. sp., co-type, Rio Betey, natural size. 

Arca Hindsi, n. sp., co-type, Hill-3,.Banana River, natural 
size. 

Arca Lloydi, n. sp., co-type, Red Cliff Creek, length 18 mm.. 192 

Arca Lloydi, n. sp., type, same locality, diameter 18 mm. 

Arca Lloydi, n. sp., co-type, same locality, length 15.5 mm, 

Arca Henekeni Maury, Water Cay, length 30 mm................ 186 

Arca Henekeni Maury, Water Cay, length 28 mm. 


PLATE 25 (28) 


Arca costaricensis, n. sp., type, Hill 1a, Banana River, nat- 
TRA SIZE Peet ede cme atigauigunecnar tas 186 


284 


Fig. 


nn 


10. 


is) 


Lal 
OR Or Oe ON Oates 2 


20106 


2s 


13. 
14. 


BULLETIN 39 456 


Page 
Arca costaricensis, n. sp., co-type, Hill 1a, Banana River nat- 
ural size. 
Arca actinophora Dall, typical form, Mt. Hope, slightly re- 
GUC CU ree Sak Shee aati eeaatate 185 


Arca MacDonaldi Dall, Hill No. 3, Banana River, nearly nat- 
ural size, interior view to show 
ihighWicardinalltancateeers sac sesesacs 

Arca MacDonaldi Dall, exterior of the same specimen. 

Arca MacDonaldi Dall, variety subreversa, n. var., Rio Betey 
exterior of left valve. 

Arca MacDonaldi Dall, variety subreversa n. var., Hill No. 3 
Banana River, interior of right 
valve, to show form and low card- 
inal area. 

Glycymeris Lloydsmithi Pilsbry and Brown, Red Cliff Creek, 
right valve, slightly reduced....... 

Glycymeris Lloydsmithi Pilsbry and Brown, same locality 
left valve. 

Glycymeris Lloydsmithi Pilsbry and Brown, same locality 
right valve. 


PLATE 26 (29) 
Tellidora cristata Recluz, Hill 3, Banana River, nearly nat- 


MAGA STZ Oars a SE Sie a terest 
Tellidora cristata Recluz, Hill 1a, Banana River, nearly nat- 
ural size. 
Tellina dariena Conrad, Water Cay, natural size.................- 


Tellina lepidota Dall, Water Cay, length 24 mm.................. 
Tellina lepidota Dall, Water Cay, length 21 mm. 

Tellina costaricana, n. sp., Banana River, about natural size 
Tellina rovala, n. sp., Water Cay, slightly enlarged............. 
Tellina rovala, n. sp., Water Cay, (type) slightly enlarged. 
Tellina costaricana, n. sp., Banana River, about natural size 
Tellina crystallina Wood, Banana River, enlarged nearly 


Macoma panamensis Dall, variety canalis n. var., Mt. Hope, 
eroverdol inky Senhiol sm hoocbondeeKd6) | coSbo00 
Tellina umbra Dall, East Grape Point Creek, enlarged about 
Bi abst) ocounsqoeegeocnubboouesHonDb05d0 
Macoma gatunensis Toula, Banana River, about natural size 
Macoma costaricana, n. sp., Rio Betey, about natural size..... 


194 


181 


251 
249 
257 
253 


257 
256 


457 Costa Rica MiIockNE—OLSSON 285 


Page 

15. Tellina umbra Dall, East Grape Point Creek, enlarged about 
BWA OBITS a Hagt eacsacoadsogoGce soud05 253 
16. Macoma costaricana, n. sp., Banana River, about natural size 256 
17. Tellina nasua, n. sp., Red Cliff Creek, about natural size...... 250 
18. Strigilla musanica, n. sp., Banana River, length 13 mm....... 255 

Ig-20. Venericardia Conradiana Gabb, variety limonensis, n. var., 
Portueinions leneth) 7mm)ys....senie 217 

21. Chione grus Holmes, variety, Red Cliff Creek, enlarged about 
AlGiTMesSeaaeiews: PAIGE A) ta ASIA Ban Ar SR 248 


22. Cardita caribbeanensis, n sp., Banana River, length 18 mm. 214 
23-24. Limopsis monilis, n. sp,, Old Man Sam Creek, length 1.75 
ED FT oe daca dtonecate s6obh bene ucnvbpocadeapeade: 177 


PLATE 27 (30) 


Fig. 1. Cardium spinosum Meuschen, var. Turtoni Dall, Port Limon 


aboutmatunallisizes nies aeeceh seieicece 228 
2. Cardium dominicense Gabb, East Grape Point Creek, slight- 

hi? SEC IWKOETE cocagcosatocdeogeqoneoosD jo6o0H 225 
3. Cardium costaricauum, n.sp., East Grape Point Creek height 

DIAN) CON en ANKE Soc S A SRO AAG rosa AHAB BABEe OnE 226 
4-5. Cardium stiriatum Brown and Pilsbry, Banana River, slight- 

Ny SSO BLES | so od bdomnododoosy sescaspanodcedo 225 
6. Cardium medium Linné, Port Limon, about natural size....... 226 
7- Protocardia gurabica Maury, Gatun, length 29 mm....... bedbesor 229 


8. Protocardica gurabica Maury, Water Cay, length 25 mm. 
g. Protocardia gurabica Maury, Water Cay, diameter 19 mm. 


1o. Cardium heredium, n. sp., Rio Betey, length 11.50 mm........ 227 

11-12. Cardium serratum Linné, Middle Creek, natural size......... 228 

13. Protocardia costaricensis, n. sp., Comadre Creek, length 21 
PADDED 3S dE MS SHIN AE UUM RNase 229 


PLATE 28 (31) 


Fig. 1. Mactra estrellana, n. sp, Estrella River, length 50 mm......... 263, 
2. Mactra exoleta Gray, Boucary Creek, about natural size....... 262 
3. Mactra alata Spengler, Banana River, about natural size...... 263 
4. Mactra sincola, n. sp., Saury Creek, length 36 mm .............. 264 
5. Echinochama yaquensis Maury, Old Harbor.............-..00:0000 219 


286 


Fig. 


19. 


20. 


BULLETIN 39 458 


Page 


Plicatula marginata Say, Water Cay, the lower or attached 
valve, enlarged about 2% time.... 
Plicatula marginata Say, Water Cay, the lower or attached 
valve, enlarged about 2% times. 
Hchinochama antiquata all R1o Blanco sececea-c-5 hoe eee 
Pteria inornata Gabb, Red Cliff Creek, natural size. ........... 
Limzea solida Dall, Red Cliff Creek, height 2.5 mm.............. 
Chama congregata Conrad, Red Cliff Creek, enlarged about 
GwiGeyaeves ibddougueaauouanpeHAsDsusooaoDdad 
Corbula oropendula, n. sp., Banana River, length 6.5 mm..... 
Corbnla oropendula, n.sp., Banana River, same valve but 
less magnified, length 6.5 mm 
Corbula oropendula, n. sp., same locality, right valve, 6.5 


mm. 

Corbula cf. vieta Guppy, Banana River, right vaive, length 
(OPA hoohes We Man ec AemaseaisnaaascasbooC0 

Corbula cf. vieta Guppy, Banana River, left valve, length 
6.25 mm. 


Leda Dalliana, n. sp., Port Limon, length 10.50 mm............. 
Mesodesma costaricensis, n. sp., Banana River, length 22 


Nucula orbicella, n. sp., Port Limon, length 9.5 mm............ 
Nucula orbicella, n. sp., Port Limon, length 11 mm. 


21-22. Yoldia bocasensis, n. sp., Bocas del Toro, length 9.5 mm... 


23. Mulinia lateralis Say, variety sauryensis, n. var., Saury 
Creek, about natural size............ 

24. Mulinia lateralis Say, variety sauryensis, n. var., Saury 
Creek, about natural size. 

25. Corbula viminea Guppy, Middle Creek, length 15 mm......... 
26-27. Corbula oropendula, n. sp., Saury Creek, length 9 mm...... 
PLATE 29 (32) 

1. Semele levis Sowerby, variety costaricensis, n. var., Hill 3, 

Banana River, length 72 mm...... 
2. Semele Claytoni Maury, Hotel Creek, natural size.............. 
3. Metis barba, n.sp., Rio Betey (type), right valve, length 
AS ier Th ea Oe ee eee OA Severe 
4. Metis barba, n. sp., Rio Betey (type), left valve, length 


48 mm. 


208 


208 
218 
195 
209 


218 
267 


266 


270 
267 


258 
259 


255 


Fig. 


Fig. 


to. 
Tes 


PIA Re SH 


ae 


© 


Costa RicA MrocENE—OLSSON 287 


Page 


Sanguinolaria alouatta, n.sp., Banana River, right valve, 
Neraverelay (eyo) seobee A secan doroAdicaechoda eeauA 260 
Sanguinolaria alouatta, n. sp., Banana River, left valve, 
length 54 mm. 
Psammosolen gatunensis Toula, variety costaricensis, n. var. 


Rio) Betev, length 63) mimis5. 5.2... .. 260 
Semele Sayi Toula, Sousi Creek, slightly enlarged .............. 258 
Tagelus cebus, n. sp., Rio Betey, length 36 mm.................. 261 
Abra triangulata Dall, Bocas del Toro, length 7.25 mm........ 259 
Crassatellites midiensis, n.sp., Middle Creek, length 3.25 

TTT Aunt setleciee eal an cea ace ma 213 
Crassatellites bowdenensis Dall, variety costaricensis, n. var. 

IKsiakestel at, eh. Ryo) saubaol, codssacon.conbannnoanaoae 21g} 


PLATE 30. (33) 


Antigona multicostata Sowerby, Water Cay.......-....c.c.ceeseeees 239 
Chione Rowleei, n. sp., Banana River, length 42 mm.......... 244 
Antigona Harrisiana, n. sp., Port Limon, length 21.5 mm... 241 
Antigona rugosa Gmelin, Port Limon, about % natural size 240 
Antigona Blandiana Guppy, Comadre Creek, length 30 mm.. 241 
Antigona Blandiana Guppy, Port Lomon, length 24 mm. 
Chione mactropsis;Conrad, Gatun, length 30 mm................. 245 
Chione mactropsis Conrad, Sousi Creek, length 51 mm. 
Chione holocyma Brown and Pilsbry, Gatun, length 36.5 

PTL CIN EO UA eS acetal 247 
Antigona tarquina Dall, Saury Creek, about natural size...... 240 


PLATE 31 (34) 


Dosinia acetabulum Conrad, Rio Betey, somewhat reduced... 231 
Cyclinella beteyesis, n. sp., Rio Betey, height 39 mm........... 242 
Pitaria circinata Born, Saury Creek, about natural size......... 236 
Clementia dariena Conrad, Sousi Creek, height 60 mm ...... 232 
Pitaria Hillii Dall, variety musanica, n. var., Banana River, 
Wesalegeol 07) S007 Aa cuubpeadu cjesoddousosoe 237 


. Macrocallista maculata Linnzeus, Middle Creek, about nat- 


ebENU SILAS. uebaabaded SIR Bette iaraaaan stan ea 234 
Cyclinella subquadrata Hanley, variety quitana, n. var., 
Quitana Creek, height 28 mm..... 243 


288 BULLETIN 39 460 


Page 

9g. Pitariacircinata Born, Saury Creek, about natural size......... 236 
10, Pitaria boucaryensis, n.sp., Boucary Creek, about natural 

. SUZ ET OR Man UNTER OU ee) Oana na 239 

11. Pitaria Guppyana Gabb, Port Limon, about natural size. ..... 237 


PLATE 32 (35) 


Fig. 1. Callocardia gatunensis Dall, Banana River, about natural 


oI She Han rela hea eeReR MEE ONS andl cach 235 
2-3. Pitaria VanWinklee, n.sp., Banana River, about natural 
SIZE a user hed ie cs eae 238 
Chione costaricensis, n. sp., Banana River, length 34 mm..... 245 
5. Phacoides costaricensis, n.sp., Middle Creek, length 11.50 
ELUTED re sres ee ee ore ee ra Mada eae Natta lale 222 
6. Phacoides bocasensis, n. sp., Bocas del Toro, length 10.00 
SET ee aM Se ON rata 223 


7. Chione hotelensis, n. sp., Hotel Creek, slightly reduced...... 247 
Cardita matima, n.sp., Old Man Sam Creek, about natural 

SIVA SHoncbdose UuAseiiBboodoued) ebsododeccemDaced 215 

g-10. Chione chiriquiensis, n. sp., Water Cay, length 13.75 mm.. 248 

11. Myrteea limoniana Dall, Red Cliff Creek, height 9.50 mm..... 221 

12-13. Venericardia Terryi, n. sp., Quitana Creek, natural size.... 216 
14. WVenericardia scabricostata Guppy, East Grape Point Creek, 

aboutematunalli sizeasessesese see eaeaet 216 

15. Diplodonta collina, n. sp., Middle Creek, height 6.75 mm..... 220 
16. Phacoides radians Conrad, variety medioamericanus, n. sp., 


Middle Creek, height 9 mm........ 221 
17. Venericardia scabricostata Guppy, Comadre Creek, about nat- 
(SOREN L SHV AD nonaat aah oaodeespesosope0nq000000 216 


18. Gafrarium limonense, n. sp., Port Limon, length 4.75 mm 233 
1g. Gafrarium altum Dall, variety costaricense, n. var., Estrella 


River, enlarged about 3 times...... 234 
20. Divaricella quadrisulcata d’Orbigny, Middle Creek, height 

TY Who ahaa Mie ANOLE Nas Ta RreRe CaS MCAN bonds 224 
21. Diplodonta insula, n. sp., Water Cay, height 6.5 mm............. 220 
22. Gafrarium altum Dall, variety costaricense, n.sp., Middle 

Creek, height 4.75 mm............... 234 
230 Moyateoa llamoniana Dall (Port Wim mses ssicesclelacsasiecnleile ae 221 
24. Phacoides actinus Dall, Bocas del Toro, height 4.50 mm....... 224 

%* * 


* 


INDEX TO VOLUME 9 


Note.—Figures refer to the continuous numbering of the volume (not 
of the separate Bulletins); Roman numerals refer to pages; [talics, to plates. 


Abra triangulata 32.........200, 431 
Acanthochites pygmeeus........... 170 
SPICULOSUS!Recscacee case) vceesemsis 169 
Acanthopleura granulata......... 169 
PIC CAE eiseeesws ne sictoistie lulnaieseewicels 169 
Nicimiceaycandeanal . eis. .c.eese. 151 
Heicoplemmaversssccoscesisns sea: 152 
muelaMnoOleucay  ves.cest ss: ferent 
PUUMTCeulatay vy. ee. dsauneeae I51I 
Actzon costaricensis /78........ 206a 
CIIDXSTICIG  sesasdcea ede uBSd eee eR sacs 45 
GMI CALS eas Siseeseseacnesceencnes 46 
Exc SMe eee eet tee BOL 45 
PAMVESLOMESIS: eee seeaccssssae se: 47 
AI CTS ISR aaere stoic te cienale sauces 46 
TURE KG INS cesaataeed en eee aEHE aH ES 46 
DETMOLALUS ee tse eens poet eee a 46 
PUNCLALUI ST oe ciiseveciclssnisisnseeess 45 
PUMCLOSERIAtUS Heese nub: 45 
PUSS eee aees saan 46 
Acteocina bullata......... 47, 190, 199 
bullata, var. costa- 
TA CATIA D7 Se bose lesan 2064 
Canali cullataeeveescchiecsoneecces 48 
CAM Ge IRs le Sd cli 48 
ThE log ooogoopbend 47, 190, 194, I9I, 
206b ; 
triticum-tritonia................ 194 
WIG WORT Shae Bae nanan AN ey 48 
INGeorbis) adamsile. ss y.csesuese sees 159 
SUP TATU S hapa wtee eee ee oe 160 
Admete microscopica................- 75 
PNESOPUSISLEATMS Ia ere neiee iancen secs 94 
PMA a LELVATICOSAN ee rcecaseuscnesel: + 138 


{GDN Gch NE RL a SME 199, 325 
AulaloiiavadiarlSitaaesese see wen ee reeas 127 
Cenithioidesye ste seen eae 128 
NIC CEMOMVACHEUS Meeeeetneace cy seeeaeee 89 
AD ae inva U AN a eau 89 
Ehoall oN eA DIDS 55a rlonsboooaqedaosHeHodos 89 
Abo a Dl eye bhTo yn oss Gouoomseonddoooaos 89 
DeatimMOntenSis daeessceeee sete 9! 
Caididissimiayserns-cee seen eoeee 89 
CONSENS USH eee emo rena ans go 
DORE SN OISV SI NG aa ee aS UC 89 
Pal VEStOMENSIS eee eeeaccase 90 
OUOKSSISH SS Thy 344. ndodghign InoueconSoses 90 
Losqidemodicay wis ees. naercuces 195 
losquiemodicajer2) eis cen. ss. 295 
TMCLCN CAN e Nie tees. cee caee 296 
ObSOLSUIS HEN eelnes west acee ae sie go 
LAM TL CUA 2 eee eee 295 
(EBUEUNWESOUIS) cote crises sodoostoduaacos go 
ball oreo. taal A SERB SNA fab NARS 89 
Auluimy Bilis erteshe meas seee eee see 202 
7a (op: Fe Hea Aa ae a aaa HARA Ma Nn else 57 
Amanra tombigbeensis.............. 26 
Amphiperas canadiensis........... 112 
Ampullaria caliginosa. ............ 141 
GISFOmRSS ss doobeoosoodconsuodunses I4I 
PUN SA Ey BNA Ny an 141 
Amusium bocasense 20.......... 378 
DEBS PA eA ae acoA sa ANAS aa 200 
Wy ODT EUS dae ce sauceaieoneuaal 187 
TL OGLOMIG eee eee 200, 377 
EO Unil ae is Si SED leon aac a 186 
Anachisiall bella ne eile, 93 
Bhaeoyalabiy sll. cod sadoocodoodseeBoboLe 92 


290 BULLETIN 39 462 


avara, var. Ssemiplicata......... 92 
Ware Sim tlisnyees see meeenesees gI 
Vat jtrams linatassd seed ssenenesee 92 
cancellatayy iia ue sea ninnane 92 
Corel Wes oleh 2 Wea Re A aa Sa 92 
un Sie Eig SoH Eee 186 
ihoOtesSseniatnal wey eeney. sneer ns 93 
Obese a rae a ees be un alie 92 
ODPM) hod edocs nc ooopabagdsay maoadoHbond 92 
OStrelcola ieee Weta 92 
h OP CEN Eines eR M Ean Ns Rue TUE 93 
Ancillaria acquaensis 70......... 266 
Chipotle ays iN Meee eu 267 
| OyU USA DUIS MP SERS NAMB USE Aeneas 267 
Subslobosay an haiaene 7 
Ancillopsis tuomeyi /7.............- 12 
IATICISEFOMASISS eee eee neato 158 
Ancystosyrinx dalli 7... ........ 232 
elegans 7a eee eer. 190, 232 
Tawa talent sue eA SEEM ABE 63 
Anomiayeplanp pile seeeeece cet 381 
Sim plexeeeeesesee 189, 194, 200, 
DOS N27 Os SR ada se nee RUnieie 381 
Nan 2 BSP MGB AA ea Sins ane UG ARE ROR Ep 35 
Antigona blandiana 33........... 431 
lean oINehoeh jasoudonsencsdadsesuedue 188 
Marrisianayiyeemeee essences 413 
multicostata 33......... 189, I9I 
AII 
TUL ZOSAM GRR ae esacaaiaae 189, 442 
tarquinia 3}...... 196, 194, 
412 
Aphera islacolonis zone............ 193 
Ja\y 0) bASN Wee Mico Gon Ao Aabdebocnucaubonob dents 53 
Arcalactinophiora 26 lier s seas 357 


auriculata 25......189, 193, 199 
362 


CACTI CANNER ee eter ncas see ssa nteaes 362 
Candiday 25) tiareaessce. 189, 355 
Chinigiiensisvysecsssersnacscse 193 
ConSObrinan yee posse seats 358 


Costaricensis 25)... see: ee 359 


darienSis! W25ncsesese secs 187, 356 
SalI CASIS me dee eeeee eeeee eer 356 
golfoyaquensis...............0+5. 193 
golfoyaquensis, var. medio- 

AySoCAiney’ | AO) ssoecoccadase0e 360 
STAN Gis wy hits eae IQI 
henekeniy 2722221934 OO 258 
Wad S27 eee owen yaneee 365 
LoKOsoKSINISIS: Abyatseodod ohandennse 359 
Pham opeKOr EAN io) Gaesnanncddocdes! sauce: 354 
UK (0 by “haa AeaMeee Aon acco Aobede 364 


macdonaldi 28...187, 191, 366 
TMAWUYeS) |) 25.25 eee AN LOS 


occidentalis 252...) 44" 189, 193 
199, 353 

OOM TENS sadcsesosaconccsogascooc 356 
Oza KENSISWMghe eee te eee 
ON OKs CIB BAL ura ha ad uRAAna sclatc IQ 
DALCISK eZ 7) meee 187, 364 
LEVETSAN cua una uaa ncbUnnn mee IQ! 
SObrimal iieshqiaenaaalaeneeere eee 358 
umbonata 25.....189, 193, 354 
WEENUO ON Alo Sho csacuaeeecouaeace 261 


Architectonica granulata 79...138 
190, I9I, 196, 200, 326 


perspectivaeeeastee ee 138, 326 
Aspella, var. obeliscus............. 117 
Vane PAallpereulaseaw ese Ree eee 117 
Scalariord eswy ene eee trees 117 
Assiminealaubertaman.ccescussteer 142 
Astralium brevispinum 78...... 190 
334 
(CHS ENGUIID d oooac Shonen sadose 190, 334 
Astyris) diaphanaeens ssa 94 
dueclosianmay (ewes eee ease aes 94 
Tuatha eer ees re 93 
Atlanta perOmtteneee-nssseeeeee cess: III 
Atty S TObUStOId esuareseaeesscacy Meccsees 26 
Ait ClANCOTON ane enn ese eC EEE EEE 56 
Oe ee Ne Pa 55 
PEllUCens es Mae eeesoasseecee 54 


Auriculina insculpta ................. 46 


463 Costa RicA MiocENE— OLSSON 291 


PMETACUIASEGUEMY V4. oa sececcaccbeesenoee 54 
/ANEVTTRIG, GLI OE mendasessnaecppeA cauaHeone 82 
FINCA DMA sscciee secrete cea escis 3 
OMUSLAL es lose callessaisio ns wesioceee 8I 
AW ACU ITMOTM ATA n+. <+-scsscccse 367 
B 
Balanophylla haleana. ............... 27 
TBiailaonientb tad a5 ae ae weer ae Ame 43 
Barneamallatovdeas. 2.0.0.0 /. 5. 7 
BaASwliSSalalttasccse cs s-0cesee- 157 
COSHUMa tae sees ccoe cle sssees sses 158 
SUPE I Darneeni see cucew suse oseees 155 
watdelicatiuilayesay cms ecel. 158 
WEEE, GIG DIKESREI A) Wocanangesdocseee 158 
BaSterotianypraima, fe 20 
Bathoxsyplalistaests. 2 vest svc alate sos 38 
Belawinlike raven anon) seavane ney cua. 68 
Betanodolinnm es eh we bE Il7 
Benthonella fischeri................. 140 
SAAR nnMea HLL Mee Va GARIN 140 
TAVISKOVED ISS Rau ema a NEE See Gan 140 
Bitinaemnboiplexs ese ase. 128 
Sint MIO ores sbecdeusuacede: ebake 128 
PaAlweStOnMensSes ny ve eee 129 
BAUS DYSeEU Baa GhAsconadeadonesneoner 128 
TEN CATTT TT AA eae Eee 128 
HAUS ALO TAUTAN es nase se ee seeise seen 186 
VIANNA soe aes al esata hee eae We 128 
DY MUCALS CATANIA) eee ateisiefoeeiels leis 135 
ISYIONEHA, GoxXGsc@lsconscdnaccoouseseuesue 132 
WBA TT ral ease ee eat NL 56 
SLC KOCIitaM ensues ovis babe Meee 57 
jOCINeHGlay San Sea sAbtae ABE) Bee uonmemaE 57 
IBOKEOLTOPMOM: 0. ssieeeslsee ocean oe bee a9 
Borsonia cocoensis S.............- 250 
Bothrocorbula viminea............ 442 
Bowden beds, Jamaica............. 199 
BUCCIMUM ODES. no sass sesesoc ace: 92 
OT DIT Sayles eres ene uses 87 
arbre) ae een AN ke 99 


ZO TS aerate nen tauan ante eau 94 


Bulings arora y eee eae ates 55 
I BY DUN ke WORN eee LH A A Ue ea 48, 49 
Di bhica tars pnd aw Weal ae aaa 50 
canalliculatame weceunrarenceueetee: 47 
Pawperenilaeemesceee keene 206d 
HEC Cer oe ae Han eee Nene 206a 
Bullariajabiyssicolas. |... sssseees: 51 
lanl AeXGIe VE) es seddan Baccioass booted 206d 
OCCidental sy eae een 50 


pauperculia 7...194, 199, 206c 
PAM SUIS CUA se wt ae eA ET 


Solid any aN hn eee ea 51 
Sta bare Ngee marci oll lat iin eral 50 
Burrs al arta as eevee maa ae gen 114 
crassa J8..... 190, 195, 200, 306 
Busycon canaliculatum ............. 85 
CATA CA re MUU AN Ab § 
Sli CSA TIS AUN oeE Me ve ae 56 
DELVERS UAT sie Weve eae eee 86 
DY AU ee Sous SaSdobRSoaHoseedes 55 
C 
Gaduilastanctusiuan cou) senna 4! 
AMT AIC Sue HOUR UN RRINS LTH 40 
CAMO EATS Seen racen se ta eeak Huet 4o 
Clo meas he Ak, Cee 40 
MTA OTA UaREE Nee UE neh aN 4I 
PA VSS es ee ea ae 4o 
POC TH NO AE oN 40 
quadnidentatusme: spsepecenees 40 
HErer HS TOTS inate eens Al 
Ce SiS eae eee 26 
CzecumWvannlatume nee 130 
bipartibuman Sarees tie esc: 130 
Carolinas eee ee eee eee 130 
COSTAE TAR Gre eee eRe nee 131 
STAD RUT: Kare ee ee 131 
TAS PIAL BUDTTD eee er eek ee 130 
TREE Ql aera Uae Ae Ms aka ee NNR 130 
TORT Cinna tls Lea eS con 132 
poeta tara eee ees 131 


292 Costa Rica MrocENE—OLsson 464 


Honida mums ee yee saat 130 
Calliostoma asperrimum........... 156 
Castile 7S ee Na aU SE 337 
CIFCUMCINCtum yon ee 154 
CORD IG TU HO Beene a 155 
Cushy ptusiy cou eeanawen aes 154 
TLPU AUCH HOWE we NN AA Ne 155 
NIMOMen Sighs. ees ena 336 
Mmaneinellan 73k Menai 336 
PUl Chery NM ee 155 
VATE AW SOM Iie see aul e eens niee 156 
TOSCO UN Cee Rea a eased 155 
CATT PD AS TNS TSP ese eee Mean ert 165 
DEN EEN He LE es es SAL MSs a 155 
VESTER SUNNY NUMA O atta 155 
yucatecanum...... SS MPa are 156 
Callocardia gatunensis 35....... 407 
Gallo gazau Zee Wns uay Me) Wiuaess 154 
Galyptreea, candeana 22) eee, 145 
JOE WENVABU NTIS Os Sonny HacenedcaoosadaueD 145 
Calyptraphorus valatus, var. com- 
PLESSIS Scans Nee Uses aw ae ea NU ee 26 
Cancellariaagassizil in oa 74 
Gell cea ee OC MIA A AUER A IA | 
barneth aig aes an 200, 253 
LODE UU Fei es ee a SIN te ee ROL 191 
CONGO Haima Sau Manag ek iets 388 
Clavatunl apse Gee ate aay 74 
COnradiana wen SE BAas 74 
COSSIMATIM I OW eee a anes 253 
COSC ba EN AUN Na Sei | 
darienay rom cananune Sos 252 
Epistomifera wisi eel O5 255 
galvestonensis................008. 74 
gillbentin i 7s we Meni oeiTi 
islacolonis 9. ..... 187, 191, 195 
196, 258 
ELL NUTSV ZO eens nares 195, 254 
TOOLSET EMM aca aN an nanny 255 
DATIOTICS yt ey ANNO Aalaaoe 7 


Plummet vomey ee eee IQI, 257 


FELICIA ta ae ae ae Ge 74 
CON KEUET)) @sdagnacssoosonnbos 195, 256 
SIMPSOn i ewe 75 
[ten oar gee eR Se Annan MEE 74 
tesselila tame neeseeeeeenes I9QI, 258 
MOSKOS | O)esaussccgodacen IQ, 256 
var. trachyostraca.............. 252 
tiberculatayn iiss) anal aNeE 191 
Cantharns limbatus...:..eeen nee 88 
ORDIQMYA as... coe 87 
CINCH See 2), eee eae 87 
Capulustintontus.:,..ee eee 143 
Cardata caribbeanensis 29...... 386 
laticostatal i... l.e semana 387 
TACIT Ai) 95.22). ae eeneene on eee 387 | 
SCAtD ia COStatar yy sseeeeeee cea 388 
Cardium, var. alternata............ 189 
APAtetICUmay. Whee ee ee eee OG 
Costaricanuml)| (GOs... eee 398 
dominicense 30.........194, 397 
fap b ab bar UR ARC Man Se REM S RAINS 186 
ferehobhalerelS,pagunddoscoasoces 186, 307 
JOKESRSONMDbOON!  EXObsadodsaonddn Boodss 399 
medium 30...189, 194, 205, 398 
Serratum Ou... £94, 205, 400 
JOO TOS I dota 189, 205 
Webeg WUNAKONON: | Or-cdccooncdecooos 400 
Stila tuna iy Ones seeeee eee 188, 397 
Sublimeatum sass ee eee ee eee 201 . 
Gareliopstsie i aMule Sei ees 110 
Caricellaldolitas anne sun a eames a 
Caryatis guppyana........ ......... 409 
@assidaria laevigata. ts.-:css0s5m 308 
CaSsisicaileo eases eee eee 118 
Alar imea eau eu gec maaan 307 
IMA atay NN) seach eee ee eee ee 118 
podYOVAHU NUKE loa donudegnacoodsoucsH4cS 307 
LECTUS AV LN Oe 307 
Snilciteralanwenauaaaes 195, 200, 306 
(HSI OTOBI EIS. Aas oadoddcodenanuoossios 118 
PuUberOsal ee Ae eae 118 


465 BULLETIN 39 293 


CAvolinaniniexay vee oe 42 
TRS PNMOSAN chase ue Us sen Uae 42 
ITACHI ALAA ant secre et ait 42 

CercadoMtornmation. (i.5...2.ceccosees 193 

Cembmicdeaicostatal..../.2.-....6.- 126 
IS RENIIONG nee n ances nee Ge sEaHaRe 127 


Scallanifonmis...-...--:-.-.4. 126 
lI Ietal beeen eee oscienindcs ooaseoes st OO, 


Wag COSAME ee esc. otes, os senn sss 2O 
Cerithiopsis burkvillensis.........123 
Chy Stallman oe. ese ese. J 123 
CHIEKSOMI nsec ie sees slain uote 124 
EstelllensiSia esate scene aS 
BTCCM OW esse scan teecessoeeteeasts 123 
TED ETISTIS Gootadcene ce a ranee eeBeeee 122 
Sto mlatarecnesecocsee, seems. <4. 124 
ai TTA sskekss. (cese se 123, 
Certthivm) algicola ...3............. 12 
COStATICEMSE| 7 e oe s25s 52s. 315 
dislocatumiye es eeetetcnsceaeoes: 59 
GID TIFT suooaudeo onoddbnoelees 125 


Ord dani tbeeesen eae ST OM 
PialWeESLOMCHSE! cre... 4. 404-402 120 


RECN e sesh sn ldeck vanes t 123 
Inte SreaTEwTT Gos souose. “Goddoesecee 125 
THATTHPTNEN TT neooobesseue sskanpeBsne- 125 
VATA SESCEMS 2.25. sac sascnes- 125 
IAT CANT yee atseis eee ew eee 125 
PROLEXstT TAY ecm ia-tas sisson ace 59 
PUNCLatwIME eens seeeeeceeeeeee 124 
tencbrallasmpen sun cesee ceca i ueeune 12 
ATS EDLOU Se eee alas aia ie 125 
Cheetopleura apiculata.............. 168 
Gama antiquata. is...) 0.0.6.5... 390 
ACIS acer mee ne eked scan tc2k 391 
CONTE Pata EG eee cme 203, 390 
congregatoides......... We ae 390 
CWhreileatequestris...22.6:.42 6225.5 .5- 143 
PME CEOIMTAM eee eat le oes eteE 186 
(CINE STHVoST ATES a nsnac Ser aaa a ESnE PERE 108 
Chesapeake Miocene............... 202 


Chione chiriquiensis 35.......... 420 


COStATICENSIS gbnestenossseaee 417 
SUSU ZO mr eee eee 189, 205, 420 
holotyanaly eopiee- nesses ses 419 
MOtELENSIS ig sanecencesMensesseee 419 
AMACLTOPSISH NG Feeleceseseee 188, 417 
THONG BGssoocudodornnsdce 188, 416 
Leg uillrmma eee eee see e eae eee eee 187 
NOWWATHEI.  sodoabeacoos 200, 201, 417 
CPO Tees LLU nae tee uni 168 
MAGI OLACUS meeeceseeseeeeeeeceee 169 
tuberculatusieeeeseeeeeeeeeaee 169 
Chi cores eas oan ta iibeaua: auatcinae 95 
Chlorostoma costaricensis 78...335 
C horistella pompholyx ........... 161 
Cicculuspliratusyeeecee eee eee 160 
[BCDB D-CA ANGR ar RE AAU HRN te 160 
Clathurelilayy ewettitnes.cs---ncessee: 70 
Clavatalaywelaiigeniaeseeeeeeee 317 
chipolama Le anes 126 
COstameanayai7ige) wcoseesess 316 
Clementia dariena 34....... 187, 404 
Cleodora Tanceolatal.:. 200. .sse eae 42 
Cliogne curva ness aus 43 
Dy Lai dataleessses estes eee 42 
Cochliolepis parasitica.............. 159 
Striata cular seas een cee. 159 
Columbella ambigua. .............. 298 
IMEDCAtOnan. sere. Aslea eee eins 93 
FAWISROKCLONC KES) co oagnoooncocsudoaqosoue 93 
submercatoriS 73............0 297 
Columbrariaytestaceaynseessee 115 
Comipressidensi tea tues 39 
Conomiitra ee Aue aes 82 
Connsiicemiulators ween enna 186 
AGASSIZ Iv. ehaes eek aun 200, 226 
Bhacly ol awieE FEES, cococabocodosnesosscoNe 62 
Der salisite ee eee een eee (eseun 205 
Vat DOSCEMSIS I MOl-eereecsee eas 223 
burckhartiviOmne ees 200, 224 
Catenatusrsn cca senses rt neasees 215 
EEMEMTT Oe as ate eeeemn eR yees 61 


294 BULLETIN 39 466 


Conus)ichipolanuss.3 62, 217 tortuosostriatus 4............ 221 
CONCAVILEC tama vee Meee es 215 NASH Ob Ny eae a ranma uuabeodoradc 216 
(LovaiKoloyGbaels) soy) papeuoonose 186, 195 Coralliophila abbreviata............10T 
var. costaricensis 6.......... 217 debungiae eee ae eeee ee eereee IOE 
C6 Fe BTA MUN IE eT NA 217 lactwealy akin) IOI 
delessentiamusieenss essen 61 Corbula dietziana 24. .....189, 440 
demiungsuse een. Lon TOES 216 dispartlis ye eee 438 
domingensis#ay- ae 186, 195 SHAN HOYOS OISTS Asjeacec Gadde coooases¢ 186 
SENSORS, 5 sAcldaonosdaonoddon aA 61 hexa qyrmmal2 7 ee eee 441, 442 
floridantarics. hse tesescere 61, 189 SHOUD UT UDIC GRAD ARM RE MARAAAA iol oeoboce 25 
BAZAN hen NL Wee Runt 226 oropendmlay io iene ee ee eee 439 
gracilissimus 4...195, 199, 224 OLOST 1927.66.22 ae aen eee eee AAI 
granozonatus 6......... 199, 222 Serica dike iy ae eee 186 
RAT TISIWMOvN Ie Rey aes me eee 225 Splenisss.20.s oheeae eee eee 186 
hay tenSishe ease ee 186, 194 Subcompressaii cee eee 
TOR PUBEN KONE yes caemaoasgcocauadd 217 WC LAR ITAL ti ane aenune 194, 438 
interstimctus) Oi. .e) se. 199, 220 viminea 37...194, 196, 200, 442 
[EOTERSIS eee Nee ene 215 Costa Rican Geosyncline ........ 205 
Nexo yah OSL 84 ae Nenee a eUG anaes 61 Costa Rica Stratigraphy..........- 181 
laVOTMeHSIS) USe7 ee ee 188, 219 Granopsisiic.saiceascccusecesceeeeee 167 
marginatus) 7..seee 195, 223 Crassatella antillarum.............. 384 
ANONMS (io. eoecoeaeetn ena 188, 214 Crassatellites bowdeninsis, var. 
Varimultilinatus: cess ane, 200 COStATICENSIS HG 2t enc eencaet 385 
ml tilinatus snes eens 195, 201 Fo712)1 6) OVW GAMe BEB ARE SoH anBeo Asbo sGb6s 25 
multistmatusi ze ence 226 TIE CACTISIS| Hg Ze ea een cies e eee 385 
HMSACTISIS qibuicsNaelie sus cana: 219 mediamericanus................ 384 
pealiee enw EULA STU ORC UR an 62 DEEVE MMM Za entee ae tent mene ne eee 384 
planiliratus 6.....188, 199, 222 Sepulcolllascessesseossereeseseeeteee 25 
proteus 5......61, 189, 195, 215 GCremiidesy\eeseeesece secs emcee meseeacee 16 
ON ASHITE DS aa agadscboeaauauducseedans 62 Crepidullayaculeatae eee I4A 
pytitormisy..2s:-4ee une le IgI, 218 COUDVERaN a rane nnccicece nee neete 144 
recognitus 5...... IQI, 195, 199 POT Ca talks slednseseeee eee 143 

518 Plana eases 144, 190, 199, 331 
GEA TS ees LuSe an eee nese 222 Crepitacella limonensis 76...... 315 
STATI deat aue mele 194, 220 Creseis subula.... cilia sieuiateinaleeeaaaetieet 43 
Solidi i or Crucibulum auricula................ 145 

RVers PARAL a AMON eE pPlamatiwins os ewesces esses 145 
SUCA TH SIN price adsaeecuese ace 62 : 

Scutalla tums eeesaeeeeeeeeeceeeee 145 

ene Shocliseh, Gh, “Gy Straten ma eee Cee a 145 

i Ua Cnyptonaticasesasse cere eceeere 147 
BROSHS I )eetn  eicosccooa acca: aha Cucullzea macrodonta............... 25 
tortuosopunctatus 6........ 226 


SatoOcday: MEAN ee 25 


467 Costa RicA MIocENE—OLSSON 295 
Cuvierina columnella................ 44 @ythexeavaltenmatayensescuscsses-ta 408 
Cyathodonta gatunensis...........383 loleraslianaes -bsdnane sadagdacacadac 200 
Wy clostrema afiine,.../......56 1-144: 161 daiienaly a arseu seme asneaetinecse. 406 
OMA AO Vex, seco enece yess LOM HIEDGKCKE) Cub dWansnadudennonpoocodoue 408 
HS etl OS UMMA pel Ws loyslfelee sjniecls= (sles 160 CAnChadiMaeeccostseeeloocsesblarss se 412 
ERO CMOTG ES ee eae eees sees eee: 161 D 
Cylichinajbidentata:..0) 5221/24).205€ 
dacs ns : Daphnella hormophora.......-+. +» 73 
TRENT Ss soso NS GRE EE SRS EE POR BBB 26 leucophlegma..............:2000+- 73 
Cyclinella beteyensis 34...191, 414 Atria Claas Ae nat aaa enemas 73 
CPARIEMENSISt tS A) ehe aN. 187 Wehr aia clay clay taemeauensectdetseceensias 72 
Hea Oyy, CUI ees seie velo sy sis Saison cise 191 perpauxilla ...............4. seeeees 72 
SMMDEAG GALA =. +...¢seraee-senne 191 Dentalium antillarum...........35, 36 
Wate iguuitanal) 34..6.5-..c00+4: AI5 barblebeivmeeee i naeeeeas elev ucts ce 36 
Cylichnella bidentata................ 50 bocasensis 18.........s.sss000 338 
var. galvestonensis.............. 50 Callipeplum............00-seee00+- 37 
bidet CUIM=trAtOMIS4s.-.6-2-0-- 22: 206¢ Callithrix..............seseeeeeeeee 36 
Cylindrobulla beaui.................. 52 CITES, catobobacsuobdaokooscags dbaaoonos 36 
Cymatium chlorostomum ........ 116 CSiAshH RNID \codoscasnIpgaadbboauOGuoUde 35 
CHy/MOS) oO mveY lubaols ear boncabosaeacne 116 COMPTESSUM.-.. 0-0-2... -eeeere nee 39 
STA CH SMe eM iy 116 GidymMUm. 0.2.22. seseeeeeeseeeeees 39 
Olearinmy Nee ee 116 Cisparile.............cceeeceeee esses 35 
Cymatosyrinx centimata............ 67 (2) DYOFMENEN Col sw osodesdonoconaadouLaHoOGNS 37 
{TUOSETY, SSSR US HHUA RHO ABHa EBON GeneUs 67 ENSICUIUS «00.0.0... veeee eeeeeeee £8 
@yphomialeibbosa...:......6-...4-5; 112 HUM ....... eee eeeeee eee eee 38 
Cypreea alminatensis 75......... 312 IBKOACIEMES,.. Nadsenosnoccocobadnous 339 
CHMEKEA 15. scdecccovsesses D134, 312 EXOTOIICLET s-.cdunodebsenocosungacaceooons 34 
dominicensis.. ............ 196, 312 laqueatum...........0...ssseeeeeeees 34 
ESESMMEMSIS. | 12... ¢5sciensceasiensee: 14 Te ptuma to coe eee enon o-noeesenstene ae 37 
ERAMUMETMAN A). se snsscaknerees 113 MHALATA, ..........0eseeeseeer eeeeeeees 37 
Wemelkenin guile 186, 196 mie diavAdenSe ss sssaeeee tse: 26 
DEIISATTT, Tel sasanealoanedeoese 311 <n ESSAI OPOEMSs ss coos gqgoaadcacosBes 36 
BALE esse ck soles. vats AS OPhiodon.........ee- sees reseeseees 39 
SPURCArestnetecesstiaccedaccsecs =: U1 perlongum.................sseeees 38 
Gyiprde CASSIS a skesceecscscesedeaecersess 118 PTESSUI, 2.020. e eee ee scence ee eeee 39 
Cythara heptagona ........... 186, 195 pseudohexagonum.............. 35 
terminula, var. costaricen- quadrangullaney ese eens 39 
SUS ONE EME Shon It 249 TEDECCACTISISH Mn Mincneecuenes 34 
Cytharella cymella.................... 713 semistriolatum...................- 37 
@alvestouensis. oe ea. 73 S@mi Ca tutes aaiecc nanan nanan 37 
irr atiay i Sereee ener ae ema yn ana: 249 sexangulare,...........00065 0.2. 34 


296 CosTA RicA MIOCENE—OLSSON 468 


’ 


Dentalium’sowenbytsceae sae 38 Carre Cay7 ii icnecen ele Maar 243 
Vat wpellicenttee eee 38 Chiriquiensisiy Sessereeeeees 236 

ey ay SLUT ee Ue hiemedaealnbe saree 35 Co) oly lB reper MES MU ESR Sa) enon aD 72 
tetragonum......... secede eae eae 39 CINCH Volcan teense eee ooate 64 
Weigh nabboolyA ea aie ye 34 CONG aby CT RORAMANne nts CCN 237 
Vaio OS tT gua wenvea sta 34 COCCOS SL ASIN Lay eee aN 236 
uscarianum /78.......... 187, 338 COCOSIM Ay Sass. see eeeiec ee 247 

DD emitis by layin o vane ee aNN Mita | MINE 156 Comatotropisy G4 eee B72 
Detra clay ae OA NO Ml ieaninn gs 56 consors 7....195, 199, 200, 233 
Dialeniawe eee BLP A eu AM ea Eh 42 Cristobaligy (6) see EEA a: 
Diplodonta: coblinay igs. seis ee 392 dalla, war. cestrota.. ose... 66 
PSU Aygo eee Ce eee 392 Getectays ye say te epeeoenneel: 64 
LED VU Se Mees ea ae Re a 7 CS] DYBE che AREAL SABES clei at enone 65 
Distortio, var. clathratus......... II5 eliveivyaly ee Aoi eas 72 
PaStuMensis eee eee saseee 306 (SabaeSnKepaskel 4555, suscdocoonqocoues 186 
HELICula tastes eeen any 115 estrella nay 77 eae smseee eres. 241 
Stet ira ae seein cela 195 CXASPeratalyi essed naesnoeae 65 

IAS COnbrioxs) sama einai a II5 S:ccfohallh Oy tuWaousdoneondnaaeusieadog 73 
Simi biases eean yn auen ie 305 eucosmia, var. canmna........... 64 
Divaricella quadrisulcata 35...189 ADIETHEONPTOHIS,, bean coagoqoou.cseadoood 238 
200, 205, 396 BUSHING MMU eas Joie evar 186 
Dolton Tee seianeatane 119 PALMMCUSIS Ey -cuseece eens ce 186 
Caminray ee DO es ALU AR ANG 196 haliostrephis............... 65, 235 
ellipticals. au nia NS as 196 AVA CHSISH ences ss caseeaeee sas 67 
ellipticuan resi a cee evan ie 311 HORNA econ dacadobqadonobes access 73 
Donax acutangula 3? .............. 22 NOPKCOVT EY orsnocopancdennscebaoobeD086 64 
Dosinia acetabulum BA LIS) 201 sbanVevaVSAISS) | Chapnoscoodoaccoosa0. 243 
203, 403 limone tian Sac ss-escnse eco 242 
delicatissima......... SHOP eee 187 IbbsaKosekerey Oi:7Ap se AReenbandgendacooos 238 
Drilliavacestran soon aon Lak 65 LiSsotropisys eee Heese eeeee ete 66 
albicoma: tN oo ce2 wale alMaenie 64 lathocolllectayenensssneseeaetstast 67 
alllaaod aitalyeren ee uMn aoe sane 65 = lithocollectoidesiioeenrcsr sce 246 
alesidota, var. macilenta...... 64 TOMS Cae eee aseashcsasenisciseses 7 
AGUZSTSIS) (iSene ee euene tee aaa 245 TT OCTISIS) Sete Un aneenaeuaeene 235 
AGMATILC AG San bie aiiin Ulalananlns 237 SPOTBISE), | Gibcbongadopea ueddooesoebooo. 246 

AUT ATLA Son nue eee nema tay 240 SHONBISHNEHWOLEY |! labo aadgouabacodonodee 241 

LRN aKOKSI a ea A IP ay Ie 72 MM CleAtan else ueenauetesen see 66 
bOcatoroensis/iSeeawew eee ess 239 OHSAS nooscu copgaspo00 ‘eobsd00 64 
IDOSCEMSIS) MiSinti inne we uee ose oe 245 POR NEROXG ABU EY“ "GoocaddacodonasesoesKsod 65 


var. carminura........ ate selma 66 WN ORME | 7/edoasacogaboanba09). cade 235 


469 Costa R1cA MrocENE—OLSSON 297 


Dida pellagial wane cscs seem ecwccecle 72 
MEUM LAS Ase e ca stetanWastes eae 71 
POINT CT I misane a reaenGaeebeses coor 65 
SHTT/OSOIO Wegasconsonscaboeosea adscos 66 
GIMIDENG Eis easepaaheaaaseaborsee se oasacs 73 
TELEK = sa pc bBo Re Ieee ae IS roaare 66 
PAEOMLOM AN nOsaeecesiensaeceeeaee 240 
lelicclleeisteectcloutiere vvislecissiciveece ceases 72 
ETS ETC Au en ese ssida as traealiniekees 65 
VAUIMIMSET I vecsceieaeiaceccsecieet 186 
\/SioS82), -Weosncmnncos 195, 200, 233 
aC etlllllmaemetitere: is Scceclome secu oases 66 
ZOOM is esene scien ssiccveneans 186 

E 

Kehinella nodulosa.................. 137 

Echinochama antiquata 37..199,390 
fccrebny dl erimiaseeeses sce ance ce se. 391 
yaquensis 37.............194, 391 
HI SCT MMA enna Weliavnseaioveces 24 

Egerella subtrigonia...... ............ 7 

Emarginula compressa............. 167 
TMT APLAR A Neaats aac tai 167 

Ey mibolusmmiblatts...s.ce.s.4-+.-csesceee 44 

Epicynia multicarinata..:......... 161 

ES MAS IOI ees cee sare nacneine Woeces anaes: 38 

Epitonium angulatum.............. IOI 
CllaiPairbbiel Aogasansnseneananeeoseer 102 
COC eae ess 103 
contorquatum.................... 104 
Genticwlatiumy se eneee cece 102 
FAIVEStOMEMSE Ms eee ctmee sesiee 103 
IbboKSrN EB NOO OS GanncconeeoonceetaeoBBe 102 
EOOTKC] EI bl 3550400 nesacuscacbenadonn LO} 
FTNOGIES DEO \aunade) qacoqneadcans 104 
saeaikailbbowbierqbbsst  Wfeodeoneacosdaso II 
TMUNGIStriAa CIITA sae aseeese cess 104 
AUIS ELA EMM 7G eee ee eel IO 
Toute EN WoT os eScHoooandacaboneceoe 102 
GANG oO vtagdooocecdeooaneasanees 102 
SCIPIO tn udneeneueesouseisetss 102 
Seni Ciiilawcayeeercms tee eeaceu aud. 103 
Suiba Crit utraleze accra ees sctines - II 
(WEIENEIE, sodcuscusbbs sasgooons onaes600C 102 


CUCINA e sl eseeeeeinesce cians 103 
EM atom lanai Stracscun coccercscten ses sete: 114 
OUNCE pao saaqeearibusoocoodhsq00 114 
EVM ge mi ti Caw ag ccsenssece-ee eee 21 
DOLICAN Ee mayictad saan seems bacceceels 22 
Mthaliameclusarnc.ys.ccctees sese sens 162 
SONG coeocasasanancooodasosoeedoac 162 
SUPPRESSAbeesceceteteccnscteas cea 162 
Ulf ail anasawars cements sclatclssclsecisesinecissces 73 
I aller a eas See eaten esis ciate 105 
Eulimella unifasciata............... 107 
A Deb aah cel ea Naas SAN U A Nea NAD an EN aig 124 
Eunaticina semisulcata............ 151 
Hupleurallican data ssss--sscssesnees 99 
var. sulcidentata........... 99 
AUG CILOLIMIS Sees esse eects: 99 
AEG S Oh ez ease dbcrecer eed sRaeHadeebenSeuocE 147 
TENE RECOV ATE 4racadecoossoobecoosouadsae™ 155 
cilia percracCiliS wancusmsscssaes Fas Of 
Fc 
Pas ciolariatdistansy.scccnnleccseseees 83 
PAG AML AM wacnesei/eaaceeeeeeeresee 83 
gorgasiana JJ........ JeCobSoqoE PG] 
macdomal digas ae asses 278 
MeClaian aye saiasewcsmeceseaetest 84 
(qEUIBGOY). | VCE odacocsouson 83, 205, 277 
Havarntiacelil ml OSaecueccssceaseasessees 96 
VaTalevi CUlassuesnssseen ce 97 
intermediarce sss PA Ge 97 
paWBKOYSNNISY CuarcuacacemosbadoN GosSsaubS 96 
Jeb Geriwtesane=s bebe Gogseepsnqgououosoodeaboe 36 
F issurella alternata........ 166 
ASTIIGIANMA Ms oseueceseee se cee sweets 167 
barbadensiseeeeenseaeeeccsecees 165 
Fissurellidea fasciata............... 165 
Fissuridea alternata.................166 
Galy,CHleNSiSweenuaecsencenceaeesnee 
ISSURISEPtamacseamaccesrneesteccseess 167 
Flabellum conoideum............... 235] 
Fusinus cepynotus...............2.055 85 
alcimiuse ee deteuasicccecince 84 
AIMTATICUS eee eee seee cee one seioae 84 


298 CosTA RicA M1ocENE—OLSSON 470 


COMIC ee ME iat sabia 34 
SUCOS TITUS ay wena een alee 84 
ELE ESS UTS NR SUITE ante Fan nts 84 
MO TIETSISI MiZpZeee en aera naa 276 
MAM OCOSIAINS pT eeeae nye eee 275 
HILO Tee ace vue cae ea Beane 26 
OLLOMIS ese yO ieee 26 
CUEKEON IS Hee renner sass DO 
eSehla tise etc w sees Died 8I 
(MOY ABTS NC AeA Gs a 26 
G 

SNOB oe oese Gacueaoes cedee doce uadocdoon 4I 

Gafrariam altum, var. costari- 
CONSE aban waa sre nenanee 406 
IiIMOMEMSE! iz 5eteeeee eee 188, 405 
Gastrochzena ovata...... PN Ag 433 
EO CULCa  eaea 200, 438 
Gatimistageieiey ease ee meeee 181, 162 
Gazairfis cheney een aes Ba, 153 
SUPER Da ees Waetae Taiaccuet 153 
WALSOMM see oeseo ses FA aia £54 
Genota didyma........ SEN Oa 63 
TATE OA ea eae ae allanol 63 
Gilbertia estellensis 2.........13, 26 
Glycymeris aviculoides...........0 25 
CADMAS Dies eee sda e ae 349 
CarbaSina)i22u eee. ahtesihes 351 
CaStaneus 270s We AT OQua 52 
decussatus 22...... SEH gaa 352 
Glycymeris decussatus. .......... 189 
jamaicensis 22...194, 196, 199 

350 

lloydsmithi 28......... Seis 352 
EDtlObICOSta Use een eee 349 
Glyphostoma dentifera 195,190,248 
(27210) OF WERE Soca n CN Ana SAu A hag ah 69 
ES HDD Pe ae SRN A «269 
aaTOVHODLSE! opeo MARU neat e 248 
Graptacmien was we Ieee ee ey er pees Bil 
Gratelupia mactrop3is..............417 


Gurabo formation................ aOR 


GyGiT Swi eee aes eae nae 115 
| 
aliavamericarialya 7a eae 251 
HMamineavantillariunm-s sss. .e eee 52 
antillarum, var guadalup- 

TIS TSA, Sa a ane en 52 
Cle sia SR eee eae Annem 5I 
Sul ara ae) 5 an ee ee 51 
OYE ICH se eae EME Hauer KAUR 52 
SUC CIMe ANU aaa Eee 52 
VARESE OTIS Heo iC: skis ee a 52 
FTAUStA LOT 0. os 2a sel eee 134 
elie depressus) see eee eee eee IAI 
Feteroschisiiialies ss heee eee ee 36 
Ely datinalaplaysise steer eee 53 

H 
Mnonitbs sob oyeblbban yo yale sodesndcadnoos 145 
Isapishebsoletaeneus sca saee 137 
Ischnochiton limaciformis........ 168 
JPEVOMINOSBIS Cecgdenocas) coucsoeosoes 169 
PULUIpLasScenis ieee ae 168 

J 
Ueborbab ee yRe ip ee ee eRe osueEaean 110 
Jiamthinacommilnisys---s- eee: 104 
fragilis ...... raph Ga eoetiertet bra) 104 
PlObOS aye ee 104 
Jamtla ina ay eee We eee ee 104 
pallidayee ae sea duane a wa 104 

L 
Labiosa lineata 25.....189, 205, 436 
Leevibuccinum harrisi 2........... 16 
Leevicardium serratum ............ 200 
Sublimeatumlbieens pecascseeseeseee 200 
Weevil denitalliurraee secon se eee 37 
WamlelWarialram oajeeeeenseeeeeereya ee I51 
Warm PUSTa ie ur ve Ces aus 116 
DET OED- 6 aie n Wen OUNAL No MAb iA aia na ine IOI 
Latirus infundibulum 77...190,195 

200, 279 


471 BULLETIN 39 299 


IL ANG WIS) Sheet) Wei gana Nan RN i 279 
(GANELECIENE) «(UC pean cmpeRE ARG e CRAG SM EG 2&0 
Webaibalbore! 275.0005... 0. 6c.c cdc 345 
Geulbvaimang zc s sca et) 347 
Wedardavalee 27. oss secede, 345 
SHMSMMONCISS  Drasessgneae, ayaa oe 346 
HMM ARMETSIS Wee cscs e see ecee 25 
DAMIR UM oat elses ce'stina dae nationcde ses 25 
GIETROOINNS -Sassdoanesnostensunseoae: 25 
LAME TASS) 27. tes locas seas 346 
SaNMO Garam Nese NN Aha 25 
Wepellay cwbitcolae se sels i. sdsesses 152 
Lepidopleurus pergranatus....... 168 
WED LO CHALOM OCR MLE cole 168 
Weptothiyraiidutas ce. sete 153 
WSU COMMAY ees neceo ho deascseseaaneed: 56 
Wencosyadimr CMlOris)) (Sse. 66e: 231 
SUSIE CIS ee te sncere essen sess, 63 
Sully ooreuama cleanse ee sees 63 
EM OCERAS Waseaiaseteeeecanccacece: 63 
RVC terol Nene aecee lh us kae gah 62 
Leucozonia cingulifera... ..... ..... 85 
OCC ate aR Mee UN alii 85 
Wevitusush@alei u...s5.. .cc60ccoccese 26 
DAOC a ey ease UNG 26 
SRST ces eS Henne SaN eB ea ncses ena 7 
iLpoongeel, GOlliG eRsececausesenedcodespaceos 194 
HimMoOpsismoniis) 29. ...../........ 349 
Ne nceacolidiay yo eee see vees. 199, 381 
ILO SGT douosenpeeoudene uaaaeiistse sober OL) 
IG rtoysla): neni rll ease deapacodepecscacsess 154 
PASTTOVANNE)) SoecotcodonpleeHoReaueuees 154 
Gri Canin atari escapee decsosuaes 154 
WiSSospita proxima.) 6.22. ses... 6. 160 
IC sheave) \sxermmoyb:e AeA sda ace ieooasane Ley, 
HVelantOStomMal cheeses ss eeee Miay7, 
Littorina augulifera.. .............. 136 
TENOTALAM Been wetteceeoteseascuer es 136 
Tao SSy Osi ebaN paduacsdoooreceeeanodae 136 
(CECI S WIE ean aaa Ho Been aBAar Le 137 
ZACZAC Hoctenainst nee ceo oe tae de 136 


ome chige Su eeten tas ees eee eA 107 
Me OVO TATLTTA AN eh ok cy wR 117 
Pucapinaadspersavoses:eessen cess 165 
Cancellatayy aes p tes 165 
Fascia tan Wen eure Ena ee 165 
omGurasensisyes.coselccescoeee 165 
SURAT Ae eM AUS Eek aN NM 165 
Mucinavenistatan wes se. eeee eee 426 
ivianiCataliis wos wean dee res 396 
PEIN OL ealhs wave oes ee eee ees 19 
Guadnisullcatal iss: ee ace 200 
MeV AtTaroecavenlovceeatee sees are sacs eee 148 
M 

Mocoma costaricana 29.....-..-.- 428 
elongata en sue weniec Avene aes 19] 
SAtWMSUSISHNZO Mes nee eee 429 
Dail ik CTSISH Ae ee cens lone esau nes 1G) 

WAT Canalis Zon see doce 429 
Macrocallista maculata 32...... 189, 

406 

Mactra alata 37........ 189, 194, 435 
Ce Kk OCTANE Bia aan ae NE EH 19! 
estrella nay 7nesseseac ss IQI, 435 
CxOletaa yee eos 189, 191, 434 
SSUBRG EAN DD ie Ay am IT 437 
SUMCO]A MOA G hs Saitou eee 436 
Mialeatcanauratsosos.s. 186, 191, 200 
ello bicaGeeeeaceen eee cteen eae 311 
TUG STIS MeN ee yee a Nia 191 
Mireur teases ae ee ee eae 149 
Manto iliaantomiat ie enst ssc ccnes 71 
ASERIC ELAS Masi Mee eae oe 69 
atmos ty las snolitacsans uasenanacs 70 

Dal bea tae eR MNOR eae Ce MAIC ep 69 
DICOMICA ee UO Baal 69 

Cos gifs re BR See SS 70 
Cerin] ae eer 71 
isitm@ rit ell aeneee sae aea eaten 70 

Mm elaniti cae eee tes 71 
Pemiplas Creriees secncseuscoeccecsas 71 
plicosa,...... SbHGHaBObacoRaHabenesess 69 


300 BULLETIN 39 472 


quadratas../.- 2. iaddagk oescod cof I 
DD OL al Y Ke Mr mes ADGA ENS 70 
Stell ata ee eee ee eae 70 
Ia O) Clary een seas lence celsceliei sa matak 193 
Margarita tllosalepeeccnesusccnicecnte 154 
Marginella apicina................0. Sel, 
AULCOCIMT Cayenne Men an a iis 79 
AVEWA Hie ONeers 80, 188, 190, 269 
Carn Cai Ne ena UN  U ae 77 
CESS eae a eS 77 
Collina pore en et nen 269 
COMEORIMIS se escort eee ene eee 268 
conoidalise: (oe 78 
denticullatay eee ee eee nians 78 
FESS Do TMi ie ain as A MPI, 79 
SULATICTISISH eae eee eee see ae 186 
Had ria aeRO: 80 
laerimelar ed oe nein eee ECOL 
VENGISSS Bank B canedenadanadorsuedadsads 268 
Lean Ger OMA acecece anaes: 270 
macdonaldiyvouse sues 187, 267 
WMINATERSISHMON ese Nee eee 268 
BIDITIATTV A ese cena enees BBE NAn ese 79 
EPRI ROYEAY Noe eno Besdode abe OR ae 79 
MAUUSACIMA aM eee 270 

Ob Ong ae RU NaN eR 80 
Opalanay ie, ve. Bo esdoSceHAnod 78 
Ovalliformis meses ee eee eee 80 
PellUCtda LOU 78 
Var pilsbryal i yige eens. 268 
TOStTA TA eau ewecs nsec sees anaes 80 

SO retry gee A Ae 80 
StOLER a ee eae A ROIs 77 
SUCCIMG Al PRD e n Nun aeien a Ne 79 
VANE ia tas eset hee een wa eae 78 
WATSON ea UE UL gaa ae 78 
AOE SEAN Bon cooboncuosuoasucaoace 80 
Marseninavamplanne the 151 
Martesiarecnrvaii2 assist ee 19 
Mathilda yucatecana............... 135 
Meioceras deshayesi............. ... 131 
TACT CL VET re on IS a ae ns 132 


Melampus bidentatus. ............... 55 
biplicatus ee eee 56 
bulloides eae 56 
eCiromalUs hese eo ON nem 55 
Cofkenset ees Lie Ee ae 54 

Var. oundila cliivesse eee 55 
COMMOnMTIS eee 
COLREN ALIS Meee secsbisece ee neee seer 55 
CTOSSCAMAREEasece seisaceeceene eee ree 56 
Havas eee oh ea 55 
Bley gle aNbIS) sso cusbeecHcoodce’ dobdocos 55 
In\ey ocho) bb Ub ly MoMA NAR Ndia uacobdoodoc 56 
f EUDIOO SS Ea AMBER MPMES corns doeeecn: 56 
lnmeatus aii...) sae eae 55 
TOT OSA erica st ah aie Manne cle 55 

Melanellaiibrlineatas ieee nes 106 
COMOHC ERE ccocbaddesoosdoboonsesse 105 
BLISUS Pca Ns Seat NH ae ale Neg 106 
Bibba es Cas eens 105 
STaciligy ye eee 105 
hemphalliy cece eessoveeeeceeeas 106 
Aten dias eee ee eee 105 
JAMLAICEMSIS) .n-eseeeeeee eee eee 105 
Subcanina tase eee uaa 105 

Melongena consors 72...195, 200, 

284 
COTO ae ee See Aare ane 86 
melongenay sees 87, 200, 284 

Mere trixadanienaeerssecen tne 404 
SRO) aka Fel HasUanoBaseecamaddbo seebeo 25 

Mesalia alabamiensis.................26 
PU a eee Ue ee ee 26 
WAESONETISIS eee eee ee eee 26 

Mesodesma costaricensis 3/.....437 

MES Or iy 1 Supe eean eee eee eae ere eee 84 

IMetiSibar baliig 2 nae eea nen 427 

Metula cancellata 7}...... 195, 200, 

287 
Sabi Mega unesaeeasw anes 287 
Joke nee chy Was eee aboneodatiacHandandac: 288 

var. limonensis 7}......288 

Mexican Mio Cene tusceweescsteneeen 200 

Microdrillia rostratula................. 7 


473 
Mitra alamagrensis............ 188, 275 
[DIB ASRIOD Acs. Sscadsaadoda) cocabpdoobe 82 
VaTeCOLralloplailal Ose seca 273 
GANG ETISIS) Oe) Saciesacellseeesc ce 273, 
GASTaUVAVIENIE) ask oho sdbdcbenpjoHoodone 82 
ALTE OTS TASTS ecules Welsioe 191 
MOTUAMI ON Meenas ulsstenenisians 195, 273 
OAS Ober menses sesierimei 188, 274 
MUGS: > Macodoo esos eos cccees lays 77/7 
GUNTANTAL OEM seindne teondanaas neorocace 82 
GS tay Ig AMP Pe ee Ns larisiss as iel asics 82 
SUVA SOMMer eis talsac scease cn se-icas 191 
swainsoni, var. limonensis 
GMa cererisunn ke nivale relents 272 
VATICOEIMSIS cessccesdecesceiseeatece 82 
Mitromorpha pygmea........ ...... 82 
(EAI OMleN  Gosnondessonobobens ncdooe 51 
Modulus floridanus.................. 129 
MITOGMINITIG eae aise sissies eniisslls soni 129 
SBTORGM ee einen uve te cuetee slant 25 
Montacuta bicuspidata 2.......... 20 


I@DSE! A SodasnsoobadsonsbodessnsAH! 
Mulinia lateralis, var. sauryen- 


SUS iene sea U ean 438 
Mitwreiencles), Teverrol onlUbl Aaee sbscosasdosse: 98 
Municideahexagonal....0 i102... 98 

ATALANTA TL A) tetnus.sssieieclins velee nee 98 
CSHIEBIIFEDIOL Gogaodbubasonnone 4oacudd6 98 
jOlawlijoyomehaley Seanseeooonadanana: souee 98 
Wine xtalS pe terATMU Shee cc ecto ces 96 
IDS AU ML i acdateGatiennats 95 
[oven aboRoyaGy Aasasaadeecs a aEner ees odtion 96 
Cab citaee wes cee ssescwiea eclespelsaleea sey 95 
@QUNDB Is acdosd oscadonhosoebauonaE 96 
COGMUTKEC EIS eases sno-ee 195, 303 
HU WESCSUS opsocoucscssoeauebdecdese 96 
MeahebhalSsaSiS.s05 Indeesadacaso, eaoodd 186 
PIKOOOSE A poaadsea qoduac on uaERRoneSaae 96 
messorius.. .... 95, 190, 195, 303 
OCI AES ee eee cetaaln cium sicietre 96 
POlyMemAatiCusiassececesdcees-: 186 
[DGPS TOs consviendces cabddanuedodasogec 96 


Costa RicA M1ockENE— OLSSON 301 


Myrtzea limoniana 35...... 200, 393 
N 
Nassa beaumontensis............--+0. gI 
INOW rel yoedopaqopeoosn6ed aodooRdod go 
OO OIENE es sonscmododdoucoopasodadasoc go 
DUC OMA Ese dosent eyielewsealetestier 12 
NiaSsaraimallo ts liitenwnescsesscccosia antes 89 
Columibellla tae cere c scala 88 
PBs A ton bo dbonenenoRD00d0 HOdodaHOC 88 
INaticaallit(calll oSals-eereteeenesticses 147 
DONS Heese se ece esse esectiss 186 
Cala zZOnaliSmesereeneecate vere. 186 
caurena 76..147, 190, 196, 
199, 327 
Spaoutratbll Wal ococacces asesapeoooobN \ocg98e 26 
FORCIATIA Mae bene secnaceecmacees I5I 
SUP Py awaw7Owevessaees. 188, 328 
J AIVIAICCH SIS see. uescnmnaseernt 146 
EVOKE so kescespaccoancnopAnecoudcac 149 
Miksysi Gl ca eee Sk aero eet 146 
IShoe hull eH are aaa anos HOH hn Asa 26 
TAT OCCA AME els acenete renee 146 
ON TRBISLE GUAM IA MEER O SA SE AM GH MeN ane 26 
jouhveene Nal seh cnoussadsasooscanads 146 
TOL TID ee sa aed 146 
jONUSTUU ele, b Sansa Dod oHoaeeSboHsonouGo 147 
IMENTS sa sodnanoadoncosebonqaudoras 26 
Sathorn vate pau Meee Saensies Woaeas 26 
Sub Clansaang esse seecaeeacereciee 329 
GRU KOH Ya conassaaasonaaceosseedadanes 328 
SOULE Ieee ncece ee actaseicerleteaist 196 
youngi, var.cocleana 76 327 
Naticimaplacteate tia ehescecernesce 149 
INGOs iitrattai ape ne ee ee eee eee 112 
Nextag pelorontasseyecce-seceeerree 162 
tESSell arta ee ee eS acee canara 162 
VETSICOL ORAS seen ee ee ae tenet 163 
INIGIALGWAE), FOYOL A cacdaaocoasooosaoosceonad 164 
HECHivatassissnceuee see uetae 163 
Showa Cerise ceecneaeeceee 164 
Sparsilimeata yeseycesseeeeee seer 164 
WAL Pal ea Sue oueeeetesnen seas 165 


302 Costa RicA MIOCENE—OLSSON 474 


viridemarisy 73s). 196, 331 
VALLE See ese UTR At ga ae 164 
ING Veritauibriinmea eye ease anene 150 
CamipechienSisniesimsseesnaes 149 
Ghuaplicatale se Meaney. 148 
foal hbWey WARES SAAN RAB AAUP UNE IQ 
moat llariseaeececoseeceeeceeeeas 150 
nmereidis eu. 187, 191, 196, 330 
TULL eee eee eee TA 
fexasiania ya Winns auiy 149 
TAD CT eA St bleu Eat ea dee Rian 149 
INTIS) Bevel SeS nos gavasdasdooss sasdowobs 106 
Splendidiuilay Wayne seecen. 106 
Nitidellatparnviuil ayaa eer ues 94 
INO ROTA VAs OU id aaah Nea) 74 
Northia morthige eer. 190, I9I 
northiz, var. miocenica 
TZ SSS sbpowesbocobesopaGEbosaaoD 296 
Nucula cahuitensis 27............ 343 
ASEH Calsheecebu ace entean ty arene 186 
ml ediawalaye sen sence eee nee 25 
Orbicellay eee eee eens 343 
OVA eee UES sal era a 25 
tenuisculptay) 27.0 193, 344 
Nyctilochus femorale............... 116 
0 
Obeliscus tesellatusy eee eeeee 107 
Meine rae PEA aaa a Go 117 
Odostomia acutidens ............... 110 
ats Chay ize ea ee 8 
CLONKONKG KYM IBERIA ABA a LL A IIo 
ATP LESSA A URE eM CaM aEm ed 110 
GesaabboneKolah Asscanedes odooboxe hesoS 110 
Oligocene: CostapRicas ayes 181 
Oliva brevispira /7o......... 195, 260 
Carolinensisw saree oe 75 
CInCiata elma NU An alae 75 
Cylidnicani7o Maes 195, 200, 260 
Oliva gatunensis  70............... 261 


SATAN see ANTRAL 261 


Nay les aR HAS ae A AIA 75 

litteratae sue rol ce aaa 75 

oo AOEMASWEY WO Aosoanecadsdoness 263 

PRELO(OWULN EIS) isbadeoncanacdooane sunoce 260 

sayana, var. immortua 70 
205, 261 


testica, var. costaricensia 
7O 190, 262 


HESIEIOEE)  docoddogseusdpoodson I9I, 192 
Olivella, var. bocasensis 70.....266 
IDOWUSSAICION 7O NS. ecco 265 
(21 bi del(o fs) ar aren aanMmRRerEcn a Sriagocaaae TW 
VOT allele nest) ee ian 76 
PalvieStOmen SiS) see eee ee eee Te 
OMAN WON teste eMeeanee le 264 
FaSpidea ys 25 ua eh Meo ua 76 
lito menSis) Zoya ee 188, 265 
SooKO DEENA Su SABU AAD BR RKO Bua I 26 
PTA Ga Cars 0) keer ata a re aa 76 
MUticoides ZO 195, 264 
TVG A Ne AU e Tenia Oe 76 
jOLOKSSUU Fa OR eran ea Lee 34k 76 
Semilie nth ca iy wees ese eee 13 
SUD temalra ais uy Seen nana ral 
LGESTUBIOEE), J ses Hoscooseccoodasqbedcaqon 192 
QMS Cia eS ean ae aed eae 118 
Oniscidia dennisoni...............068 118 
Oocorys abyssorum......... Aeaseiene Il7 
Ostrea costaricensis 24...20I1, — 
368 
PuehdeboleralsilSyarnodiehasdsoanoscboke 186 
interme d O1desiizeeeeea se eae 23 
MEesod on 27 eNas 189, 191, 
194, 199, 367 
Scull pturatay ese meee 201 
Ovula regularoidea 7......... alee 14 
(Oran lotankananveteal mL A eee ie 112 
EhotoUblawebbaal aAuranog ;yonsoenaocaney 112 
BROWNED HONS BosaanGansedddas Géods 12 
@RUROI ETNIES A onodoosdenanee wsboce 112 
Subrostratumecees eee 112 


475 BULLETIN 39 303 


Oxygyrus keraudreni............... It 
P 
Pachycheilus anagrammatus.....129 
Gaynillierarere sca noneancobeseeoncseace 130 
SUA WAS een eskascovadenaasenes 130 
Raludestrina aldrichi)......00..... I4I 
AND CTTATI AH eee ee LeU a 142 
CNM OMA bay wenense see vecevncces 141 
TAU IR eysls ea seielaetesemeloies 142 
SD eNta Ae eee ees ecsnieclcis are cee sibesis sive elss I4I 
LUIGI CU Deeps aiaala'se's'sna/ seelesidagiai 142 
RanopeambelllSensis) | 3..0..Jes6..-) 22 
JERR NEO) os cage cod oeeoeBeaeedn ne sbeebs? at (0) 
IPEISHUE AKER, ABNER elo pocanpoaneosenceoneore 7 
Reartellanasstcnijeseledesoaccadeegeg ee 58, 151 
SOMES ETS oases at eecustsccusomnisesi 143 
ROM Cabal eeesen ce cecccsues ie ssteise 143 
Pay TAU eAatlameecneasee cess ce seseise 149 
RectentallabarienSisicscsscescceccce 25 
AlminattensiSs 27s .0) ose. 375 
OLMIS MOTE Meco catce acts ceeculees 376 
Coraliliplailan) Zones. 188, 371 
COStATICEMSIS! (27 yeeneeene ees 373 
ChOSAN ee Sasi uils weedeouusaebens 186 
SALMMSM SIS Oscars eecses 369 
levicostatus 9...187, 200, 
371 
ISITE) AO eA AM an PRED acon Tel 
MACH OMA WOM nsees ssece oe: 370 
Olli@OllejontS.njsabanynodooons poseseo be 375 
operculariformis ............... 186 
IRSA OES) AOsssescadodsocsssas 374 
TRSILIG|ENBIS, bao sceesbepdedoedeanee 186 
Santarosanus.............- 200, 371 
WEMLTANCOS EIS souoscaquducoseasceons 192 
Pectunculus castaneus.............. 352 
GISCDISEENENIS) co sonadadedsseoos 350, 352 
[OSI TOSS jeeaoes dosegone 350, 352 
inedicullariavallbiday we eeeneesceses. 113 
decussataee se teess RBA E ATED 113 
Redipeselonsatusecrsscese-ce ees 54 
TaTRbRAN ONDE Senasciou MocoBocoECESeeEeae 54 


SOME COHNGKES,,.. Sonaénoodsobusqobseccbes 54 


OVA SHeeeeeriencee ee sneiies cies 54 
GidadinidenS ees scsatecrsomeees ee 54 
CHIGENS Baars ccewthos tocecerecaees 54 
Reracleneticuilataycsssscencese esse. 44 
Peristernia insula 77.............. 282 
WON UbIEXEN) | Wiay lencooasop se stoans 282 
Reristichiaraortee masa cssseancscers> Il 
WOKE) LOAN be BBS Gacodo dobocd cncoadeon III 
Personmay stra lilitadase ees cccs came sce 305 
Petaloconchus domingensis 204,318 
ELECCEUS ee ereruricieles|sleniselsleniserlseis.9 133 
eae Cu atl Sueeeetarsloletsetetsals oat 133 


sculpturatus 77...196, 199, 
203, 204, 318 


VAMTTATIS Hy Wei uennani cic ain cratstare 133 
Petricola millestriata.. ......,..... 187 
RetrOtTnO Chi S eee scc eens 164 
Phacoides actinus 35....... 200, 396 

DOCASETISISHMpoestersceseeeseas ce 395 

COSEALICENSISMGSieesseteie sees 304 

var. medioamericanus........189 

RACHA SoM eee RTA 189, 205 

radians, var. medioameri- 

(CANONS | 267%55) wdoogudtioooobdec 393 

(ero IGANEWIS) ncausocose 189, 205, 394 

moiuiliferum 75......... 200, 307 
Phasian ela ieee neck asc seceme 136 

DRE VAS se seU ni eats ue ee 152 

pOMOVUHIS, 2) oda anu sudonaecBogoceo Secbe: 232 

ORIN ey, Gsosaqcooson506 adoooabAb 152 

PONE C ENG BoGocogsssddconnseecadde 333 
Dhabi beavers >>:UORE\ cased dqsascuns aéana600 53 
Phorus agglutinans.................. BGO 
Phos) aritillarcammy ese. eenes | cesteslas 88 

IbeteivjenSiSimi/2aaee ese aasnstes 294 

(SUAKG Erloncosasaceod! eodusuan oaasdoae 88 

elegans eases ancor ... 188 

elegans, var. limonensis 

D RAR EEE EAE DER BSG Nae 291 

EStrellensisimZeesceecoreee see 292 

PalvestomeMmsis seeeeces ss eeeee 88 

RYOTE | MEeasss5050 000980805 289 

grateloupenSiSwerssssacesssess ee 88 


304 BULLETIN 29 476 


ATCT CALUIS Sse ee Neen Magee ene nnauas 88 
00K Mb KO KEKE) VAZERS een asoscoBee 294 
mexicanus /72.....200, 201, 289 
Moorei, var. costaricensis 


EDREAMS Me ASAE AGREE 290 
PAT VS Wie ean MeN et Aerials 88 
subsemicostatus 72........... 293 
NSEC OE een a dogabe 293 
Bh yWonOrs eaastencecromeene acre 96 
Pitaria boucaryensis }34%........... 4It 
CIF CIMA tay eZ. ses IgI, 194, 408 
COT AE AR SGOT AG ee rater 187 
SUP Pavlos ae eee 409 
hilli, var. musacina 34.....409 
SDV VOUN AK EI Oe son Ousnh oda conedesoude 409 
Va mwatelese ry Gemma 410 
Placunanomia lithobleta 27...194, 
199, 382 
Rlanorbismopiisun vue cunnca nse) 56 
Palartay;s clad Gest ie ya nana Murase ease 4o 
PLCUROLO MTA NA ROE AAS LON eresaue nat, 63 
ACCOM a RUE ON a SUNN Si 26 
MNO Ke Leben ine ie ANE L NARA TANS 230 
BLESTMO Lata yi ana 209, 234 
DTT ae INT A Aa a tie 66 
haley Manabbonl) oooh dun esnogadoouededc 220 
Pater tetera eaten Ne NG Wain 230 
blalkeamalsci ee ram waa alien. 68 
nyebey loyeobal ole sce ous ddWoonoe 70 
COR TTI AI Ue MIG MUNA RIS Nae Ne 70 
COMSO Ras Cea OA Et aR ds 234 
Far Mery Wie wee er ukeserene 8 
FoR eeeT BLO Eee ACS ee ASE REA ea 69 
AAT NSIS nhieu ne Boerne 230 
Sve! a porn ds MAA RIAN en ea 8 
DOKSH ea HOKORY Ae oA nKe a ogsonmadsadedaae yf 
Morekeah oYSrasHIS. Won ldadsnodpdsoaaddaqae 26 
WENT, TOBKE OO Ascoasbdoakoe 201 
MMSE AVA Le aula naan 26 
PETS ale OE TU MOMMA ali ee catia 26 
PLC ALA ARSC HNMR eae a a 69 
GM SECONLISH Eye Naan teEEnu ts 26 
SIMPSON Ai Nee ne nicelcatat 66 


Streptoploraa a. haa eley aan 68 
VERUS TAMU eae 233 
VITO en a ee 230 
Pleurotomaria quoyana............ 164 
Plaurotomella agassizi............... 68 
blakeanay i Cae 68 
IDREVES ANNs U3 aa 68 
diomedece siege eee 72 
EXTECH SAN Mes sevais «ose ceneeeeeeeen 68 
IGUBUH EY 21 os MONEE MER EES oad 00.c 68 
eal eee lie 5.) a) a 68 
ATT CTT SURE No 1 RN Ve 69 
TeWCO nda tasers see Se 67 
Wales s00\e> OTE s osaopodosce 68 
packardaye ces... iss eeaee eres 67 
Piicatulaydensatas-.\ 5-5 380 
Manoiwatay) Gece 203, 380 
Polinices eminuloides..... asta 148 
HAbaKeAU ey PN ocosadoaaso dos BARES 148 
JOKES TANGO] DHIhg dogtdocosbocessooes$ 148 
Veptale ayes si lan a aE USN ea eae 148 
stanislas-meunieri 76...196, 329 
Subclausaly e7Olsnaeeeee 196, 329 
PET TTS eh SU ee AE 147 
POULT Aaa as ASR UMS Ray ROD Aaa ae SR 87 
JEL YSCLODIGIES). | osoocatasaccsdaadeaeono6ces 4o 
Potamides, var gracilior ......... 127 
ATL ATS O TMA re se en a 127 
Satillensisueemen parece ecens 130 
Protocardia costaricensis 30...401 
gurabica 30........ 194, 196, 401 
harrisisccuaaeceeescosseeaveseeeaen 25 
Psammobia harrisi ?............... 24 
GyasbUeoll i eos LA penacdoodacsbuouudabodse 23 
Psammosolen gatunensis, var. 
costaricensis 32.......:. 432 
Pseudolivalscalinaeess--eees eee 26 
Evab (CeO DONEAL Sngnoqhacagsadaedecoos 26 
Rteriainormatayy i eeeeree 194, 367 
Ptychosalpinx dentalis 7S...... 187, 
281 
Puncturella circulars. eerscse 166 


triancullata we pos eeeee re reeaes 167 


477 Costa R1icA MiocENE—OLSSON 305 


Paneturellay trifolimimis yess 166 
WIVES OPIMUE i nna Mclain large 166 
JPARN FONTAN Dies RAS era ate ae 100 
TO CUI OS are eves) pei alenin ene an ys Iol 
TAO COU Bb Ibe esayronesenadeuncnase 305 
Ryramuidellal candida... 0h.) 20 107 
Crermiullanca ye Sea Aan Manure Sn 107 
Golalorataye ee i ue le 107 
aA CLO SUNOS de aeaHalannoussenbien: 10 
ODEWSOIGES ie 7s eee eee a(@) 
SVTTHHTS co aa ee 107 
isa ita areas oe sk lobe es Soka oe 108 
Ryiulagabbreviata... 04. 2..6s0.se.: 101 
PAUP AIAN e 6 .scciaaneeccuensnce 119 
R 
TRavaSI Des oN UAE Bane aii LOS eAM TEA! 
Clavelmrartaaen ey ede sire ba 99 
CRASS a ee ris tte nicislerecia aaa Slearels 306 
IRGCHNSAL IBMIIS ues cso sdedandadeduasedse 49 
AZO CIMT SE eee ey cen och eeene ene IO 
Rucinmlalnodulosa...)62)5....0- 305 
IRGBEMTTD IE seo yeaa SER EEE MGR HEM Sena E Man: 167 
AS HUTS ATA geese eecste aaa! 167 
Ringicula dominicana.......194, 196, 
206d 
IOS OPA TARO aioh hodnmbecocakbasene 186 
Ke ptochielansy eye eae 47 
FTW lh, ososoeaeaaACMeUoreeebe aeolas 47 
SEMIS tslaicaa meee eee nie ces 206d 
Rissoa acutiocostata................. 139 
CMSA A IN Pee Ee PENAL 139 
PSRCIPICAT AL. le aes keels 138 
Rissoina alabamensis................. 26 
LOONATENOE, ccosnagds soso suosdeneE 139 
Chesmeliprara tts ssce socks este 139 
GECHSSA tA MUU ea: 139 
elegamitissimialeneeeo-ce ose 140 
Ben raeren czy OUP Sa RAISE ee a 139 
SAG AIAM Aes MN nee en seal lne, 140 
Rochefortia minuta 2............... 20 
Rotella cryptospira.....-:.......... 162 
S) 


Sabatia bathymophila............... 50 


Sanguinolaria alouata 32........ 432 
Santo Domingo Miocene........... 92 
Saxicavella alabamensis 3........21 
aly Sa OUe ie nha e yk etn Kant asian 56 
Scallania raat Woe iy webs es OM 102 
Scaphander librarius.................49 
161(0) Oy LIS ape AnABdrlespacbeonaanocbeen 49 
PMU CLOSER EMS Heese enee nee Re esoe 49 
WALES OTM! ML eee Lia ad ana 49 
Scaphella costaricana............... 187 
COStamICenSis 7Speeeeseeecer eee 272 
[MELON aN e eee ee ae es 81 
Scobinellaymorieretim7scee ee 251 
Sconsia bocasensis 75.........-.. 309 
Cocleamayynseeeesedsshe 187, 310 
ligewlcatalateeecaeees 195, 200, 201 
Wate Sallis ieeeenceace 308 
7ZAOL OK US UNEASE ENE 193 
GUIDE SN ERMA conusanaddstoesaeeaso 200 
SYSTUADVSTOWAIE. ys Srisissoguasonocabeqcbabesec: 158 
fORMOSA Nasa nsenseacaceeeeeeeens 120 
ACOMICAL Maa vensirescustacsedsn nue 121 
MONO CINTA tale tenes neee: 120 
(UPIGFODEDOS 27, saons adnooadoosbeasdcas 120 
Senlavadian sien eesec hase eres 124 
Semele claytoni 32....194, 196, 431 
DEE VAS HESS UNE rae maa I9l 
var. costaricensis 32....430 
iewoverckoratlalialey 1 Beso ccuseba ionsseos 18 
ONOVOVTOLESUIEHS) | Zocsacuecoccosasocooss 19 
SEN Ol S72) Osea ncosdacdecosidasgacee 430 
S Era CASSIS sees een ue tiaeed Mane een Neen 118 
Semiunellalstearisiy wesw ete seen elt 94 
Serpulorbis papulosa 75...196, 200, 
317 
SCulpPLUravUS ssseeeasese see heee 200 
SHEEN TEESE wesc bogne! ob: Hops opoddsmonon. 150 
SALUMEMSISM ses aeseee lee sacra eee 186 
TEMAS vAptEwOVbbOOl p audod codonoossnde 150 
Zab bie ee eoe auan ees naeee ys 186 
Siliquaria squamata..............-... 133 
IveoKOWOVETHIGTS) cosconaccpoceos TOS 
MIO esta ena wi we 134, 188 


306 BULLETIN 39 478 


Siliquaria modesta, var. limo- 


NEMSIS |) 7G i eee: Eta 319 
Sinppiapiaterme diay ee eeu sees 112 
UBIplicatar i. wees. bb SbodeHOdO 112 
Sinum pmacula tums essen eee 150 
Spoahbhaol saoNWOKOS Me ndoas4Auas /abrcoaduade 150 
PEKSPECEIVU MIME eae sees 150 


Siphonalia quadrilineata 2........15 
Siphonaria alternata...........0......58 


lanate@ arta eae UST ORs Me neN 58 
Siphonium nebulosum.. ..... ..... 133 
Sistrum nodulosum 7}............ Lol 

190, 305 

TOS CUTIE MIURA ile aes IoI 
Skeneaplanonbis: eeeseselasceaces 140 
Solaricllatzeclecsi) uaa eaE5 7 

alaibamenSisneees yuna a aeene tate 26 

AIMADILIS eee e ee Meee Mie sal 156 

Wars ClavaitaaleMuteniM sect. 157 
Varo cideal sna nen ie 157 

lissoconchiayy swear ie 157 

Dub ri Sa se eM ee UNS Ae) 157 

SCalbrclus culliaaene en nae ee 156 
Solarium gatuniense ie 326 

Scat UTM hue ease sstanets 326 

STAGES NG hbWaOL  oneghdoogonedadodon 26 

svatll areal Loni e eu nee meme 200, 326 
Solecurtus gatunensis.............. 432 

Strieabla tus yea ee aaa 186 
Solenosteira chiriquiensis 77...286 

dallas: UTD AUS ERR Lee ers 285 

var. medioamericano...200 

Vvaul shanty ea tenon aua ny 200 

var. medioamericana 

UA hc cchbaCoo Han todaaugcHECondsSod 285 
SPintalisinrostrallisn yay eee eM eE 44 
Spondylus americanus.............. 379 

SlaheapenlesmeyG’) ey ee 378 

gumanomocon 24...... 194, 379 
Strepsidura heilprini........... DRL 25 
Strigilila) musamicalwi2o4 eae 427 


Pisilonmiicieey eye Heey 194, 200, 426 


PIStoriis eee eee wanes 189 
Strombina ambigua }........... 298 
Chiniqmiensis i7opr esses eee 302 
costaricensis 7}...... Bae 300 
CaCO) | 7 careanocor=cassanoe 299 
gibberula, var. galvestonen- 
SSRN ae dc cA A a aN gI 
NESSEPONEITE, | Ro asrbacdeoussconedc 2098 
UL AGLIMV AN OTge eta a. 8h 299 
var. musanica 7}........ 301 
Sim COL aig 0.55) sete een 301 
Strombus accipitrinus.............. 120 
(COSWERGBIS) Gadsobote qooscddes sovocuaKe 120 
SACUMESISIS| Ose eae 188, 313 
pugiloides /6..... 187, 196, 314 
UA S ee ee eee 119, 200, 314 
Stylio lag wea a ae UVM NaN nae 43 
WAL OUD A a aces ea eat e 43 
SHsMoOe), Ghashablodoueosooeseacueac sooder: 107 
T 
Macelisice bushi eee eee 433 
Ma Pes Syl Saks Nae Mile Beara 420 
Maranisnpulchieliiay assays woneenee 72 
Tectarius municatus.../2222.0.04.5;. URz 
Teinostoma cryptospira....... eee O2 
ROSES). Wicaasecaodsao) dopoocessos 12 
subangulata, var. smithi 7...11 
Tellidora cristata 29........ 189, 426 
HeliinaibelistanaG 25s seeeeeee eee 17 
cercadicaniaqa eae 194, 196, 425 
Costanicanal Zone cceeees 423 
ery stalllimay2oeeeneec 189, I9I, 
194, 421 
CyMOSLOSSANN eer aceeeneeattee 17 
dariena) Zo eerste: 424 
equitermiima tae eee peerearee 186 
esteliltenmsisiy 2 eeneeeseeee 17, 25 
seAN A DNOVOSH 3 54s2ndhoedoue yuocpedoe 429 
KEW ake MONA LUEBH ann MAM TU Mm 7 
leprcota yi Zones aaeaee 200, 423 
VAST APL U. catalase edb geeata 422 


479 Costa RicA MI0CRENE—COLSSON 307 


THONVEUIEN 21a) AON ee saat 421 

TRON AYRE OKG bg ayy Marae R Hn ta aD a 424 
SQM GOVAG case nantesnececue ee ees 424 
GTA ORDA, VAAN Sadao sot sodocd 18 
SemmlinOpit Gama veh ses sae seeeeeeet 18 
MOAN OME tanh eet es 2OGM AG 
Tenuiactzeon pertenuis 2......... 16 
herebrasbembmMalis.) ssc.) 202s 189 
var. boscasensis {4...... 213 
loerliimeces ) Veaneuoocacuounn 194, 211 
ipancitayyegs) sc... 194, 199, 207 
Wali OCASCHSIS He sneccees. sees 189 
CATO MIMMETISIS Hacer essen ueaee 60 
(CINGVETEE. ona oS sen BABAR eH bee car 58 
COMEA VAR aE MMe a uel ae NPE LL 59 
COSHATICEMSIS/ WI Zin nese tessa: 212 
iSO cata weno une Ch a 59 
Smell vaey | ess codmoneouosscon 212 
SalVeEStOMenSiShess.. sho eeeee nce 60 
SALUT SMSISI ua esecces ince scee 208 
PAUSAP AVANTE seesieictslerleeiceles leis 210 
HMAItEMSIS NZ sce ss scees 194, 207 
HATMATCETMSISH (eh. Ub seskiless ess: 58 

Te sayexGVovratly od sdeeae ciated an uRal 60 
sinner total eee LA ee ce ae 60 
INBHtESOSTINS A oodans, eocsduoaseseoancHe 60 
War. tmiddenSis) |) ae. 211 

ASSUME Ne aseaeatisraiens dete Saige ess 60 
TOSENGT a cb eunt gem asnconALaResPE sa onect 59 
]OIROUED XU eacnouce  encdonod noses uoEsnod 59 
PULA DILSTSIS| ieee a EEE Bua 
TRIBKG IS ae AN A Se a Men esi 59 
SANSA ecu e Gace Wee Lies 58 
Gyovberbiese 72 Goosaasoouaoss 194, 210 
subsulcifera.......... PN Oi 

USS EEE (SOSH ER SER aM EARS 60 

\/GUT COSTED un nai nO MSE ee 59 

Tr Olbieteavest |" Has Aaabebocoboe 208, 209 
Teredo ringens 2............... 1725 
ANS WAS FOROS Gav daoee asusodusodsonue 53 
WWBWUICob-ald. Beau doncoal sbedemaeaeeHeen 53 
MblTanSiG elitoid Casneeewseecseausccees oi 100 


Juleehaavalsyovanlalys phan oqunooaeasandane 100 
Wags anon shale soceoodcocgoseoc 100 
JOENABI EL ouh ca wenoobood aggesonodudodne 99 
MINEO COX Ses tee ecenceestelecees sais 163 
Thracia gastunensis.............-..- 187 
Hite SEA TINY 2G ese elesiselelsleletertsl/=le'or 383 
Aone) AVI Ws oaodosagosbesopassede: 119 
PETC UKEP sese.speescenelnnnns sons 119 
AbloNenl Fal aAsuansodovsdosaosaeadspescesoene 134 
orinianCanalikeraseneesscsetasscet 138 
CylimGriCa. so... ses eecns ener oxen 138 
Tornatella quercollis................. 26 
Tornatina) Ovi a cee canoe AT 
coix-lacryma.............---+-- 206b 
Trailrancdencuilatarcessweseecericelscicts 57 
THATS CU) Aeekecmesecescsecelelece 57 
MY OSOLIS. ....06.....cceceeereeeee ees 57 
PUSIlla..........e ee eee eee seen seen 57 
triplicata..........+..-seeeseeeeeee es Ti 
Trigonostma ........2.-sesseeee sree eee 74 
Triphora abrupta...........--..++0+- 122 
bigemma........--....-eseeeeer ee: 122 
ENG iecaodehoundaaseoasadoccH ones 121 
cylindrella.............2+2++---- 23 
GIACOHFEI| sadognosdoscuouedpsoaeusee 121 
ISURCLBIS) gouooancdossobs pdgssbaasasonbc 122 
sabi  NVEbies KES icacHanaundossuoune 2 
Hole Ntlsnadcavossbandoass boobeasednde 122 
perversa, var. nigrocincta...121 
(HONMUKCDU 2), 5 odod6e suoeeeoonDoboSERs 122 
RUIeIGYeVe ULE A Gante Hadonanee aaaceogse 121 
GBray quence mays dseehosedasaasoadeszeoosaso5500 44 
P Rrerkuova i Gdav DMNA AON AR eSB ate oMsGN eee 115 
CEhelBEDTSL .Hoguongaas Gs5d “323495 117 
lacunellus...........-- -.--.- ----- 99 
PAL VUSHesanseor sere ce scent censecian 88 
pulchellus...............-.2:+2++- 116 
Showaltetivnescwssee beseca scr 26 
GyoambUbbadtbISy, onsso5e5 posoHooDoCORRbE 305 
A WaltoyaoUl@el. do ouuscoaas4secosonSuaocHcenac 87 
Trivia approximans......... ..-.+-- 114 


@arverabiolblley candadsosbsupssan Goose 114 


308 BULLETIN 39 480 


MirivaayclODOSai sens) dear eea asa sacres II4 
OLOT a eee OA Matyi alae IIl4 
Peddculus yy eee deweNewesidas 113 
UL aye UO ee ere ea rale ee are II4 
Sphere ee ence eau ieee 114 
Suih Sa eee Tue alas 114 

Trochus agglutinans. .............-. 332 
Ebon iel) on bi aao4 hon Hse ceaete HaEBRaGOu 156 
Venticulanise estate: 129 

Truncatella bilabiata............... 142 
CALIMAETISISH ee see eee eee 142 
Pulchellanwrscaseeesstnece seers 142 
subcylindricay ey ies.ceteseeese X43 

ar boalllpicdateeee sie eeesene castes 153 
CAStATT CUS Mp e ssa asceecne estes clieres 152 
Crenulatus ee ee eeaeease 153 
aA |bbA(CRRUDISYS denadanan | Caseoucaouce 137 
oul chre ll ae ee eee es seneleeeter 152 
Salts ie Tor wea eaasi nee 333 
Spengleniamuse see) canes lene 333 

AUR D OMI ae Mey anaaie Nan gna Hl ema ats 107 
belothecane ry... Te HeOSHE LT 108 
CE GbKO Sa ducdudodedseeeasue acueneac IIo 
Chinensisi ii ee ea Gia 9 
COTTA CAR eee eee 109 
CUT Beem eeu clacie aaecivasactetcleiscts/af= 109 
Ge-cU ba riscaelad donachtiag sescadauader 108 
INCE LHW play Mey eeee sascue eee eet 108 
ObeliScusH see eee. BUEN SA 10g 
[DLEDTTEEOEEY | dines Acanopaosecosaauddeda 109 
Leb Culatarc ees seesecectse sacl 109 
G) nS) bralg SAadsdan ecaadeadacedadaedade sexe) 
StylifOrmis ............2- seeeeeeee sate) 
tardisculusiy ese cesses eer 9 
ilep:< GU DIS RENO BeM EAB Ba SS SRO HeRE 108 
tuseahomenSis zee eed eecce= 9 
PUREE sen saotueaseooabecasosoocdasrood 109 
Vinbaks FERGIE Goobocoa Jacoepeouodneseeec 108 

Aurcricuilavlav Amare eeeeen es ee eee IgI 
lawanO1deSun7aenssases see IQl, 227 

var. limonensis 7....... 228 
(gapbariate), |) bicgdqsacos, waceoouos6ae06 229 


TIP eyAL V igbyabarckoodaceacedan seed 229 


Turris albida 7...62, 190, 194, 199, 


230 

Webes WAU 6 Gocuseda Keodoodce 62 
binronmtatus ene eee ee seraccet eer Ti 
denticwla tay eee eee sea serene 6 
TmaKOVIHVVENEBI ssacengag dacuenocoosodso 8 
TVOYOIONGIENG) Zo ccouddcabe: sadnovdoe- 5 
SPOXCHIS | Ua socandonagssnnocoo Buse Ge 6 
Miarpitellayacroporayeysseeeeeeeeeee 134 
alalpatmienSiSs ve-cs-ceeneeeserece 25 
evlonbbieh, 17775545 akaoe [SS B2l 322 


var. chiriquiensis 77...322 
var. costaricensis 77...323 


Chipolama aes Pas eeeeee cree 135 
COUTAGT AIRS 2 5) eee met 320 
ExOletarn alia a aeeEnees 134, 188 
var. limonensis 77.....224 
Pal DU OS eek cen ena 221 
SALT SISIS 7 pears eee eee 320 
lohbhasVSCost le ccdabou=ob sorodoondooos0d 25 
var. limonensis........... 188 
UONHATSWES | IH cae ooaasooapegadehes 321 
TNOLLO MI yee eane ener ecerase 25 
OMAXOOOD 17? docchongonascecyeons 324 
Sata ea Cha se ues eal eye 135 
Subexumdiheras se. tane eee 135 
tenebmatoriadsh eee eeeanere 135 
CeTmMeESSeEMSISH 4. coe neceee aoe 25 
WOMEN HEI de GugencconadUsesconeooeoud 323 
NW/BNEISTINE) | Lobo sgnaseonsed opoccca0n 134 
VWillCalecaiasee nasser cere 134 
diy plhisvalbatiism eee erste 195, 304 
eFN ODP UAE Aap banabuoodassangoouspoe 186 

linguliferus, var, costarien- 
se aedannnsndenonascHooGood 304 

U 

(OMS TEES SAcgusdedenweGasesess cosoccoco000 112 
Uimbonium jbaimdieeens.. eee 158 
Wrosalpitsx| cinereus: pencnaeen 97, 282 
OVATE Coos setnnoudgcssoodoosdeos 98 
perrugatus......-....0............-. 97 
{iehan OB DOSS So go nndoodcosecedocosoade 97 


48X Costa RicA MIocENE—OLSSON 309 


TONES SAE a AINE 48 
V 
Vaginella chipolana .................. 43 
Wateoatnttll aise ansciecsiaisccevetulelceusscsseee 58 
Venericardia conradiana, var. 
ItmonmenSiS: 9/29... ..e.sce- 389 
scabricostata 35...194, 200, 388 
CaTaysIMMME OM ss eaaldceciesece se as 388 
WHULGOREIIGIS peauetosesueBedosouesoae 23 
Wemusmblanidiana. 62 .5.... stec eens ne 
CIOSRBEMIVAT soaacase wsdeoneeodeseay 200 
AITACIUAL ARE Ne ccs) scciioecies wetioes 4oo 
AINA SHAM CARMA setae cients eee 412 
TTMUNELCOS teas asc eee ees ee 4it 
PU SOS ARM Ae siete) eslerceieisw eis susiee( 412 
Vermetus papulosus................. 317 
UChr reo soem scate sacar 200 
SCHIP EMA LTS Ie ese eee eee aes 318 
Vermicularia nigricans............ 133 
Sioitertaemaudey unas sa! ALUN 132 
Vermiculus spirata............-..-.- 132 
Veronicella floridana.................58 
Vitrinella hemphilli........... Naa 159 
TIDNESHCTT Bl ORE Seounoneaneeacencosoaaon 159 
TNOOREAM As scone ne wen scene meee 159 
AMAUtCaTAMatas.- 2). 4h.ss.e os 161 


IGEN WOME SAL aA eae RRS ream alotay 
Chill ONAN sc pApcuocoseodéocuceoudsonDdceN. 81 
VATESCENS patina us rovesesecesnnenans 81 
Volutilithes limopsis ................. 25 
TOBA GNSS Gagos sdadogsosdonodoooudAnAge 25 
WOE) oobaGdasssobanbooDsHoccdoodaded 80 
Wel Aichi ke Wee aaioRtrricoc ha caiSceSHensbanoaea 48 
WO EWaITelS ooscodbaopoade acodonquaccacogace So 
Volvula cylindrica.....190, 194, 199, 
206¢ 
Volvulella micratracta...... ..... 186 
WwW 
Wolliliatratulenr eb Sipeeereean ease iessesce: 58 
X 
Xan CUSMS COIS Hees seseielssiece eet 284 
SOO) DUNKS 7.22.55 Socoseo \aobensn0n 283 
Xenophora agglutinans............ 146 
Cais DaSalyeniasaee woes nace aenioceercisee 136 
conchyliophora........... 145, 332 
Y 
Yoldia bocasensis 37/........ ...-. 348 
eborea........ GON I AN ea ca Naan 25 
Z 
TINA AO WBMP 4, Seocbaneanpooosode Se 36000° 154 


" 


ADDENDA ET CORRIGENDA 


NotEe.—-All page references are to the Volume and not the separate 


Page 


47 
58 


WS 


A 
81 


IOI 
104 


164 
188 
206a 
275 
306 


318 
405 


Bulletins. 


For RNIGICULA read RINGICULA. 

For Siphonaria lineclata d’Orb. (reme frecectziec) 
substitute S. xaufraga Stearns. 

For Olftva circinata Marrat substitute O. sayana Ra- 
venal (Cat. p. 19, 1834) and place civcinata in 
synonymy. 

For Marginella aptcinca read apicina. 

For Scaphella junonia substitute Maculopeplum junon- 
zum, this species being the type of Maculopeplum 
Dall, 1906. 

For Epztomium read Epitonium. 

Janthina communts is regarded by Johnson (Nautilus, 
1922) asidentical with /. janthina. Bartsch, how- 
ever, considered communis and fragilis (equals 
7anthina) to be distinct (Bull U. S. N. M. 1915). 
The latter opinion is followed here. 

For Petrotrochus read Perotrochus. 

For Chione Rowelli read ChioneRowleet. 

After Acteon costaricensis, n..sp._ insert Pl. 15, fig. 15. 

For fuses mtocosmins read fusus miocosmius. 

After Bursa crassa, tor Pl. 15, fig. 15 read Pl. 15 fig. 19. 

Home Belwea readily GaWeas 

For Gafrarium limonensts read Gafrarium limonense. 


No. 39, Pil 


Bull. Amer. Paleont. 


9, 


Pl. 4, Vol. 


<————e 
ee 


Se 
Sea ee 


Ha, Se 
ee We 
he, ‘ 


PI. 5, Vol. 9, Bull. Amer. Paleont. No. 39, Pl. 2 


OLSSON, PHOT. 


ITHACA ENGR. CO. 


Seonli 


PI. 6, Vol. 9, Bull. Amer. Paleont. No. 39, PI. 


OLSSOW, PHOT. ITHACA ENGR. CO. 


‘ 
' 
4 
a 
iY 


4 


No. 39, Pl. 


Bull. Amer. Paleont. 


9, 


PI. 7, Vol. 


ITHACA ENGR. CO 


OLSSON, PHOT. 


i : ; cearne! uf 
; : cere Heat a 
. et Sy f 
, pala’ ? 3 ig 
y y 
Maer 4 + 
ithe ieee ‘ < 
4) Li ca 
‘ 4 i 
; +e f vi 
& ty bx MT ) 
2 f 
res ¥ ie 
F hS 
5 Kae 
¥ f cn 
a 
5 x Mr 5 
Pity 
rat 
‘ - 
* rns 
- ig ~ 
¥ ty 
} : 2 é 
us a i 
¥ sy 
mint 
“ > 
* pte 
‘ Fa at P 
NX 
= 4 i 
‘ - * 
£ 5 6 
a 
ye } 
te! ¢ j 
hay ae 
- MDs rF 
1 


Bull. Amer. Paieont. No. 39, Pl. 5 


Pl. 8, Vol. 9, 


ITHACA ENGR. CO 


OLSSON, PHOT. 


han 
FiSteeD) 
rae 


ee 


is 
alates 
7a 


Hes 


No. 39, Pl. 6 


Bull. Amer. Paleont. 


ITHACA ENGR, CO 


OLSSON, PHOTO. 


Pl. 10, Vol. 9, Bull. Amer. Paleont. No. 39, Fl. 7 


OLSSON, PHOTO. 


ITHACA ENGR. CO 


No. 39, Pl. 8 


Bull. Amer. Paleont. 


Pl. tl, Vol. 9, 


ITHACA ENGR. CO. 


OLSSON, PHOTO 


fed oto” 


No. 39, Pl. 9 


Bull. Amer. Paleont. 


PI. 12, Vol. 9, 


ITHACA ENGR. CO. 


OLSSON, PHOTO 


era 


i 
Vals 
i de lan 

pea 
ie 


10 


No. 39, PI. 


Bull. Amer. Paleont. 


PI. 13, Vol. 9, 


ITHACA ENGR, CO. 


OLSSON, PHOTO 


peace lg 
“4 
Te v 


anes 


PI. 14, 


Vol. 9, 


Bull. Amer. Paleont. 


No. 39, 


Pl. 


No. 39, Pl. 12 


Bull. Amer. Paleont. 


PI. 15, Vol. 9, 


ITHACA ENGR. CO- 


OLSSON, PHOTO 


Pl. 16, Vol. 9, Bull. Amer. Paleont. No. 39, Pl. 13 


Se 


OLSSON, PHOTO ‘ITHACA ENGR, CO 


Pl. 17, Vol. 9, Bull. Amer. Paleont. No. 39, Pl 14 


OLSSON, PHOTO ITHACA ENGR. CO 


GON 
Mod 


No. 39, Pl. 15 


Bull. Amer. Paleont. 


PI. 18, Vol. 9, 


/THACA ENGR. CO 


OLSSON, PHOTO 


~ PILI9, Vol. 9, Bull. Amer. Paleont. No. 39, Pl. 16 


1 


- OLSSON, PHOTO : ITHACA ENGR. CO 


i 
opie ete 
phe 


© 


FP) 
ne TO 
Net ae 


Pl. 20, Vol. 9, Bull. Amer. Paleont. No. 39, Pl. 17 


OLSSON, PHOTO ITHACA ENGR. CO 


Pl. 21, Vol. 9, Bull. Amer. Paleont. No. 39, Pi. 18 


OLSSON, PHOTO : ITHACA ENGR. CO 


Pi. 22, Vol. 9, Bull. Amer. Paleont. No. 39, Pl. 19 


OLSSON, PHOTO ‘ ITHACA ENGR. CO 


No. 39, Pl. 20 


Bull. Amer. Paleont. 


9, 


Pl. 23, Vol. 


ITHACA ENGR. CO 


OLSSON, PHOTO 


Pl. 24 Vol. 9, Bull. Amer. Paleont. No. 39, Pl. 2l 


OLSSON, FHOTO. : /THACA ENGR. CO 


Pi. 25 Vol. 9, Bull. Amer. Paleont. No. 39, Pl. 22 


OLSSON, PHOTO, ; (THACA ENGR. CO 


Pg 


Seas jativ4 
ETN 
hatin 


© 


eebis 3 
es ga: 


Pi. 26, Voi. 9, Bull. Amer. Paleont. No. 39, Pl. 23 


avulosninae 


oo 


8 a , ; ow 
“OLE Ei 


OLSSON, PHOTO ‘ ITHACA ENGR. CO 


We 
eit 
Tet aan 


eS 
es Maen 


No. 39, Pl. 24 


Bull. Amer. Paleont. 


Pl. 27 Vol. 9, 


ITHACA ENGR. CO 


OLSSON, PHOTO 


m 
i 


me Beas 
Raa 


Pl. 28, Vol. 9, Bull. Amer. Paleont. No. 39, Pl. 25 


OLSSON, PHOTO /THACA ENGR. CO 


Pl, 29 Vol. 9, Bull. Amer. Paleont. No. 39, Pl 26 


fe 


pa 


é Spo 
Go: ommaie 


OLSSON, PHOTO /THACA ENGR. CO 


Bs 


Pl. 30, Vol. 9, Bull. Amer. Paleont. No. 39, Pl. 27 


OLSSON, PHOTO : ; ITHACA ENGR. CO 


7°, 


UAT 


ae 


Pl. 31, Vol. 9, Bull. Amer. Paleont. No. 39, Pl. 28 


OLSSON, PHOTO /THACA ENGR. CO 


POO REL, 
SAGO Ha er 


Pui 
ae 


pyce 


Mint” 


Pl. 32 Vol. 9, 


OLSSON, PHOTO. 


Bull. Amer. Paleont. 


No. 39, Pl. 29 


/THACA ENGR. CO 


Pl. 33 Vol. 9, Bull. Amer. Paleont. No. 39, Pl. 30 


OLSSON, PHOTO ITHACA ENGR. CO 


i 


} ¥ 
Bs i 
2 
(5 
fi} 
SS 
t 
ne 
. 
aU) 


Pl. 34 Vol. 9, Bull. Amer. Paleont. ; No. 39, Pl. 31 


OLSSON, PHOTO ; ITHACA ENGR. CO 


No. 39, Pl. 32 


Bull. Amer. Paleont. 


9, 


Pl, 35 Vol. 


ITHACA ENGR. CO 


OLSSON, PHOTO 


Hl} | 


Ill || H] II IM HI | || | |